Worker bee
Encyclopedia
A Worker bee is any female eusocial bee that lacks the full reproductive capacity of the colony's queen bee
Queen bee
The term queen bee is typically used to refer to an adult, mated female that lives in a honey bee colony or hive; she is usually the mother of most, if not all, the bees in the hive. The queens are developed from larvae selected by worker bees and specially fed in order to become sexually mature...

; under most circumstances, this is correlated to an increase in certain non-reproductive activities relative to a queen, as well. Worker bees occur in many bee species other than honey bee
Honey bee
Honey bees are a subset of bees in the genus Apis, primarily distinguished by the production and storage of honey and the construction of perennial, colonial nests out of wax. Honey bees are the only extant members of the tribe Apini, all in the genus Apis...

s, but this is by far the most familiar colloquial use of the term.

Honey bee workers keep the hive temperature uniform in the critical brood area (where new bees are raised). Workers gather pollen into the pollen basket
Pollen basket
The pollen basket or corbicula is part of the tibia on the hind legs of the four related lineages of apid bees that used to comprise the family Apidae: the honey bees, bumblebees, stingless bees, and orchid bees...

s on their back legs, to carry back to the hive where it is used as food for the developing brood
Brood (honeybee)
In entomology, the term brood is used to refer to the embryo or egg, the larva and the pupa stages in the life of holometabolous insects. The brood of honey bees develops within a bee hive. In man-made, removable frame hives, such as Langstroth hives, each frame which is mainly brood is called a...

. Pollen carried on their bodies may be carried to another flower where a small portion can rub off onto the pistil, resulting in cross pollination
Pollination
Pollination is the process by which pollen is transferred in plants, thereby enabling fertilisation and sexual reproduction. Pollen grains transport the male gametes to where the female gamete are contained within the carpel; in gymnosperms the pollen is directly applied to the ovule itself...

. Almost all of civilization's food supply (maize
Maize
Maize known in many English-speaking countries as corn or mielie/mealie, is a grain domesticated by indigenous peoples in Mesoamerica in prehistoric times. The leafy stalk produces ears which contain seeds called kernels. Though technically a grain, maize kernels are used in cooking as a vegetable...

 is a noteworthy exception) depends greatly on crop pollination by honey bees, whether directly eaten or used as forage crops for animals that produce milk and meat. Nectar is sucked up through the proboscis, mixed with enzymes in the stomach, and carried back to the hive, where it is stored in wax cells and evaporated into honey
Honey
Honey is a sweet food made by bees using nectar from flowers. The variety produced by honey bees is the one most commonly referred to and is the type of honey collected by beekeepers and consumed by humans...

. Workers must maintain the hive's brood chamber at 34.4 degrees C to incubate the eggs. If it is too hot, they collect water and deposit it around the hive, then fan air through with their wings causing cooling by evaporation. If it is too cold, they cluster together to generate body heat. The life of all honey bees starts as an egg, which is laid by the queen in the bottom of a wax cell in the brood area of a hive. A worker egg hatches after three days into a larva. Nurse bees feed it royal jelly
Royal jelly
Royal jelly is a honey bee secretion that is used in the nutrition of larvae, as well as adult queens. It is secreted from the glands in the hypopharynx of worker bees, and fed to all larvae in the colony....

 at first, then pollen and honey for six days. It then becomes an inactive pupa.
Honeycombs have hexagonal cells on both sides of a vertical central wall. As shown in the photo, these cells are inclined upward, primarily to retain liquid nectar and honey. During its 14 days as a pupa, sealed in a capped cell, it grows into a worker (female) bee, emerging on the 20th day. In most species of honey bees, workers do everything but lay eggs and mate, though Cape honey bee workers can lay eggs. They build the comb from wax extruded from glands under their abdomen. They clean, defend, and repair the hive. They feed the larva, the queen, and the drones. They gather nectar, pollen, water, and propolis. They ventilate, cool and heat the hive.

When a colony absconds (all bees leave the colony) or divides and so creates a swarm and then establishes a new colony, the bees must regress in their behavior in order to establish the first generation in the new home. The most urgent task will be the creation of new beeswax
Beeswax
Beeswax is a natural wax produced in the bee hive of honey bees of the genus Apis. It is mainly esters of fatty acids and various long chain alcohols...

 for comb. Beekeepers take advantage of this by introducing swarms into new or existing colonies where they will draw comb. Comb is much more difficult to come by than honey and requires about six times the energy to create. A newly hived swarm on bar bars (top bar hive) or empty foundation (Langstroth box hive
Langstroth hive
The Langstroth bee hive, patented in October 1852, is the standard beehive used in many parts of the world for beekeeping. The advantage of this hive is that the bees build honeycomb into frames, which can be moved with ease. The frames are designed to prevent bees from attaching honeycombs where...

) will often be fed sugar water, which they can then rapidly consume to create wax for new comb (Mature hives cannot be so fed as they will store it in place of nectar, although a wintering hive may have to be fed if insufficient honey was left by the beekeeper.)

Cell cleaning (Day 1-2)

Brood cells must be cleaned before the next use - cells will be inspected by the queen and if unsatisfactory will not be used. Worker bees in the cleaning phase will perform this cleaning.
If the cells are not clean, the worker bee must do it again.

Nurse bee (Day 3-11)

Nurse bees feed the worker larvae worker jelly which is secreted from glands that produce royal jelly.
  • Advanced Nurse Bees (Day 6-11)
    • Feed royal jelly
      Royal jelly
      Royal jelly is a honey bee secretion that is used in the nutrition of larvae, as well as adult queens. It is secreted from the glands in the hypopharynx of worker bees, and fed to all larvae in the colony....

       to the queen larva and drones receive worker jelly for 1 to 3 days at which time they are started on a diet of honey and pollen.

