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Zooplankton

Zooplankton

Overview



Zooplankton are the heterotroph
Heterotroph
A Heterotroph is an organism that uses organic carbon for growth. This contrasts with autotrophs, such as plants, which are able to directly use sources of energy, such as light to produce organic substrates from inorganic carbon dioxide.- Ecology :Heterotrophs are known as consumers in food...

ic (sometimes detritivorous
Detritivore
Detritivores, also known as detritus feeders or saprophages, are heterotrophs that obtain nutrients by consuming detritus . By doing so, they contribute to decomposition and the nutrient cycles....

) type of plankton
Plankton
Plankton consist of any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. Plankton are defined by their ecological niche rather than their phylogenetic or taxonomic classification...

. Plankton are organisms drifting in the water column
Pelagic zone
Any water in the sea that is not close to the bottom is in the pelagic zone. The word pelagic comes from the Greek πέλαγος or pélagos, which means open sea....

 of ocean
Ocean
An ocean is a large body of saline water, and a principal component of the hydrosphere. Approximately 75% of the Earth's surface is covered by ocean, a continuous body of water that is customarily divided into several principal oceans and smaller seas.More than half of this area is over 3,000...

s, sea
Sea
A sea is any large amount of water filled with animals such as crabs, whales, sharks, and fish, but there is inconsistency as to its precise definition and application. Most commonly, a sea may refer to a large expanse of saline water connected with an ocean, but it is also used sometimes for a...

s, and bodies of fresh water
Fresh Water
Fresh Water is the debut album by Australian rock and blues singer Alison McCallum, released in 1972. Rare for an Australian artist at the time, it came in a gatefold sleeve...

. The name of zooplankton is derived from the Greek zoon , meaning "animal
Animal
Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously...

", and , meaning "wanderer" or "drifter". Many zooplankton are too small to be seen individually with the naked eye
Naked eye
The naked eye is a figure of speech referring to human visual perception that is unaided by enhancing equipment, such as a telescope or microscope. Vision corrected to normal acuity using corrective lenses is considered "naked"...

.

Zooplankton is a broad categorisation spanning a range of organism
Organism
In biology, an organism is any living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homeostasis as a stable whole...

 sizes that includes both small protozoa
Protozoa
Protozoa or Cornelius protozoans Protozoa or Cornelius protozoans Protozoa or Cornelius protozoans (from Greek πρῶτον proton "first" and ζῷα zoa "animals"; singular protozoon; (the word "protozoan" is originally an adjective, used as a noun) are microorganisms classified as unicellular eukaryotes....

ns and large metazoans.
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Zooplankton are the heterotroph
Heterotroph
A Heterotroph is an organism that uses organic carbon for growth. This contrasts with autotrophs, such as plants, which are able to directly use sources of energy, such as light to produce organic substrates from inorganic carbon dioxide.- Ecology :Heterotrophs are known as consumers in food...

ic (sometimes detritivorous
Detritivore
Detritivores, also known as detritus feeders or saprophages, are heterotrophs that obtain nutrients by consuming detritus . By doing so, they contribute to decomposition and the nutrient cycles....

) type of plankton
Plankton
Plankton consist of any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. Plankton are defined by their ecological niche rather than their phylogenetic or taxonomic classification...

. Plankton are organisms drifting in the water column
Pelagic zone
Any water in the sea that is not close to the bottom is in the pelagic zone. The word pelagic comes from the Greek πέλαγος or pélagos, which means open sea....

 of ocean
Ocean
An ocean is a large body of saline water, and a principal component of the hydrosphere. Approximately 75% of the Earth's surface is covered by ocean, a continuous body of water that is customarily divided into several principal oceans and smaller seas.More than half of this area is over 3,000...

s, sea
Sea
A sea is any large amount of water filled with animals such as crabs, whales, sharks, and fish, but there is inconsistency as to its precise definition and application. Most commonly, a sea may refer to a large expanse of saline water connected with an ocean, but it is also used sometimes for a...

s, and bodies of fresh water
Fresh Water
Fresh Water is the debut album by Australian rock and blues singer Alison McCallum, released in 1972. Rare for an Australian artist at the time, it came in a gatefold sleeve...

. The name of zooplankton is derived from the Greek zoon , meaning "animal
Animal
Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously...

