Staphylininae
Encyclopedia
Staphylininae are a subfamily of rove beetle
Rove beetle
The rove beetles are a large family of beetles, primarily distinguished by their short elytra that leave more than half of their abdomens exposed. With over 46,000 species in thousands of genera, the group is the second largest family of beetles after the Curculionidae...

s (family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 Staphylinidae). They contain the typical rove beetles with their long but fairly robust blunt-headed and -tipped bodies and short elytra, as well as some more unusually-shaped lineages.

Systematics

As it seems, the Staphylininae are part of a large clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 together with the subfamilies Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Paederinae
Paederinae
The Paederinae is a large subfamily of rove beetles . In terms of phylogenetic relationships, the 14 subfamilies of Staphylinidae can be separated into three subgroups; such a classification system links Paederinae most closely with the subfamilies Stapholinidae and Xantholinidae.Paederinae...

, Pseudopsinae, Scydmaeninae, Solieriinae, Steninae, and the extinct Protactinae which are only known from fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

s.

The numerous Staphylininae genera
Genera
Genera is a commercial operating system and development environment for Lisp machines developed by Symbolics. It is essentially a fork of an earlier operating system originating on the MIT AI Lab's Lisp machines which Symbolics had used in common with LMI and Texas Instruments...

 are divided into six tribe
Tribe (biology)
In biology, a tribe is a taxonomic rank between family and genus. It is sometimes subdivided into subtribes.Some examples include the tribes: Canini, Acalypheae, Hominini, Bombini, and Antidesmeae.-See also:* Biological classification* Rank...

s. Some notable genera and species are also listed:
  • Diochini Casey, 1906
    • Antarctothius Coiffait & Saiz
    • Coomania Cameron
    • Diochus Erichson, 1839
      • Diochus electrus
        Diochus electrus
        Diochus electrus is an extinct species of rove beetle in genus Diochus, the only definitive fossil species in subfamily Staphylininae. The species is known only from the middle Eocene, Lutetian stage Baltic amber found in the Blaue Erde deposits, Baltic region, Northern Europe.-History and...

        Chatzimanolis & Engel
        Michael S. Engel
        Michael S. Engel is an American paleontologist and entomologist. He has undertaken field work in Central Asia, Asia Minor, and the Western Hemisphere, and published more than 300 papers in scientific journals. He was trained at the University of Kansas where in 1993 he received a B.S. in Cellular...

        , 2011
        (Lutetian
        Lutetian
        The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between and . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene subepoch...

        , Baltic amber
        Baltic amber
        The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite, with about 80% of the world's known amber found there. It dates from 44 million years ago...

        )
  • Maorothiini
  • Othiini C.G.Thomson, 1859
  • Platyprosopini Lynch, 1884
  • Staphylinini Latreille, 1802 (including Philonthini, Tanygnathinini)
    • Cafius
    • Creophilus
      • Creophilus maxillosus
        Creophilus maxillosus
        Creophilus maxillosus belongs to the order Coleoptera and is most commonly referred to as the hairy rove beetle. The rove beetles are a large family, having almost 2,900 species in North America. This species can be found in woods and wherever carrion is found, usually from the spring to autumn...

        – hairy rove beetle
    • Eucibdelus
    • Gabrius
    • Jurecekia
    • Ocypus
      • Ocypus olens – devil's coach-horse beetle
      • Ocypus ophthalmicus
        Ocypus ophthalmicus
        Ocypus ophthalmicus is a species of rove beetle belonging to the family Staphylinidae, subfamily Staphylininae.These beetles are present in most of Europe, in the East Palearctic ecozone, in North Africa and in the Near East....

    • Ontholestes
    • Paederomimus
    • Philonthus
      • Philonthus cognatus
        Philonthus cognatus
        Philonthus cognatus is a rove beetle. Adults are 8 to 10 mm long. They are mainly black in colour, although the elytra have a metallic sheen, and are profusely pitted. A distinctive feature of this species is that the underside of the first antennal segment is yellow, contrasting with the...

    • Quedius
      • Quedius curtipennis
        Quedius curtipennis
        Quedius curtipennis is a beetle found in Britain. It is similar in appearance to Quedius fuliginosus, which is a less common species, however, the eyes of fuliginosus are more convex and the basal antennal segments darker. The punctures on the elytra are slightly stronger in Q. curtipennis....

      • Quedius fuliginosus
        Quedius fuliginosus
        Quedius fuliginosus is a beetle found in Britain. It is similar in appearance to Quedius curtipennis, which is a more common species, however, the eyes of fuliginosus are more convex and the basal antennal segments darker. The punctures on the elytra are slightly stronger in curtipennis....

    • Rhynchocheilus
    • Staphylinus
      Staphylinus
      Staphylinus is a genus of rove beetles.-Selected species from Europe:*S. agarici*S. alatus*S. anceps*S. angustatus*S. arnicae*S. biclavatus*S. bicolor*S. brevipes*S. caesareus*S. cantharellus...

    • Stevensia
    • Thinopinuspictured rove beetle
      Pictured rove beetle
      The pictured rove beetle is a wingless rove beetle which lives on the sandy beaches of the West Coast of the United States from southern Alaska to Baja California. They are nocturnal, and emerge at night from temporary sand burrows to feed on beach hoppers .-Identification:Like other rove beetles, T...

    • Velleius
      • Velleius dilatatus
        Velleius dilatatus
        The rove beetle Velleius dilatatus lives together with the European hornet Vespa crabro crabro.Velleius dilatatus is up to 26 mm long. It feeds on detritus produced by the hornets . The beetles cannot survive without the hornets themselves, even if there is enough detritus present.It finds...

  • Xantholinini Erichson, 1839
    • Leptacinus
      • Leptacinus intermedius
        Leptacinus intermedius
        Leptacinus intermedius is a European rove beetle, described in 1935 by Horace Donisthorpe. Like most staphylinids, L. intermedius and its larvae are predatory upon other insects. Members of this family can usually be identified by their very short elytra, which leave three to six abdominal segments...

    • Xantholinus
      Xantholinus
      Xantholinus is a genus of the beetle family Staphylinidae.-Species:* Xantholinus appenninicola * Xantholinus baeticus * Xantholinus balearicus * Xantholinus bonomettoi...

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