Vipera
Encyclopedia
Common names: Palaearctic vipers, Eurasian vipers.

Vipera is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of venomous
Venomous snake
"Poisonous snake" redirects here. For true poisonous snakes, see Rhabdophis.Venomous snakes are snakes which have venom glands and specialized teeth for the injection of venom...

 viper
Viperinae
The Viperinae, or viperines, are a subfamily of venomous vipers found in Europe, Asia and Africa. They are distinguished by their lack of the heat-sensing pit organs that characterize their sister group, the Crotalinae. Currently, 12 genera and 66 species are recognized...

s. It has a very wide range, being found from North Africa
North Africa
North Africa or Northern Africa is the northernmost region of the African continent, linked by the Sahara to Sub-Saharan Africa. Geopolitically, the United Nations definition of Northern Africa includes eight countries or territories; Algeria, Egypt, Libya, Morocco, South Sudan, Sudan, Tunisia, and...

 to just within the Arctic Circle
Arctic Circle
The Arctic Circle is one of the five major circles of latitude that mark maps of the Earth. For Epoch 2011, it is the parallel of latitude that runs north of the Equator....

 and from Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

 to Pacific Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

. The name is possibly derived from the Latin words vivus and pario, meaning "alive" and "bear" or "bring forth"; likely a reference to the fact that most vipers bear live young. Currently, 23 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 are recognized.

Description

Members are usually small and more or less stoutly built. The head is distinct from the neck, of triangular shape, and covered with small scales in many species, although some have a few small plates on top. The dorsal scales
Dorsal scales
In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales.When counting dorsal scales, numbers are often given for three points along the body, for example 19:21:17...

 are strongly keeled
Keeled scales
Keeled scales refer to reptile scales that, rather than being smooth, have a ridge down the center that may or may not extend to the tip of the scale, making them rough to the touch...

, the anal scale
Anal scale
In snakes, the anal scale is the scale just in front of and covering the cloacal opening. This scale can be either single or paired . When paired, the division is oblique. It is preceded by the ventral scales and followed by the subcaudal scales....

 is divided and the subcaudals paired.

Geographic range

They can be found in Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

 and nearly all of continental Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, on some small islands of the Mediterranean (Elba
Elba
Elba is a Mediterranean island in Tuscany, Italy, from the coastal town of Piombino. The largest island of the Tuscan Archipelago, Elba is also part of the National Park of the Tuscan Archipelago and the third largest island in Italy after Sicily and Sardinia...

, Montecristo
Montecristo
Montecristo is a small Italian islet, approximately 4.3 kilometers across at its widest point, located about halfway between Corsica and mainland Italy; south of Elba and west of Giglio. It is part of the Tuscan Archipelago...

, Sicily
Sicily
Sicily is a region of Italy, and is the largest island in the Mediterranean Sea. Along with the surrounding minor islands, it constitutes an autonomous region of Italy, the Regione Autonoma Siciliana Sicily has a rich and unique culture, especially with regard to the arts, music, literature,...

) and the Aegean Sea
Aegean Sea
The Aegean Sea[p] is an elongated embayment of the Mediterranean Sea located between the southern Balkan and Anatolian peninsulas, i.e., between the mainlands of Greece and Turkey. In the north, it is connected to the Marmara Sea and Black Sea by the Dardanelles and Bosporus...

, as well as in northern Africa in Morocco
Morocco
Morocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...

, Algeria
Algeria
Algeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...

 and Tunisia
Tunisia
Tunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...

. It also occurs across the Arctic Circle
Arctic Circle
The Arctic Circle is one of the five major circles of latitude that mark maps of the Earth. For Epoch 2011, it is the parallel of latitude that runs north of the Equator....

 and eastwards though northern Asia to Sakhalin Island and northern Korea
Korea
Korea ) is an East Asian geographic region that is currently divided into two separate sovereign states — North Korea and South Korea. Located on the Korean Peninsula, Korea is bordered by the People's Republic of China to the northwest, Russia to the northeast, and is separated from Japan to the...

.

Habitat

Most species prefer cooler environments. Those found at lower latitudes tend to prefer higher altitudes and dryer, rocky habitats, while the species that occur at more northern latitudes prefer lower elevations and environments that have more vegetation and moisture.

