Dinichthys
Encyclopedia
Dinichthys herzeri is an extinct, giant, marine arthrodire placoderm from the Late Devonian
Devonian
The Devonian is a geologic period and system of the Paleozoic Era spanning from the end of the Silurian Period, about 416.0 ± 2.8 Mya , to the beginning of the Carboniferous Period, about 359.2 ± 2.5 Mya...

 (Famennian
Famennian
The Famennian is one of two faunal stages in the Late Devonian epoch. It lasted from 374.5 ± 2.6 million years ago to 359.2 ± 2.5 million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage and is named after Famenne, a natural region in southern Belgium.It was...

 stage) of Ohio
Ohio
Ohio is a Midwestern state in the United States. The 34th largest state by area in the U.S.,it is the 7th‑most populous with over 11.5 million residents, containing several major American cities and seven metropolitan areas with populations of 500,000 or more.The state's capital is Columbus...

 and Tennessee
Tennessee
Tennessee is a U.S. state located in the Southeastern United States. It has a population of 6,346,105, making it the nation's 17th-largest state by population, and covers , making it the 36th-largest by total land area...

. It was comparable in size, shape and ecological role
Hypercarnivore
A hypercarnivore is an animal which has a diet that is more than 70% meat, with the balance consisting of non-animal foods such as fungi, fruits or other plant material. Some examples include the big cats, dolphins, eagles, snakes, marlin, most sharks, and such invertebrates as octopuses and sea...

 to the better known Dunkleosteus
Dunkleosteus
Dunkleosteus is a genus of prehistoric fish, one of the largest arthrodire placoderms ever to have lived, existing during the Late Devonian period, about 380-360 million years ago.This hunter, measuring up to and weighing , was a hypercarnivorous apex predator...

. Originally described in 1868 by John Newberry on the basis of an incomplete skull roof and mandibles (AMNH
American Museum of Natural History
The American Museum of Natural History , located on the Upper West Side of Manhattan in New York City, United States, is one of the largest and most celebrated museums in the world...

 81), this species remains imperfectly known to this day.

For much of the 20th century, many unrelated large arthrodires were classified together within this genus, including species now assigned to Dunkleosteus, Eastmanosteus
Eastmanosteus
Eastmanosteus is a fossil genus of dinichthyid placoderms. It was closely related to the giant Dunkleosteus, but differed from that genus in possessing a distinctive tuberculated bone ornament, a differently shaped nuchal plate and a more zig-zagging course of the sutures of the skull roof.Species...

and Titanichthys
Titanichthys
Titanichthys agassizi was a giant, aberrant marine placoderm from the Late Devonian. It approached Dunkleosteus in size and build. Unlike its relative, however, T. agassizi had small, ineffective-looking mouth-plates that lacked a sharp cutting edge...

. Today, Dinichthys is considered a monotypic genus, containing only the type species, D. herzeri. Similarly, in a 2010 analysis, the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

Dinichthyidae that once held a wide range of arthrodire genera was redefined as comprising only Dinichthys.

The type species of Dunkleosteus was originally described as Dinichthys terrelli by Newberry in 1873. After complete exoskeletons of this species were discovered in the early 20th century Din. terrelli served as the basis for life reconstructions of the genus as opposed to the fragmentary Din. herzeri, even long after terrelli was separated into Dunkleosteus by Jean Pierre Lehman in 1956. As a result, most illustrations captioned as Dinichthys are actually pictures of Dunkleosteus
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