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Winter Wren
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The Winter Wren (Troglodytes troglodytes), also known as the Northern Wren, is a very small bird, a member of the mainly New World wren family Troglodytidae. It is the only one of nearly sixty species in the family which occurs in the Old World; in Europe it is commonly known simply as the Wren. They are noteworthy among songbirds both because of their amazingly long and complex songs and because they are one of the few passerine species that has a distribution spanning both North America and Eurasia.
It occurs in Europe, a belt of Asia from northern Iran and Afghanistan across to Japan, and North America. It is only migratory in the northern parts of its range.
The scientific name is taken from the Greek word "troglodytes" (from "trogle" a hole, and "dyein" to creep), meaning "cave-dweller", and refers to its habit of disappearing into cavities or crevices whilst hunting arthropods or to roost. Taxonomy of the genus Troglodytes is currently unresolved, as recent molecular studies have suggested that Cistothorus spp. and Thryorchilus spp. are within the clade currently defined by Troglodytes . A study of individuals in an overlap zone between two divergent subspecies of winter wren in North America (Troglodytes troglodytes hiemalis in Eastern North America and Troglodytes troglodytes pacificus in Western North America), which differ substantially in song and genetics, found strong evidence for reproductive isolation and suggested that the pacificus subspecies be promoted to the species level designation of Troglodytes pacificus .
DescriptionThe 9–10.5 cm long Wren is rufous brown above, greyer beneath, barred with darker brown and grey, even on wings and tail. The bill is dark brown, the legs pale brown. Young birds are less distinctly barred.
The plumage is subject to considerable variation, and where populations have been isolated, the variation has become fixed in one minor form or another. There are around 27 Eurasian subspecies of this taxonomically complex bird, with around 12 more in North America. The disputed subspecies orii, the Daito Winter Wren, became extinct around 1940 – if it is indeed a valid taxon and not merely based on an anomaly.
Thus in the British Isles, in addition to the typical bird, T. t. indigenus, there are two distinct insular forms; one, T. t. hirtensis, is confined to the island of St Kilda, and another, T. t. zetlandicus to Shetland. The St. Kilda Wren is greyer above, whiter beneath, and with more abundant bars on the back; the Shetland Wren is darker.
TaxonomyRecent taxonomies of the Troglodytes genus have shown that winter wrens are the most distantly related of all species within Troglodytes, and have shown that two other groups, the timberline wren (Thryorchilus browni) and the four species within the genus Cistothorus, are within the clade defined by all of the Troglodytes. To make Troglodytes monophyletic, Rice et al. (1999) proposed that winter wrens could be placed in their own genus, Nannus. Alternatively, as suggested by Gómez et al. (2005) , Troglodytes could be made more inclusive by assigning the Troglodytes genus to the current Thryorchilus and Cistothorus genera.
By studying the songs and genetics of individuals in an overlap zone between two subspecies of winter wren within North America (Troglodytes troglodytes hiemalis in Eastern North America and Troglodytes troglodytes pacificus in Western North America), a study by Toews and Irwin (2008) found strong evidence of reproductive isolation between the two. It was suggested that the pacificus subspecies be promoted to the species level designation of Troglodytes pacificus with the common name of ‘Pacific wren’. By applying a molecular clock to the amount of mitochondrial DNA sequence divergence between the two , it was estimated that Troglodytes pacificus and Troglodytes troglodytes last shared a common ancestor approximately 4.3 million years ago, long before the glacial cycles of the Pleistocene, thought to have promoted speciation in many avian systems inhabiting the boreal forest of North America .
Breeding The male Wren builds several nests, up to 6 or 7 in Europe, but fewer in North America. These are called "cock nests" but are never lined until the female chooses one to use.
The normal round nest of grass, moss, lichens or leaves is tucked into a hole in a wall, tree trunk, crack in a rock or corner of a building, but it is often built in bushes, overhanging boughs or the litter which accumulates in branches washed by floods.
Five to eight white or slightly speckled eggs are laid in April, and second broods are reared. The eggs of the St. Kilda Wren are marginally larger and often more boldly spotted; six is the usual number.
In popular culture
External links- (for 19 issues) with Circum-Polar Range-Map
- on the Internet Bird Collection
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