Venom

Most Vipera species have venom that contains both neurotoxic
Neurotoxin
A neurotoxin is a toxin that acts specifically on nerve cells , usually by interacting with membrane proteins such as ion channels. Some sources are more general, and define the effect of neurotoxins as occurring at nerve tissue...

 and hemotoxic
Hemotoxin
Hemotoxins, haemotoxins or hematotoxins are toxins that destroy red blood cells , disrupt blood clotting, and/or cause organ degeneration and generalized tissue damage. The term hemotoxin is to some degree a misnomer since toxins that damage the blood also damage other tissues...

 components. Bites vary widely in severity. Smaller, northern species, such as V. berus
Vipera berus
Vipera berus, the common European adder or common European viper, is a venomous viper species that is extremely widespread and can be found throughout most of Western Europe and all the way to Far East Asia. Known by a host of common names including Common adder and Common viper, adders have been...

, have only slightly less toxic venom, but inject very little. Others, such as V. ammodytes
Vipera ammodytes
Vipera ammodytes is a venomous viper species found in southern Europe through to the Balkans and parts of the Middle East. It is reputed to be the most dangerous of the European vipers due to its large size, long fangs and high venom toxicity...

, are capable of injecting much more with devastating results. However, bites from Vipera species are rarely as severe as those from larger Macrovipera
Macrovipera
Macrovipera is a genus of venomous vipers that inhabit the semideserts and steppes of North Africa, the Near and Middle East, and the Milos Archipelago in the Aegean Sea. These snakes are responsible for a number of bites in Africa and western Asia every year...

 or Daboia
Daboia
Daboia is a monotypic genus of venomous Old World viper. The single species, D. russelii, is found in Asia throughout the Indian subcontinent, much of Southeast Asia, southern China and Taiwan...

.

Species

Species Taxon author Subsp.* Common name Geographic range
V. albicornuta
Vipera albicornuta
Vipera albicornuta is a venomous viper species endemic in Iran. No subspecies are currently recognized.-Description:Grows to a maximum length of 66 cm....

Nilson & Andrén, 1985 0 Iranian mountain viper The Zanjan
Zanjan
Zanjan may refer to:* Zanjan Province, Iran* Zanjan County, an area within Zanjan Province* Zanjan, Iran, the capital of Zanjan County and Zanjan Province* Zanjan University, located in the city of Zanjan* Senjan, a city in Markazi Province, Iran...

 Valley and surrounding mountains in northwestern Iran
Iran
Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...

.
V. albizona
Vipera albizona
Vipera albizona is a venomous viper species endemic to the mountainous regions of central Turkey. No subspecies are currently recognized.-Description:The maximum is a little less than 78 cm , although most specimens are smaller....

Nilson, Andrén & Flärdh, 1990 0 Central Turkish mountain viper Central Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

.
V. ammodytes
Vipera ammodytes
Vipera ammodytes is a venomous viper species found in southern Europe through to the Balkans and parts of the Middle East. It is reputed to be the most dangerous of the European vipers due to its large size, long fangs and high venom toxicity...

(Linnaeus
Carolus Linnaeus
Carl Linnaeus , also known after his ennoblement as , was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of the fathers of modern ecology...

, 1758)
4 Horned viper North-eastern Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

, southern Slovakia
Slovakia
The Slovak Republic is a landlocked state in Central Europe. It has a population of over five million and an area of about . Slovakia is bordered by the Czech Republic and Austria to the west, Poland to the north, Ukraine to the east and Hungary to the south...

, western Hungary
Hungary
Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...

, Slovenia
Slovenia
Slovenia , officially the Republic of Slovenia , is a country in Central and Southeastern Europe touching the Alps and bordering the Mediterranean. Slovenia borders Italy to the west, Croatia to the south and east, Hungary to the northeast, and Austria to the north, and also has a small portion of...

, Croatia
Croatia
Croatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...

, Bosnia-Herzegovina, Serbia
Serbia
Serbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...

, Montenegro
Montenegro
Montenegro Montenegrin: Crna Gora Црна Гора , meaning "Black Mountain") is a country located in Southeastern Europe. It has a coast on the Adriatic Sea to the south-west and is bordered by Croatia to the west, Bosnia and Herzegovina to the northwest, Serbia to the northeast and Albania to the...

, Albania
Albania
Albania , officially known as the Republic of Albania , is a country in Southeastern Europe, in the Balkans region. It is bordered by Montenegro to the northwest, Kosovo to the northeast, the Republic of Macedonia to the east and Greece to the south and southeast. It has a coast on the Adriatic Sea...