", and , meaning "wanderer" or "drifter". Many zooplankton are too small to be seen individually with the naked eye
Naked eye
The naked eye is a figure of speech referring to human visual perception that is unaided by enhancing equipment, such as a telescope or microscope. Vision corrected to normal acuity using corrective lenses is considered "naked"...

.

Ecology


Zooplankton is a broad categorisation spanning a range of organism
Organism
In biology, an organism is any living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homeostasis as a stable whole...

 sizes that includes both small protozoa
Protozoa
Protozoa or Cornelius protozoans Protozoa or Cornelius protozoans Protozoa or Cornelius protozoans (from Greek πρῶτον proton "first" and ζῷα zoa "animals"; singular protozoon; (the word "protozoan" is originally an adjective, used as a noun) are microorganisms classified as unicellular eukaryotes....

ns and large metazoans. It includes holoplankton
Holoplankton
Holoplankton are organisms that are planktonic for their entire life cycle. Examples of holoplankton include diatoms, radiolarians, dinoflagellates, foraminifera, amphipods, krill, copepods, and salps.-Sources:Asexual Holoplankton:1. Copepod...

ic organisms whose complete life cycle
Biological life cycle
A life cycle is a period involving all different generations of a species succeeding each other through means of reproduction, whether through asexual reproduction or sexual reproduction...

 lies within the plankton, and meroplankton
Meroplankton
Meroplankton are organisms that are planktonic for only a part of their life cycles, usually the larval stage. Examples of meroplankton include the larvae of sea urchins, sea stars, crustaceans, marine worms, some marine gastropods and most fish....

ic organisms that spend part of their life cycle in the plankton before graduating to either the nekton
Nekton
Nekton refers to the aggregate of actively swimming aquatic organisms in a body of water able to move independently of water currents....

 or a sessile
Sessility (zoology)
In zoology, sessility is a characteristic of animals which are not able to move about. They are usually permanently attached to a solid substrate of some kind, such as a rock, or the hull of a ship in the case of barnacles. Corals lay down their own substrate....

, benthic
Benthos
Benthos are the organisms which live on, in, or near the seabed, also known as the benthic zone. They live in or near marine sedimentary environments, from tidal pools along the foreshore, out to the continental shelf, and then down to the abyssal depths....

 existence. Although zooplankton are primarily transported by ambient water currents, many have some power of locomotion
Animal locomotion
Animal locomotion, which is the act of self-propulsion by an animal, has many manifestations, including running, jumping and flying. Animals move for a variety of reasons, such as to find food, a mate, or a suitable microhabitat, and to escape predators...

 and use this to avoid predators (as in diel vertical migration
Diel vertical migration
Diel vertical migration refers to a pattern of movement that some organisms living in the ocean's photic zone undertake each day. The word diel comes from the Latin dies , and means a 24-hour period....

) or to increase prey encounter rate.

Ecologically
Ecology
Ecology is the interdisciplinary scientific study of the interactions between organisms and the interactions of these organisms with their environment....

 important protozoan zooplankton groups include the foraminifera
Foraminifera
The Foraminifera, or forams for short, are a large group of amoeboid protists with reticulating pseudopods, fine strands of cytoplasm that branch and merge to form a dynamic net. They typically produce a test, or shell, which can have either one or multiple chambers, some becoming quite elaborate...

ns, radiolarians and dinoflagellate
Dinoflagellate
The dinoflagellates are a large group of flagellate protists. Most are marine plankton, but they are common in fresh water habitats as well. Their populations are distributed depending on temperature, salinity, or depth. About half of all dinoflagellates are photosynthetic, and these make up the...

s (the latter are often mixotroph
Mixotroph
A mixotroph is a organism that can use a mix of different sources of energy and carbon. Possible are alternations between photo- and chemotrophy, between litho- and organotrophy, between auto- and heterotrophy or a combination of it...

ic). Important metazoan zooplankton include cnidaria
Cnidaria
Cnidaria is a phylum containing over 10,000 species of animals found exclusively in aquatic, mostly marine, environments. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey...

ns such as jellyfish
Jellyfish
Jellyfish are free-swimming members of the phylum Cnidaria. Jellyfish have several different morphologies that represent several different cnidarian classes including the Scyphozoa , Staurozoa , Cubozoa , and Hydrozoa Jellyfish (also known as jellies or sea jellies) are free-swimming members of...

 and the Portuguese Man o' War
Portuguese Man o' War
The Portuguese Man o' War , also known as the blue bubble, blue bottle, man-of-war, or the Portuguese man of war, is a jelly-like, marine invertebrate of the family Physaliidae....