Wax production (Day 12-17)

Wax Bees - build cells from wax, repair old cells, and store nectar and pollen
brought in by other workers. Early in the worker's career she will exude wax from the space between several of her abdominal segments. Four sets of wax glands, situated inside the last four ventral segments of the abdomen, produce wax for comb construction.

Honey sealing

Mature honey
Honey
Honey is a sweet food made by bees using nectar from flowers. The variety produced by honey bees is the one most commonly referred to and is the type of honey collected by beekeepers and consumed by humans...

, sufficiently dried, is sealed tightly with wax to prevent absorption of moisture from the air by workers deputized to do same.

Queen attendants

The attendants groom and feed the queen. They also collect QMP (Queen Mandibular Pheromone
Queen mandibular pheromone
Queen Mandibular Pheromone, or QMP, is a honey bee pheromone produced by the queen and fed to her attendants who share it with the rest of the colony that gives the colony the sense of being queenright. Chemically QMP is very diverse with at least 17 major components and other minor ones...

) from the queen and share it with the bees around them who also share it spreading its effects through the hive.

Honeycomb building

Workers will take wax from wax producing workers and build the comb with it.

Pollen packing

Pollen brought into the hive for feeding the brood is also stored. It must be packed firmly into comb cells and mixed with a small amount of honey so that it will not spoil. Unlike honey, which does not support bacterial life, stored pollen will become rancid
Rancidification
Rancidification is the chemical decomposition of fats, oils and other lipids . When these processes occur in food, undesirable odors and flavors can result. In some cases, however, the flavors can be desirable . In processed meats, these flavors are collectively known as "warmed over flavor"...

 without proper care. It has to be kept in honey cells.

Propolizing

The walls of the hive are covered with a thin coating of propolis
Propolis
Propolis is a resinous mixture that honey bees collect from tree buds, sap flows, or other botanical sources. It is used as a sealant for unwanted open spaces in the hive. Propolis is used for small gaps , while larger spaces are usually filled with beeswax. Its color varies depending on its...

, a resinous substance obtained from plants. In combination with enzymes added by the worker this has antibacterial and antifungal properties.

Propolis is used to aide with ventilation and at the entrances of hives.

Mortuary bees

Dead bees and failed larvae must be removed from the hive to prevent disease and allow cells to be reused. They will be carried some distance from the hive by mortuary bees.

Fanning bees

Worker bees fan the hive, cooling it with evaporated water brought by water carriers. They direct airflow into the hive or out of the hive depending on need.

Water carriers

When the hive is in danger of overheating, these bees will obtain water, usually from within a short distance from the hive and bring it back to spread on the backs of fanning bees. The worker bee has a crop
Crop (anatomy)
A crop is a thin-walled expanded portion of the alimentary tract used for the storage of food prior to digestion that is found in many animals, including gastropods, earthworms, leeches, insects, birds, and even some dinosaurs.- Bees :Cropping is used by bees to temporarily store nectar of flowers...

 separate from the nectar crop for this purpose.

Foraging bees (Days 22 - 42)

The forager and scout bees travel (up to 1.5 miles) to a nectar source
Nectar source
A nectar source is a flowering plant that produces nectar as part of its reproductive strategy. These plants create nectar, which attract pollinating insects and sometimes other animals such as birds....

, pollen source
Pollen source
The term pollen source is often used in the context of beekeeping and refers to flowering plants as a source of pollen for bees or other insects. Bees collect pollen as a protein source to raise their brood. For the plant, the pollinizer, this can be an important mechanism for sexual reproduction,...

 or to collect propolis
Propolis
Propolis is a resinous mixture that honey bees collect from tree buds, sap flows, or other botanical sources. It is used as a sealant for unwanted open spaces in the hive. Propolis is used for small gaps , while larger spaces are usually filled with beeswax. Its color varies depending on its...

.

Genetic characteristics

In most common bee species, worker bees are infertile and thus never reproduce. They are nevertheless considered female for anatomical and genetic reasons. Genetically, a worker bee does not differ from a queen bee and can even become a laying worker bee
Laying worker bee
A laying worker bee is a worker bee that lays unfertilized eggs usually in the absence of a queen bee. Only drones develop from the eggs of laying worker bees . A beehive cannot survive with only a laying worker bee....

, but in most species will produce only male (drone) offspring. Whether a larva becomes a worker or a queen depends on the kind of food it is given after the first three days of its larval form.

The stinger

The worker bee's stinger is a complex organ that allows a bee to defend itself and the hive from most mammals. Attacking bees aim for the face by sensing regions with high levels of carbon dioxide (like mosquito
Mosquito
Mosquitoes are members of a family of nematocerid flies: the Culicidae . The word Mosquito is from the Spanish and Portuguese for little fly...

s.) Bee sting
Bee sting
A bee sting is strictly a sting from a bee . In the vernacular it can mean a sting of a bee, wasp, hornet, or yellow jacket. Some people may even call the bite of a horse-fly a bee sting...

s against mammals and birds typically leave the stinger embedded in victim due to the structure of flesh and the stinger's barbs. In this case, the venom bulb stays with the stinger and continues to pump. Upon losing its stinger, the bee will subsequently die since the portion where the stinger bulb was removed draws out part of its insides.

The barbs on the stinger will not catch on most animals besides mammals and birds. This means that such animals can be stung many times by the same bee.

Stingless bees

There are many bees in this group, native to all continents except for Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 and Antarctica, that have workers which do not have stingers. These bees are not defenseless, however, as they can bite with their mandibles, occasionally releasing caustic secretions at the same time, similar to the defenses of some ant
Ant
Ants are social insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. More than...

s.
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