, Republic of Macedonia
Republic of Macedonia
Macedonia , officially the Republic of Macedonia , is a country located in the central Balkan peninsula in Southeast Europe. It is one of the successor states of the former Yugoslavia, from which it declared independence in 1991...

, Greece
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

 (including Macedonia
Macedonia (Greece)
Macedonia is a geographical and historical region of Greece in Southern Europe. Macedonia is the largest and second most populous Greek region...

 and Cyclades
Cyclades
The Cyclades is a Greek island group in the Aegean Sea, south-east of the mainland of Greece; and a former administrative prefecture of Greece. They are one of the island groups which constitute the Aegean archipelago. The name refers to the islands around the sacred island of Delos...

), Romania
Romania
Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

, Bulgaria
Bulgaria
Bulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...

, Turkey, Georgia
Georgia (country)
Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...

 and Syria
Syria
Syria , officially the Syrian Arab Republic , is a country in Western Asia, bordering Lebanon and the Mediterranean Sea to the West, Turkey to the north, Iraq to the east, Jordan to the south, and Israel to the southwest....

.
V. aspis
Vipera aspis
Vipera aspis is a venomous viper species found in southwestern Europe. Bites from this species can be more severe than from the European adder, V. berus; not only can they be very painful, but about 4% of all untreated bites are fatal...

T
(Linnaeus, 1758) 4 Asp viper France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

, Andorra
Andorra
Andorra , officially the Principality of Andorra , also called the Principality of the Valleys of Andorra, , is a small landlocked country in southwestern Europe, located in the eastern Pyrenees mountains and bordered by Spain and France. It is the sixth smallest nation in Europe having an area of...

, northeastern Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

, extreme southwestern Germany
Germany
Germany , officially the Federal Republic of Germany , is a federal parliamentary republic in Europe. The country consists of 16 states while the capital and largest city is Berlin. Germany covers an area of 357,021 km2 and has a largely temperate seasonal climate...

, Switzerland
Switzerland
Switzerland name of one of the Swiss cantons. ; ; ; or ), in its full name the Swiss Confederation , is a federal republic consisting of 26 cantons, with Bern as the seat of the federal authorities. The country is situated in Western Europe,Or Central Europe depending on the definition....

, Monaco
Monaco
Monaco , officially the Principality of Monaco , is a sovereign city state on the French Riviera. It is bordered on three sides by its neighbour, France, and its centre is about from Italy. Its area is with a population of 35,986 as of 2011 and is the most densely populated country in the...

, the islands of Elba
Elba
Elba is a Mediterranean island in Tuscany, Italy, from the coastal town of Piombino. The largest island of the Tuscan Archipelago, Elba is also part of the National Park of the Tuscan Archipelago and the third largest island in Italy after Sicily and Sardinia...

 and Montecristo
Montecristo
Montecristo is a small Italian islet, approximately 4.3 kilometers across at its widest point, located about halfway between Corsica and mainland Italy; south of Elba and west of Giglio. It is part of the Tuscan Archipelago...

, Sicily
Sicily
Sicily is a region of Italy, and is the largest island in the Mediterranean Sea. Along with the surrounding minor islands, it constitutes an autonomous region of Italy, the Regione Autonoma Siciliana Sicily has a rich and unique culture, especially with regard to the arts, music, literature,...

, Italy, San Marino
San Marino
San Marino, officially the Republic of San Marino , is a state situated on the Italian Peninsula on the eastern side of the Apennine Mountains. It is an enclave surrounded by Italy. Its size is just over with an estimated population of over 30,000. Its capital is the City of San Marino...

 and northwestern Slovenia.
V. barani
Vipera barani
Vipera barani is a venomous viper species endemic to Turkey. No subspecies are currently recognized.-Description:Stoutly built, adult females are reported to reach a maximum length of 60 cm. Adult males are shorter with a maximum of 54.5 cm....

Böhme & Joger, 1984 0 Baran's adder Northwestern Turkey.
V. berus
Vipera berus
Vipera berus, the common European adder or common European viper, is a venomous viper species that is extremely widespread and can be found throughout most of Western Europe and all the way to Far East Asia. Known by a host of common names including Common adder and Common viper, adders have been...

