Vitis
Encyclopedia
Vitis is a genus of about 60 species of vining plants in the flowering plant
Flowering plant
The flowering plants , also known as Angiospermae or Magnoliophyta, are the most diverse group of land plants. Angiosperms are seed-producing plants like the gymnosperms and can be distinguished from the gymnosperms by a series of synapomorphies...

 family Vitaceae
Vitaceae
Vitaceae are a family of dicotyledonous flowering plants including the grapevine and Virginia creeper. The family name is derived from the genus Vitis...

. The genus is made up of species predominantly from the Northern hemisphere. It is economically important as the source of grape
Grape
A grape is a non-climacteric fruit, specifically a berry, that grows on the perennial and deciduous woody vines of the genus Vitis. Grapes can be eaten raw or they can be used for making jam, juice, jelly, vinegar, wine, grape seed extracts, raisins, molasses and grape seed oil. Grapes are also...

s, both for direct consumption of the fruit and for fermentation to produce wine
Wine
Wine is an alcoholic beverage, made of fermented fruit juice, usually from grapes. The natural chemical balance of grapes lets them ferment without the addition of sugars, acids, enzymes, or other nutrients. Grape wine is produced by fermenting crushed grapes using various types of yeast. Yeast...

. The study and cultivation of grapevines is called viticulture
Viticulture
Viticulture is the science, production and study of grapes which deals with the series of events that occur in the vineyard. When the grapes are used for winemaking, it is also known as viniculture...

.

Biology

Vitis is distinguished from other genera of Vitaceae by having petals which remain joined at the tip and detach from the base to fall together as a calyptra or 'cap'. The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are unisexual or modified to act functionally as unisexual, they are pentamerous with a hypogynous disk. The calyx is greatly reduced or nonexistent in most species and the petals are joined at the summit into one unit but separated at the base. Flower buds are formed later in the growing season and overwinter for blooming in spring of the next year. There are two types of flowers produced, sterile flowers with five long filaments and erect stamens with undeveloped pistils and fertile flowers with well-developed pistils and that have five undeveloped reflexed stamens. The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is a berry
Berry
The botanical definition of a berry is a fleshy fruit produced from a single ovary. Grapes are an example. The berry is the most common type of fleshy fruit in which the entire ovary wall ripens into an edible pericarp. They may have one or more carpels with a thin covering and fleshy interiors....

, normally produced with four or less per flower by way of aborted embryos, ovoid in shape and juicy.

In the wild, all species of Vitis are normally dioecious
Plant sexuality
Plant sexuality covers the wide variety of sexual reproduction systems found across the plant kingdom. This article describes morphological aspects of sexual reproduction of plants....

, but under domestication, variants with perfect flowers appear to have been selected.

Most Vitis species have 38 chromosomes (n=19), but 40 (n=20) in subgenus Muscadinia. In that respect the Muscadinia are the same as other Vitaceae such as Ampelocissus
Ampelocissus
Ampelocissus is a genus of Vitaceae having 90 or more species found variously in tropical Africa, Asia, Central America, and Oceania. The type species, A...

, Parthenocissus
Parthenocissus
Parthenocissus , creepers, is a genus of climbing plants from the grape family, Vitaceae. It contains about 12 species, from Asia and North America. Several are grown for ornamental use....

, and Ampelopsis
Ampelopsis
Ampelopsis is a genus of climbing shrubs, in the grape family Vitaceae. The name is derived from the Greek word ampelos, which means "vine"...

.

Uses

The fruit of several Vitis species are grown commercially for consumption as fresh grape
Grape
A grape is a non-climacteric fruit, specifically a berry, that grows on the perennial and deciduous woody vines of the genus Vitis. Grapes can be eaten raw or they can be used for making jam, juice, jelly, vinegar, wine, grape seed extracts, raisins, molasses and grape seed oil. Grapes are also...

s and for fermentation into wine
Wine
Wine is an alcoholic beverage, made of fermented fruit juice, usually from grapes. The natural chemical balance of grapes lets them ferment without the addition of sugars, acids, enzymes, or other nutrients. Grape wine is produced by fermenting crushed grapes using various types of yeast. Yeast...

