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Coreidae

Coreidae

Overview
Coreidae are a large family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus...

 of insects of the order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

 Hemiptera
Hemiptera
Hemiptera is an order of insects, comprising around 80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, and others. They range in size from 1 mm to around 15 cm, and share a common arrangement of sucking mouthparts ....

 (the "true bugs"), including some of the largest members (> 4 cm/nearly 2 inches) of that group. There are over 1,800 species in some 250 genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

. They generally resemble shield bug
Shield bug
Pentatomoidea is a superfamily of insects in the Heteroptera suborder of the Hemiptera order, which are commonly referred to as shield bugs or stink bugs. Shield bugs have glands in their thorax between the first and second pair of legs which produce a foul smelling liquid...

s (Pentatomoidea) which are fairly, but not extremely close relatives, and include the insect
Insect
Insects are arthropods, having a hard exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae. They are the most diverse group of animals on the planet and include more than a million species that are already described. Insects represent more than half of all...

s known as leaf-footed bugs and squash bugs. These names do not refer to different taxa in this family, but strictly speaking "leaf-footed bugs" is the most unequivocal name.

"Squash bugs" may be any common species that include Cucurbita
Cucurbita
Cucurbita is a genus in the gourd family Cucurbitaceae first cultivated in the Americas and now used in many parts of the world. It includes species grown for their fruit and edible seeds , as well as some species grown only as gourds. They have bicollateral vascular bundles...

(squash
Squash (fruit)
Squashes generally refer to four species of the genus Cucurbita native to Mexico and Central America, also called marrows depending on variety or the nationality of the speaker. It is also natively grown in other parts of North America, and in Europe, India, and Australia...

 and pumpkin
Pumpkin
Pumpkin is a gourd-like squash of the genus Cucurbita and the family Cucurbitaceae . It is a common name of or can refer to cultivars of any one of the species Cucurbita pepo, Cucurbita mixta, Cucurbita maxima, and Cucurbita moschata. They are typically orange or yellow and have many creases...

s) among their foodplants.
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Encyclopedia
Coreidae are a large family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus...

 of insects of the order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

 Hemiptera
Hemiptera
Hemiptera is an order of insects, comprising around 80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, and others. They range in size from 1 mm to around 15 cm, and share a common arrangement of sucking mouthparts ....

 (the "true bugs"), including some of the largest members (> 4 cm/nearly 2 inches) of that group. There are over 1,800 species in some 250 genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

. They generally resemble shield bug
Shield bug
Pentatomoidea is a superfamily of insects in the Heteroptera suborder of the Hemiptera order, which are commonly referred to as shield bugs or stink bugs. Shield bugs have glands in their thorax between the first and second pair of legs which produce a foul smelling liquid...

s (Pentatomoidea) which are fairly, but not extremely close relatives, and include the insect
Insect
Insects are arthropods, having a hard exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae. They are the most diverse group of animals on the planet and include more than a million species that are already described. Insects represent more than half of all...

s known as leaf-footed bugs and squash bugs. These names do not refer to different taxa in this family, but strictly speaking "leaf-footed bugs" is the most unequivocal name.

"Squash bugs" may be any common species that include Cucurbita
Cucurbita
Cucurbita is a genus in the gourd family Cucurbitaceae first cultivated in the Americas and now used in many parts of the world. It includes species grown for their fruit and edible seeds , as well as some species grown only as gourds. They have bicollateral vascular bundles...

(squash
Squash (fruit)
Squashes generally refer to four species of the genus Cucurbita native to Mexico and Central America, also called marrows depending on variety or the nationality of the speaker. It is also natively grown in other parts of North America, and in Europe, India, and Australia...

 and pumpkin
Pumpkin
Pumpkin is a gourd-like squash of the genus Cucurbita and the family Cucurbitaceae . It is a common name of or can refer to cultivars of any one of the species Cucurbita pepo, Cucurbita mixta, Cucurbita maxima, and Cucurbita moschata. They are typically orange or yellow and have many creases...

s) among their foodplants. For example, in North America
North America
North America is the northern continent of the Americas, situated in the Earth's northern hemisphere and in the western hemisphere. It is bordered on the north by the Arctic Ocean, on the east by the North Atlantic Ocean, on the southeast by the Caribbean Sea, and on the west by the North Pacific...

 the orange-tipped leaf-footed bug Anasa tristis is "the" squash bug, while in Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural River, the Caspian Sea, the Caucasus Mountains , and the Black Sea to the southeast...

 it is the dock bug (Coreus marginatus
Coreus marginatus
Coreus marginatus, commonly called the dock bug, is a species of squash bug. It is a speckled brown insect with a broad abdomen, and is common in most parts of Europe...

). Other well-known members of the Coreidae are the Florida leaf-footed bug (Leptoglossus phyllopus
Leptoglossus phyllopus
Leptoglossus phyllopus or Florida leaf-footed bug is a species of leaf-footed bug in the same genus as the western conifer seed bug ....

) and the western conifer seed bug
Western conifer seed bug
The western conifer seed bug, Leptoglossus occidentalis, sometimes abbreviated as WCSB, is a species of true bug in the family Coreidae...