(Linnaeus, 1758) 2 Common European adder From western Europe
Western Europe
Western Europe is a loose term for the collection of countries in the western most region of the European continents, though this definition is context-dependent and carries cultural and political connotations. One definition describes Western Europe as a geographic entity—the region lying in the...

 (Great Britain
Great Britain
Great Britain or Britain is an island situated to the northwest of Continental Europe. It is the ninth largest island in the world, and the largest European island, as well as the largest of the British Isles...

, Scandinavia
Scandinavia
Scandinavia is a cultural, historical and ethno-linguistic region in northern Europe that includes the three kingdoms of Denmark, Norway and Sweden, characterized by their common ethno-cultural heritage and language. Modern Norway and Sweden proper are situated on the Scandinavian Peninsula,...

, France) across central
Central Europe
Central Europe or alternatively Middle Europe is a region of the European continent lying between the variously defined areas of Eastern and Western Europe...

 (Italy, Albania, Bulgaria and northern Greece) and eastern Europe
Eastern Europe
Eastern Europe is the eastern part of Europe. The term has widely disparate geopolitical, geographical, cultural and socioeconomic readings, which makes it highly context-dependent and even volatile, and there are "almost as many definitions of Eastern Europe as there are scholars of the region"...

 to north of the Arctic Circle
Arctic Circle
The Arctic Circle is one of the five major circles of latitude that mark maps of the Earth. For Epoch 2011, it is the parallel of latitude that runs north of the Equator....

, and Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 to the Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

, Sakhalin Island, North Korea
North Korea
The Democratic People’s Republic of Korea , , is a country in East Asia, occupying the northern half of the Korean Peninsula. Its capital and largest city is Pyongyang. The Korean Demilitarized Zone serves as the buffer zone between North Korea and South Korea...

, northern Mongolia
Mongolia
Mongolia is a landlocked country in East and Central Asia. It is bordered by Russia to the north and China to the south, east and west. Although Mongolia does not share a border with Kazakhstan, its western-most point is only from Kazakhstan's eastern tip. Ulan Bator, the capital and largest...

 and northern China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

.
V. bornmuelleri
Vipera bornmuelleri
Vipera bornmuelleri is a venomous viper species found in Lebanon and Syria. No subspecies are currently recognized.-Description:Grows to a maximum length of about 75 cm, but usually much less. Males tend to be larger than females in some populations. In specimens from Mt. Lebanon, the maximum...

Werner
Franz Werner
Franz Werner was an Austrian zoologist and explorer. Specializing as a herpetologist and entomologist, Werner described numerous species and other taxa of frogs, snakes, insects, and other organisms....

, 1898
0 Bornmuellers viper Golan Heights, southern Lebanon
Lebanon
Lebanon , officially the Republic of LebanonRepublic of Lebanon is the most common term used by Lebanese government agencies. The term Lebanese Republic, a literal translation of the official Arabic and French names that is not used in today's world. Arabic is the most common language spoken among...

 and Syria
Syria
Syria , officially the Syrian Arab Republic , is a country in Western Asia, bordering Lebanon and the Mediterranean Sea to the West, Turkey to the north, Iraq to the east, Jordan to the south, and Israel to the southwest....

.
V. bulgardaghica
Vipera bulgardaghica
Vipera bulgardaghica is a venomous viper species endemic in the mountains of southern Turkey. No subspecies are currently recognized.-Geographic range:The Bulgar Dagh mountains, Nigde Province, south central Anatolia, Turkey...

Nilson & Andrén, 1985 0 Bulgardagh viper The Bulgar Dagh (Bolkar Dagi) mountains, Nigde Province
Nigde Province
Niğde Province is a small rural province in the southern part of Central Anatolia, Turkey. Population is 337.931 of which 100.418 live in the city of Niğde. The population was 348,081 in 2000 and 305.861 in 1990. It covers an area of 7,312 km²...

, south central Anatolia
Anatolia
Anatolia is a geographic and historical term denoting the westernmost protrusion of Asia, comprising the majority of the Republic of Turkey...

, Turkey.
V. darevskii
Vipera darevskii
Vipera darevskii is a venomous viper species found in Armenia and Turkey and possibly also in Georgia. No subspecies are currently recognized.-Geographic range:...

Vedmederja, Orlov & Tuniyev, 1986 0 Darevsky's viper The southeastern Dzavachet Mountains in Armenia
Armenia
Armenia , officially the Republic of Armenia , is a landlocked mountainous country in the Caucasus region of Eurasia...

 and adjacent areas in Georgia
Georgia (country)
Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...