. Vitis vinifera
Vitis vinifera
Vitis vinifera is a species of Vitis, native to the Mediterranean region, central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran....

 is the most important species.

The leaves of the grape vine
Grape leaves
Grape leaves are used in the cuisines of a number of cultures, including Turkish cuisine, Greek cuisine, Bulgarian cuisine, Arab cuisine, Romanian cuisine, and Vietnamese cuisine. They are most often picked fresh from the vine and stuffed with a mixture of rice, meat, and spices, and then cooked by...

 itself are considered edible and are used in the production of dolmades and Vietnamese lot leaves.

Species

Most Vitis species are found in the temperate regions of the Northern Hemisphere in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 and Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 with a few in the tropics. The wine grape Vitis vinifera originated in southern Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 and southwestern Asia. The species occur in widely different geographical areas and show a great diversity of form. However they are sufficiently closely related to allow easy interbreeding and the resultant interspecific hybrids are invariably fertile and vigorous. Thus the concept of a species is less well defined and more likely represents the identification of different ecotypes of Vitis that have evolved in distinct geographical and environmental circumstances.

The exact number of species is not certain, with species in Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 in particular being poorly defined. Estimates range from 40 to more than 60. Some of the more notable include:
  • Vitis vinifera
    Vitis vinifera
    Vitis vinifera is a species of Vitis, native to the Mediterranean region, central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran....

    , the European grapevine. Native to the Mediterranean and Central Asia.
  • Vitis labrusca
    Vitis labrusca
    Vitis labrusca is a species of grapevines belonging to the Vitis genus in the flowering plant family Vitaceae. The vines are native to the eastern United States and are the source of many grape cultivars, including Catawba and Concord grapes, and many hybrid grape varieties such as Agawam,...

    , the Fox grapevine, sometimes used for wine. Native to the Eastern United States
    Eastern United States
    The Eastern United States, the American East, or simply the East is traditionally defined as the states east of the Mississippi River. The first two tiers of states west of the Mississippi have traditionally been considered part of the West, but can be included in the East today; usually in...

     and Canada
    Canada
    Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

    .
  • Vitis riparia
    Vitis riparia
    Vitis riparia Michx, also commonly known as River Bank Grape or Frost Grape, is a native American climbing or trailing vine, widely distributed from Quebec to Texas, and Montana to New England. It is long-lived and capable of reaching into the upper canopy of the tallest trees...

    , the Riverbank Grapevine, sometimes used for winemaking and for jam. Native to the entire Eastern U.S. and north to Quebec
    Quebec
    Quebec or is a province in east-central Canada. It is the only Canadian province with a predominantly French-speaking population and the only one whose sole official language is French at the provincial level....

    .
  • Vitis aestivalis
    Vitis aestivalis
    Vitis aestivalis is a species of grape native to eastern North America from southern Ontario east to Vermont, west to Oklahoma, and south to Florida and Texas. It is a vigorous vine, growing to 10 m or more high in trees...

    , the Summer Grape, native to the Eastern United States
    Eastern United States
    The Eastern United States, the American East, or simply the East is traditionally defined as the states east of the Mississippi River. The first two tiers of states west of the Mississippi have traditionally been considered part of the West, but can be included in the East today; usually in...

    , especially the Southeastern United States
    Southeastern United States
    The Southeastern United States, colloquially referred to as the Southeast, is the eastern portion of the Southern United States. It is one of the most populous regions in the United States of America....

    .
  • Vitis rotundifolia (syn. Muscadinia rotundifolia), the Muscadine
    Muscadine
    Muscadines are a grapevine species native to the present-day southeastern United States that has been extensively cultivated since the 16th Century. It differs from Vitis spp. in its number of chromosomes and its morphology...

