Cuban Ivory-billed Woodpecker
Encyclopedia
The Cuban Ivory-billed Woodpecker or Carpintero real (Campephilus principalis bairdii) is, or was, a Cuban
Cubans
Cubans or Cuban people are the inhabitants or citizens of Cuba. Cuba is a multi-ethnic nation, home to people of different ethnic and national backgrounds...

 subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 of the Ivory-billed Woodpecker
Ivory-billed Woodpecker
The Ivory-billed Woodpecker is or was one of the largest woodpeckers in the world, at roughly 20 inches in length and 30 inches in wingspan. It was native to the virgin forests of the southeastern United States...

 (Campephilus principalis). Once classified as a separate species, recent research has indicated that C. p. bairdii may in fact be sufficiently distinct from the nominate to once again be regarded as a species in its own right.

No specimens have been seen since 1987, and it is now considered to be extinct.

Taxonomy and appearance

C. p. bairdii, originally designated a separate species (Campephilus bairdii) by John Cassin
John Cassin
John Cassin was an American ornithologist.He is considered to be one of the giants of American ornithology, and was America's first taxonomist, describing 198 birds not previously mentioned by Alexander Wilson and John James Audubon...

 based on suggestions by Spencer Fullerton Baird
Spencer Fullerton Baird
Spencer Fullerton Baird was an American ornithologist, ichthyologist and herpetologist. Starting in 1850 he was assistant-secretary and later secretary of the Smithsonian Institution in Washington, D.C...

, was similar in appearance to the American birds, being slightly smaller and having minor variations in plumage such as a longer white stripe on the head. A more recent study by Fleischer, Kirchman et al. has however suggested that not only are the Cuban and American forms genetically distinct, but also that they and the Imperial Woodpecker
Imperial Woodpecker
The Imperial Woodpecker is – or was – a member of the woodpecker family Picidae. If it is not extinct, it is the world's largest woodpecker species at 56-60 cm long...

 form a distinct North American clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 within Campephilus
Campephilus
Campephilus is a genus of large American woodpeckers in the family Picidae.This genus was created by George Robert Gray, who suggested taking Campephilus principalis as the type for the genus. The name Campephilus means "lover of grubs" - an allusion to the diet of these birds, many of which feed...

that appeared in the Mid-Pleistocene
Pleistocene
The Pleistocene is the epoch from 2,588,000 to 11,700 years BP that spans the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....

. The methods adopted by the study suggest that the split between C. principalis and the lineage represented by C. p. bairdii and C. imperialis occurred first, and that all three birds should be regarded as representing distinct species. The American Ornithologists' Union Committee on Classification and Nomenclature, while describing the data as "intriguing", has indicated that it is not yet ready to list the American and Cuban forms as two separate species.

Cassin described the species as:


Much resembling C. principalis, but smaller and with the black anterior feathers of the crest longer than those succeeding, which are scarlet. White longitudinal line on the neck reaching quite to the base of the bill. [...] It appears to be one of the singular insular species which have become well known to naturalists.


The first detailed description of the Cuban form's behaviour and habitat was not published until 1893, when Juan Gundlach
Juan Gundlach
Juan Cristóbal Gundlach , born Johannes Christoph Gundlach was a Cuban naturalist and taxonomist.-Biography:...

 included it in Volume 1 of his Ornitología Cubana. In Cuba it was usually known by the name of carpintero real (Royal Woodpecker), although this name was also used for other birds.

Habitat

As with C. principalis, the Cuban Ivory-bill was thought to inhabit old-growth forests with a plentiful supply of dead or dying trees; these were a source of the cerambycid and other beetle larvae that formed the bulk of its diet. Most of Cuba's lowland deciduous forests had been cleared by the early twentieth century, and the species became restricted to the montane pine forests in the north-east of the island. Its original range was given as through the Organ Mountains, in the lowland forests of the Ensenada de Cochinos and along the Hanabana River.

Behaviour

There are relatively few accounts of the bird's behaviour in the wild. The ornithologist John V. Dennis located a few birds in 1948 and noted some of their feeding and other habits, commenting that "they spent so much time [preening and scratching] that I considered it unusual". He observed that they were not especially shy or elusive once they had become used to his presence, eventually "seeming positively lethargic", although a male bird intervened quickly to drive a sparrow-hawk away from the nesting site.

The breeding season of C. p. bairdii occurred from March–June. It has been surmised that the woodpecker's foraging specialisation may have led to its forming small groups, much like Campephilus imperialis which was occasionally reported in groups of eight or more individuals; this behaviour would enable the birds to best exploit a patchily-occurring food source.

Status

Although once common on the island, C. p. bairdii was already very rare by the late 1940s, when Dennis located a small population in a remnant of forest in the Cuchillas de Moa range which had already been cut-over for timber some years previously. George Lamb found six territories still there in 1956, and recommended that a conservation plan was implemented, but the 1959 Cuban Revolution
Cuban Revolution
The Cuban Revolution was an armed revolt by Fidel Castro's 26th of July Movement against the regime of Cuban dictator Fulgencio Batista between 1953 and 1959. Batista was finally ousted on 1 January 1959, and was replaced by a revolutionary government led by Castro...

 was to intervene.

Although there have been several alleged sightings of C. principalis in recent years, C. p. bairdii has not been seen since 1987, when a single female specimen was identified in the mountains of eastern Cuba by Giraldo Alayón and Aimé Pasada, following a handful of observations of both male and female birds by a team of ornithologists, including Lester L. Short
Lester L. Short
Lester Leroy Short is an American ornithologist. His main research field is the order Piciformes.-Biography:Short was born in Port Chester, New York. In 1955 he graduated to Bachelor of Science at the Cornell University...

, in the area of Ojito de Agua, a hilly pine forest. Although the area was immediately designated as protected by the Cuban government, searches in 1991 and 1993 failed to find any further traces of the bird, and it became clear that the specimens seen in 1986-87 had already been in "dire" circumstances. The Cuban Ivory-bill is therefore presumed to have become extinct around 1990 at the latest, though the area given protection in the 1980s is now part of the Alejandro de Humboldt National Park
Alejandro de Humboldt National Park
Parque Nacional Alejandro de Humboldt is a national park in the Cuban provinces of Holguín and Guantánamo. It is named after the German scientist Alexander von Humboldt who visited the island in 1800 and 1801...

.

The IUCN Red List notes that calls were reportedly heard in 1998 in the highest reaches of the Sierra Maestra
Sierra Maestra
Sierra Maestra is a mountain range that runs westward across the south of the old Oriente Province from what is now Guantánamo Province to Niquero in southeast Cuba, rising abruptly from the coast. Some view it as a series of connecting ranges , which joins with others extending to the west...

, but that a subsequent search failed to find any trace of the species or of good potential habitat: it is considered a (remote) possibility that some individuals may survive, as around 80% of suitable habitat in Cuba has yet to be searched.
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