; crustaceans such as copepod
Copepod
Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. Many species are planktonic , but more are benthic , and some continental species may live in limno-terrestrial habitats and other wet terrestrial places, such as swamps, under leaf fall in wet forests,...

s and krill
Krill
Krill is a type of shrimp-like marine invertebrate animal. These small crustaceans are important organisms of the zooplankton, particularly as food for baleen whales, manta rays, whale sharks, crabeater seals, and other seals, and a few seabird species that feed almost exclusively on them. Another...

; chaetognaths (arrow worms); molluscs
Mollusca
MolluscsSpelled mollusk in the USA, see reasons given in Rosenberg's ; for the spelling "mollusc" see the reasons given by . are animals belonging to the phylum Mollusca. There are around 93,000 recognized extant species, making it the largest marine phylum with about 23% of...

 such as pteropods; and chordate
Chordate
Chordates are animals which are either vertebrates or one of several closely related invertebrates. They are united by having, for at least some period of their life cycle, a notochord, a hollow dorsal nerve cord, pharyngeal slits, an endostyle, and a post-anal tail...

s such as salp
Salp
A salp is a barrel-shaped, free-floating tunicate. It moves by contracting, thus pumping water through its gelatinous body...

s and juvenile fish
Fish
A fish is any aquatic vertebrate animal that is typically ectothermic , covered with scales, and equipped with two sets of paired fins and several unpaired fins...

. This wide phylogenetic range includes a similarly wide range in feeding
Feeding
Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffix -vore from Latin vorare, meaning 'to devour', or phagy, from Greek φαγειν, meaning 'to eat'....

 behavior: filter feeding
Filter feeder
Filter feeders are animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feeding are clams, krill, sponges, some fish and sharks, and baleen whales. Some birds, such...

, predation
Predation
In ecology, predation describes a biological interaction where a predator feeds on its prey, . Predators may or may not kill their prey prior to feeding on them, but the act of predation always results in the death of the prey...

 and symbiosis
Symbiosis
The term symbosis commonly describes close and often long-term interactions between different biological species...

 with autotrophic phytoplankton
Phytoplankton
Phytoplankton are the autotrophic component of the plankton community. The name comes from the Greek words φυτον , or "plant", and πλαγκτος , meaning "wanderer" or "drifter". Most phytoplankton are too small to be individually seen with the unaided eye...

 as seen in corals. Zooplankton feed on bacterioplankton
Bacterioplankton
Bacterioplankton refers to the bacterial component of the plankton that drifts in the water column. The name comes from the Ancient Greek word , meaning "wanderer" or "drifter" , and , a Latin neologism coined in the 19th century by Christian Gottfried Ehrenberg...

, phytoplankton, other zooplankton (sometimes cannibalistically
Cannibalism (zoology)
In zoology, cannibalism is the act of one individual of a species consuming all or part of another individual of the same species as food. Cannibalism is a common ecological interaction in the animal kingdom and has been recorded for more than 1500 species .Unlike previously believed, cannibalism...

), detritus
Detritus
Detritus is a biological term used to describe dead or waste organic material.Detritus may also refer to:* Detritus , a geological term used to describe the particles of rock produced by weathering...

 (or marine snow
Marine snow
In the deep ocean, marine snow is a continuous shower of mostly organic detritus falling from the upper layers of the water column. Its origin lies in activities within the productive photic zone. Consequently, the prevalence of marine snow changes with seasonal fluctuations in photosynthetic...

) and even nektonic organisms. As a result, zooplankton are primarily found in surface waters where food resources (phytoplankton or other zooplankton) are most abundant.

Through their consumption and processing of phytoplankton (and other food sources), zooplankton play an important role in aquatic food webs, both as a resource for consumers on higher trophic level
Trophic level
In ecology, trophic dynamics is the system of trophic levels , which describes the position that an organism occupies in a food chain — what an organism eats, and what eats the organism.- Energy economy :...

s (including fish), and as a conduit for packaging the organic material in the biological pump
Biological pump
In oceanic biogeochemistry, the biological pump is the sum of a suite of biologically-mediated processes that transport carbon from the surface euphotic zone to the ocean's interior.-Overview:...

