Dvinosaurus
Encyclopedia
Dvinosaurus is a genus of extinct temnospondyl amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

 from the Late Permian of Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

. The first three species, D. primus (the type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

), D. secundus, and D. tertius, were described by Russian paleontologist Vladimir Prokhorovich Amalitskii
Vladimir Prokhorovich Amalitskii
Vladimir Prokhorovich Amalitskii was a Russian Paleontologist and Professor at Warsaw University, who was involved in the discovery and excavation of the famous Late Permian fossil vertebrate fauna from the North Dvina River, Arkhangelsk District, Northern European Russia.He died in December 1917...

 in 1921. Dvinosaurus was first found near the village of Sokolki in Arkhangelsk
Arkhangelsk Oblast
Arkhangelsk Oblast is a federal subject of Russia . It includes the Arctic archipelagos of Franz Josef Land and Novaya Zemlya, as well as the Solovetsky Islands in the White Sea....

 and named after the Northern Dvina River, which was close to the locality where it was discovered. The holotype
Holotype
A holotype is a single physical example of an organism, known to have been used when the species was formally described. It is either the single such physical example or one of several such, but explicitly designated as the holotype...

 is cataloged as PIN2005/35, and consists of only a skull. Amalitskii's D. tertius is now considered synonymous with D. secundus. Two additional species, D. egregius, and D. purlensis, were named by Mikhail Shishkin
Mikhail Shishkin
Mikhail Pavlovich Shishkin is a Russian writer. He is widely considered as one of the best contemporary Russian writers and praised for depth and complexity of his books and for his perfect command of Russian literary language.-Biography:...

in 1968. These were named from a fossilized skull and a partial lower jaw, respectively.

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