    , used for jams and wine. Native to the Southeastern United States
    Southeastern United States
    The Southeastern United States, colloquially referred to as the Southeast, is the eastern portion of the Southern United States. It is one of the most populous regions in the United States of America....

     from Delaware
    Delaware
    Delaware is a U.S. state located on the Atlantic Coast in the Mid-Atlantic region of the United States. It is bordered to the south and west by Maryland, and to the north by Pennsylvania...

     to the Gulf of Mexico
    Gulf of Mexico
    The Gulf of Mexico is a partially landlocked ocean basin largely surrounded by the North American continent and the island of Cuba. It is bounded on the northeast, north and northwest by the Gulf Coast of the United States, on the southwest and south by Mexico, and on the southeast by Cuba. In...

    .
  • Vitis coignetiae
    Vitis coignetiae
    Vitis coignetiae is a plant belonging to the genus Vitis that is native to the temperate climes of asia, where it can be found in Russian Federation ; Korea; and Japan...

     Crimson Glory Vine, a species from East Asia grown as an ornamental plant
    Ornamental plant
    Ornamental plants are plants that are grown for decorative purposes in gardens and landscape design projects, as house plants, for cut flowers and specimen display...

     for its crimson autumn foliage.
  • Vitis amurensis
    Vitis amurensis
    Vitis amurensis, the Amur grape, is a species of grape native to the Asian continent. Its name comes from the Amur Valley in Russia and China.It is very resistant to frost, but is not tolerant to drought...

    , native to the Asian continent, including parts of Siberia and China.
  • Vitis vulpina
    Vitis vulpina
    Vitis vulpina is a herbaceous perennial vine of the family Vitaceae . The genus name Vitis comes from the Latin word for "vine" and the species name vulpina comes from the Latin word for "fox-like" or belonging to a fox...

    , the Frost Grape, native to the Eastern United States
    Eastern United States
    The Eastern United States, the American East, or simply the East is traditionally defined as the states east of the Mississippi River. The first two tiers of states west of the Mississippi have traditionally been considered part of the West, but can be included in the East today; usually in...

    , from Massachusetts to Florida, and west to Nebraska, Kansas, and Texas. Treated by some as a synonym
    Synonym (taxonomy)
    In scientific nomenclature, a synonym is a scientific name that is or was used for a taxon of organisms that also goes by a different scientific name. For example, Linnaeus was the first to give a scientific name to the Norway spruce, which he called Pinus abies...

     of V. riparia.


There are many cultivars of grapevines; most are cultivars of V. vinifera.

Hybrid grapes also exist, and these are primarily crosses between V. vinifera and one or more of V. labrusca, V. riparia or V. aestivalis. Hybrids tend to be less susceptible to frost and disease (notably phylloxera
Phylloxera
Grape phylloxera ; originally described in France as Phylloxera vastatrix; equated to the previously described Daktulosphaira vitifoliae, Phylloxera vitifoliae; commonly just called phylloxera is a pest of commercial grapevines worldwide, originally native to eastern North America...

), but wine from some hybrids may have a little of the characteristic "foxy" taste of V. labrusca.

Commercial distribution

According to the "Food and Agriculture Organization" (FAO), 75,866 square kilometres of the world is dedicated to grapes. Approximately 71% of world grape production is used for wine, 27% as fresh fruit, and 2% as dried fruit. A portion of grape production goes to producing grape juice to be used as a sweetener for fruits canned "with no added sugar" and "100% natural". The area dedicated to vineyards is increasing by about 2% per year.

The following list of top wine-producers shows the corresponding areas dedicated to grapes for wine making:
  • Spain 11,750 km²
  • France 8,640 km²
  • Italy 8,270 km²
  • Turkey 8,120 km²
  • United States 4,150 km²
  • Iran 2,860 km²
  • Romania 2,480 km²
  • Portugal 2,160 km²
  • Argentina 2,080 km²
  • Australia 1,642 km²
  • Lebanon 1,221 km²


Sources: FAO, Organisation Internationale de la Vigne et du Vin (pdf), Australian Wine and Brandy Corporation.

