Tapetum (botany)
Encyclopedia
The tapetum is a layer of cells that is found within the pollen sac, a nutritive tissue within the sporangium
Sporangium
A sporangium is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. All plants, fungi, and many other lineages form sporangia at some point in their life cycle...

, particularly within the anther. Tapetum is important for the development of pollen grains. The cells are usually bigger and normally have 2 nuclei per cell. It helps in pollenwall formation, transportation of nutrients to inner side of anther, synthesis of callase enzyme for separation of microspore tetrads. Orbicules
Orbicule
Orbicules are small a-cellular structures of sporopollenin Orbicules (syn. Ubisch bodies, con-peito grains) are small a-cellular structures of sporopollenin Orbicules (syn. Ubisch bodies, con-peito grains) are small a-cellular structures of sporopollenin (known size range from Orbicules (syn....

originate as small lipid droplets, called pro-orbicules, in the rough endoplasmatic reticulum of the tapetum.
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