. Since they are typically of small size, zooplankton can respond relatively rapidly to increases in phytoplankton abundance, for instance, during the spring bloom
Spring bloom
The spring bloom is a sudden and strong bloom of phytoplankton such as diatoms or dinoflagellates in the spring in temperate and sub-polar bodies of water. In winter, the waters are well mixed, i.e., water is circulated from the bottom to the top of the water column because the water is relatively...

.

Aside from this role in aquatic food webs, zooplankton can also act as an important disease
Infectious disease
An infectious disease is a clinically evident disease resulting from the presence of pathogenic microbial agents, including pathogenic viruses, pathogenic bacteria, fungi, protozoa, multicellular parasites, and aberrant proteins known as prions...

 reservoir
Natural reservoir
Natural reservoir or nidus, refers to the long-term host of the pathogen of an infectious disease. It is often the case that hosts do not get the disease carried by the pathogen or it is carried as a subclinical infection and so asymptomatic and non-lethal...

. They have been found to house the bacterium Vibrio cholerae
Vibrio cholerae
Vibrio cholerae is a motile gram negative curved-rod shaped bacterium with a polar flagellum that causes cholera in humans. V. cholerae and other species of the genus Vibrio belong to the gamma subdivision of the Proteobacteria.  There are two major strains of V. cholerae, classic and El Tor,...

, causative agent of cholera
Cholera
Cholera, sometimes known as Asiatic or epidemic cholera, is an infectious gastroenteritis caused by enterotoxin-producing strains of the bacterium Vibrio cholerae. Transmission to humans occurs through eating food or drinking water contaminated with Vibrio cholerae from other cholera patients...

, by allowing the cholera vibrios to attach to their chitinous exoskeleton
Exoskeleton
An exoskeleton is an external skeleton that supports and protects an animal's body, in contrast to the internal endoskeleton of, for example, a human. Some animals, such as the tortoise, have both an endoskeleton and an exoskeleton...

s. This symbiotic relationship greatly enhances the bacterium's ability to survive in an aquatic environment, as the exoskeleton provides the bacterium with an abundant source of carbon and nitrogen.

See also

  • Bacterioplankton
    Bacterioplankton
    Bacterioplankton refers to the bacterial component of the plankton that drifts in the water column. The name comes from the Ancient Greek word , meaning "wanderer" or "drifter" , and , a Latin neologism coined in the 19th century by Christian Gottfried Ehrenberg...

  • Biological pump
    Biological pump
    In oceanic biogeochemistry, the biological pump is the sum of a suite of biologically-mediated processes that transport carbon from the surface euphotic zone to the ocean's interior.-Overview:...

  • Diel vertical migration
    Diel vertical migration
    Diel vertical migration refers to a pattern of movement that some organisms living in the ocean's photic zone undertake each day. The word diel comes from the Latin dies , and means a 24-hour period....

  • Gelatinous zooplankton
    Gelatinous zooplankton
    Gelatinous zooplankton is the term used to describe the fragile animals that live in the water column in the ocean. They have very delicate bodies that are easily damaged or destroyed. Gelatinous zooplankton are often transparent. All jellyfish are gelatinous zooplankton, but not all gelatinous...

  • Iron fertilization
    Iron fertilization
    Iron fertilization is the intentional introduction of iron to the upper ocean to stimulate a phytoplankton bloom. This is intended to enhance biological productivity, which can benefit the marine food chain and sequester carbon dioxide from the atmosphere...

  • Ocean acidification
    Ocean acidification
    Ocean acidification is the name given to the ongoing decrease in the pH of the Earth's oceans, caused by their uptake of anthropogenic carbon dioxide from the atmosphere...

  • Phytoplankton
    Phytoplankton
    Phytoplankton are the autotrophic component of the plankton community. The name comes from the Greek words φυτον , or "plant", and πλαγκτος , meaning "wanderer" or "drifter". Most phytoplankton are too small to be individually seen with the unaided eye...

  • Plankton
    Plankton
    Plankton consist of any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. Plankton are defined by their ecological niche rather than their phylogenetic or taxonomic classification...

  • Primary production
    Primary production
    400px|thumb|Global oceanic and terrestrial photoautotroph abundance, from September [[1997]] to August 2000. As an estimate of autotroph biomass, it is only a rough indicator of primary production potential, and not an actual estimate of it...


External links