Pests and diseases

Phylloxera
Phylloxera
Grape phylloxera ; originally described in France as Phylloxera vastatrix; equated to the previously described Daktulosphaira vitifoliae, Phylloxera vitifoliae; commonly just called phylloxera is a pest of commercial grapevines worldwide, originally native to eastern North America...

 is an American root aphid that devastated V. vinifera vineyards in Europe when accidentally introduced in the late 19th century. Attempts were made to breed in resistance from American species, but many winemakers didn't like the unusual flavour profile of the hybrid vines. Fortunately, V. vinifera grafts readily onto rootstocks of the American species, and most commercial production of grapes now relies on such grafts.

The Black vine weevil
Black vine weevil
Otiorhynchus sulcatus, commonly known as the black vine weevil, is native to Europe, but common in North America as well. It is a pest of many garden plants.-Overview:...

 is another root pest.

Grapevines are used as food plants by the larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e of some Lepidoptera
Lepidoptera
Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

 species - see list of Lepidoptera that feed on grapevines.

Symbolism

The grape vine (typically Vitis vinifera
Vitis vinifera
Vitis vinifera is a species of Vitis, native to the Mediterranean region, central Europe, and southwestern Asia, from Morocco and Portugal north to southern Germany and east to northern Iran....

) has been used as a symbol since ancient times. In Greek mythology
Greek mythology
Greek mythology is the body of myths and legends belonging to the ancient Greeks, concerning their gods and heroes, the nature of the world, and the origins and significance of their own cult and ritual practices. They were a part of religion in ancient Greece...

, Dionysius
Dionysius
The Graeco-Roman name Dionysius, deriving from the name of the Thracian god Dionysus, was exceedingly common, and many ancient people, famous and otherwise, bore it. It remains a common name today in the form Dennis . The modern Greek form of the name is Dionysios or Dionysis. The Spanish form of...

 (and with the Romans Bacchus
Bacchus
Bacchus is the Roman name for Dionysus, the god of wine and intoxication.Bacchus can also refer to:* Temple of Bacchus, a Roman temple at a large classical antiquity complex in Baalbek, Lebanon...

) was god of the vintage and, therefore, a grape vine with bunches of the fruit are among their attributes. Their attendants on the Bacchanalian festivals — the Bacchanals — hence had the vine as an attribute, together with the thyrsus
Thyrsus
In Greek mythology, a thyrsus or thyrsos was a staff of giant fennel covered with ivy vines and leaves, sometimes wound with taeniae and always topped with a pine cone. These staffs were carried by Dionysus and his followers. Euripides wrote that honey dripped from the thyrsos staves that the...

, the latter often entwined with vine branches. For the same reason, the Greek wine cup (cantharos) is commonly decorated with the vine and grapes, wine being drunk as a libation
Libation
A libation is a ritual pouring of a liquid as an offering to a god or spirit or in memory of those who have died. It was common in many religions of antiquity and continues to be offered in various cultures today....

 to the god.

In Christian
Christian
A Christian is a person who adheres to Christianity, an Abrahamic, monotheistic religion based on the life and teachings of Jesus of Nazareth as recorded in the Canonical gospels and the letters of the New Testament...

 iconography
Iconography
Iconography is the branch of art history which studies the identification, description, and the interpretation of the content of images. The word iconography literally means "image writing", and comes from the Greek "image" and "to write". A secondary meaning is the painting of icons in the...

, the vine also frequently appears. It is mentioned several times in the New Testament
New Testament
The New Testament is the second major division of the Christian biblical canon, the first such division being the much longer Old Testament....