 (L. occidentalis) which is an invasive species
Invasive species
'Invasive species' is a phrase with several definitions. The first definition expresses the phrase in terms of non-indigenous species that adversely affect the habitats they invade economically, environmentally or ecologically...

 currently colonizing Europe (typically, "seed bug
Seed bug
The Lygaeidae are a family in the Hemiptera , comprising twenty-one genera. The family includes insects commonly known as chinch bugs and also some of those known as seed bugs...

s" are members of the not too distantly related Lygaeoidea
Lygaeoidea
The Lygaeoidea are a sizeable superfamily of true bugs, containing the seed bugs and allies. The ash-gray leaf bug family , usually placed herein, might be closer to the Aradoidea.-References:...

 however).
The hindlegs
Arthropod leg
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments are of Latin origin, and may be confused with terms for bones: coxa , trochanter , femur, tibia, tarsus, ischium, metatarsus, carpus, dactylus The arthropod...

 in members of this family are often modified, sometimes in elaborate ways; hence the common name. It appears that males of at least some species use these hind legs in combat over territories. All members of the family are exclusively phytophagous. Leaf-footed bugs enter houses only by accident, when they seek hibernation
Hibernation
Hibernation is a state of inactivity and metabolic depression in animals, characterized by lower body temperature, slower breathing, and lower metabolic rate. Hibernating animals conserve energy, especially during winter when food is short, tapping energy reserves, body fat, at a slow rate...

 quarters for the cold season.

Systematics


This group is most often divided into 3-4 subfamilies; some selected genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

 are also listed here:

Agriopocorinae Miller, 1953 (often included in Coreinae)
  • Agriopocoris Miller, 1953


Coreinae
Coreinae
This sub family is represented in Europe by five tribes :*Anisoscelini*Coreini*Gonocerini*Phyllomorphini*Prionotylini...

Leach, 1815

  • Acanthocephala Laporte, 1833
  • Acanthocerus Palisot, 1818
  • Althos Kirkaldy, 1904
  • Amblyomia Stål, 1870
  • Anasa
    Anasa
    Anasa is a genus of leaf-footed bug in the Coreinae subfamily. Species within this genus are:* A. andresii* A. armigera * A. bellator* A. cornuta* A. maculipes* A. repetita* A. scorbutica* A. tristis * A. uhleri...

    Amyot & Serville, 1843
  • Catorhintha Stål, 1859
  • Chariesterus Laporte, 1833
  • Chelinidea Uhler, 1863
  • Chondrocera Laporte, 1832
  • Cimolus Stål, 1862
  • Coreus
    Coreus
    Coreus is a genus of leaf-footed bug in the Coreinae subfamily. Species within this genus are:* C. marginatus* C. spinigerus...

  • Dallacoris Osuna, 1981
  • Euthochtha Mayr, 1865
  • Ficana
    Ficana
    Ficana was an ancient city of Latium, which figures in Roman history only on the occasion of its conquest by Ancus Marcius, who is said to have removed the inhabitants to Rome, and destroyed the city itself. Ficana was an ancient city of Latium, which figures in Roman history only on the occasion...

    Stål, 1862
  • Hypselonotus Hahn, 1833
  • Leptoglossus
    Leptoglossus
    Leptoglossus is a genus of true bugs in the leaf-footed bug family. Several species are economic pests of agricultural crops. Like members of some other genera in the family, these bugs have leaflike dilations of the hind tibia.Species include:...

    Guérin-Méneville, 1831 – conifer seed bugs

  • Madura Stål, 1860
  • Mamurius Stål, 1862
  • Mozena Amyot & Serville, 1843
  • Namacus Amyot & Serville, 1843
  • Narnia Stål, 1862
  • Nisoscolopocerus Barber, 1928
  • Phthia Stål, 1862
  • Piezogaster Amyot & Serville, 1843
  • Sagotylus Mayr, 1865
  • Savius Stål, 1862
  • Scolopocerus Uhler, 1875
  • Sephina Amyot & Serville, 1843
  • Sethenira Spinola, 1837
  • Spartocera Laporte, 1833
  • Thasus
    Thasus
    Thasus , a son of Poseidon, or Cilix or Agenor, was one of those who set out from Phoenicia in search of Europa, and thus founded the town of Thasos on the homonym island.-References:*Herod, ii. 44, vi. 47 ; Paus. v. 25. § 7 ; Apollod. iii. 1. § 1.) [L. S.]...

    Stål, 1865
  • Zicca Amyot & Serville, 1843



Meropachydinae Stål, 1867
  • Merocoris Perty, 1833

Pseudophloeinae Stål, 1867
  • Ceraleptus Costa, 1847
  • Coriomeris Westwood, 1842


Numerous tribe
Tribe (biology)
In biology, a tribe — or infrafamily — is a taxonomic rank between family and genus. It is sometimes subdivided into subtribes.-See also:* Rank * Rank *International Code of Botanical Nomenclature...

s of the Coreinae have previosuly been proposed for elevation to subfamily rank; for example the Agriopocorini, Colpurini, Hydarini, Phyllomorphini and Procamptini. But the only one of these changes accepted at least by a significant minority of researchers today is the first, and even recent reviews generally tend to treat the proposed Agriopocorinae as a tribe again, recognizing only the three subfamilies that were known by 1867. In addition, at least the genus Eubule is of decidedly indeterminate placement.

External links