. We have the parable of the kingdom of heaven likened to the father starting to engage laborers for his vineyard. The vine is used as symbol of Jesus Christ based on his own statement, “I am the vine.” In that sense, a vine is placed as sole symbol on the tomb of Constantia
Flavia Julia Constantia
Flavia Julia Constantia was the daughter of the Roman Emperor Constantius Chlorus and his second wife, Flavia Maximiana Theodora....

, the sister of Constantine the Great, and elsewhere. In Byzantine art
Byzantine art
Byzantine art is the term commonly used to describe the artistic products of the Byzantine Empire from about the 5th century until the Fall of Constantinople in 1453....

, the vine and grapes figure in early mosaic
Mosaic
Mosaic is the art of creating images with an assemblage of small pieces of colored glass, stone, or other materials. It may be a technique of decorative art, an aspect of interior decoration, or of cultural and spiritual significance as in a cathedral...

s, and on the throne of Maximianus of Ravenna
Maximianus of Ravenna
Maximianus of Ravenna, or Maximian was bishop of Ravenna.Born in the Istrian city of Pula , Maximianus was consecrated bishop of Ravenna in 546 by Pope Vigilius in Patras, Greece. Maximianus was a forty-eight year old deacon from Pola when he became the twenty-sixth bishop of Ravenna...

 it is used as a decoration.

The vine as symbol of the chosen people
Chosen people
Throughout history and even today various groups of people have considered themselves as chosen by a deity for some purpose such as to act as the deity's agent on earth. In monotheistic faiths, like Abrahamic religions, references to God are used in constructs such as "God's Chosen People"...

 is employed several times in the Old Testament
Old Testament
The Old Testament, of which Christians hold different views, is a Christian term for the religious writings of ancient Israel held sacred and inspired by Christians which overlaps with the 24-book canon of the Masoretic Text of Judaism...

. The vine and wheat
Wheat
Wheat is a cereal grain, originally from the Levant region of the Near East, but now cultivated worldwide. In 2007 world production of wheat was 607 million tons, making it the third most-produced cereal after maize and rice...

 ear
Ear (botany)
An ear is the grain-bearing tip part of the stem of a cereal plant, such as wheat or maize. It can also refer to "a prominent lobe in some leaves".The ear is a spike, consisting of a central stem on which grows tightly packed rows of flowers...

 have been frequently used as symbol of the blood and flesh of Christ, hence figuring as symbols (bread and wine) of the Eucharist
Eucharist
The Eucharist , also called Holy Communion, the Sacrament of the Altar, the Blessed Sacrament, the Lord's Supper, and other names, is a Christian sacrament or ordinance...

 and are found depicted on ostensories. Often the symbolic vine laden with grapes is found in ecclesiastical decorations with animals biting at the grapes. At times, the vine is used as symbol of temporal blessing.

Art

In decorative art, while not as favored as many other motifs, the vine is frequently found in works of the Middle Ages
Middle Ages
The Middle Ages is a periodization of European history from the 5th century to the 15th century. The Middle Ages follows the fall of the Western Roman Empire in 476 and precedes the Early Modern Era. It is the middle period of a three-period division of Western history: Classic, Medieval and Modern...

, and in Renaissance
Renaissance
The Renaissance was a cultural movement that spanned roughly the 14th to the 17th century, beginning in Italy in the Late Middle Ages and later spreading to the rest of Europe. The term is also used more loosely to refer to the historical era, but since the changes of the Renaissance were not...

 works the vine ornaments frieze
Frieze
thumb|267px|Frieze of the [[Tower of the Winds]], AthensIn architecture the frieze is the wide central section part of an entablature and may be plain in the Ionic or Doric order, or decorated with bas-reliefs. Even when neither columns nor pilasters are expressed, on an astylar wall it lies upon...

s, pilaster
Pilaster
A pilaster is a slightly-projecting column built into or applied to the face of a wall. Most commonly flattened or rectangular in form, pilasters can also take a half-round form or the shape of any type of column, including tortile....

s, etc. In the art of the early 20th century, the grape bunch was prolifically used in the decorative arts.
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