.
V. dinniki
Vipera dinniki
Vipera dinniki is a venomous viper species found in Russia, Georgia and Azerbaijan. No subspecies are currently recognized.-Description:Of the 49 Russian specimens examined by Orlov and Tuniyev , 29 were males and the largest measured 41.2 cm...

Nikolsky, 1913 0 Dinnik's viper Russia (Great Caucasus
Caucasus Mountains
The Caucasus Mountains is a mountain system in Eurasia between the Black Sea and the Caspian Sea in the Caucasus region .The Caucasus Mountains includes:* the Greater Caucasus Mountain Range and* the Lesser Caucasus Mountains....

) and Georgia (high mountain basin of the Inguri River
Inguri
The Inguri , is a river in western Georgia. It is 213 km long, originates in northeastern Svaneti near the region of Racha and plays an important role providing hydroelectric power to the area....

), eastward to Azerbaijan
Azerbaijan
Azerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to...

.
V. kaznakovi
Vipera kaznakovi
Vipera kaznakovi is a venomous viper species found in Turkey, Georgia and Russia. No subspecies are currently recognized.-Description:A stoutly built species, of which the males are usually shorter and more slender than the females. Adults may reach a maximum length of 65-70 cm, but are usually less...

Nikolsky, 1909 0 Caucasus viper Northeastern Turkey, Georgia and Russia (eastern Black Sea
Black Sea
The Black Sea is bounded by Europe, Anatolia and the Caucasus and is ultimately connected to the Atlantic Ocean via the Mediterranean and the Aegean seas and various straits. The Bosphorus strait connects it to the Sea of Marmara, and the strait of the Dardanelles connects that sea to the Aegean...

 coast.
V. latastei
Vipera latastei
Vipera latastei is a venomous viper species found in extreme southwestern Europe and northwestern Africa. Two subspecies are currently recognized, including the nominate subspecies described here.-Description:...

Bosca, 1878 1 Lataste's viper Extreme southwestern Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

 (France, Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

 and Spain) and northwestern Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

 (the Mediterranean region of Morocco
Morocco
Morocco , officially the Kingdom of Morocco , is a country located in North Africa. It has a population of more than 32 million and an area of 710,850 km², and also primarily administers the disputed region of the Western Sahara...

, Algeria
Algeria
Algeria , officially the People's Democratic Republic of Algeria , also formally referred to as the Democratic and Popular Republic of Algeria, is a country in the Maghreb region of Northwest Africa with Algiers as its capital.In terms of land area, it is the largest country in Africa and the Arab...

 and Tunisia
Tunisia
Tunisia , officially the Tunisian RepublicThe long name of Tunisia in other languages used in the country is: , is the northernmost country in Africa. It is a Maghreb country and is bordered by Algeria to the west, Libya to the southeast, and the Mediterranean Sea to the north and east. Its area...

).
V. latifii
Vipera latifii
Vipera latifii is a venomous viper species endemic in Iran. No subspecies are currently recognized.-Description:For adult males, the maximum length is 78 cm, for females 70 cm.Holotype: SMF 62585.-Geographic range:...

Mertens, Darevsky & Klemmer, 1967 0 Latifi's viper Iran: upper Lar Valley in the Elburz Mountains.
V. lotievi
Vipera lotievi
Vipera lotievi is a venomous viper species found in Russia, Georgia and Azerbaijan. No subspecies are currently recognized.-Geographic range:...

Nilson et al., 1995 0 Caucasian meadow viper The higher range of the Big Caucasus
Greater Caucasus
Greater Caucasus , sometimes translated as "Caucasus Major", "Big Caucasus" or "Large Caucasus") is the major mountain range of the Caucasus Mountains....

: Russia, Georgia and Azerbaijan.
V. monticola
Vipera monticola
Vipera monticola is a venomous viper species endemic in Morocco. No subspecies are currently recognized.-Description:A very small species with a maximum length of less than 40 cm. Spawls and Branch describe it as the smallest member of the genus Vipera, reaching a length of only 345 mm.-Geographic...

Saint-Girons, 1954 0 Atlas mountain viper High Atlas Mountains
High Atlas
High Atlas, also called the Grand Atlas Mountains is a mountain range in central Morocco in Northern Africa.The High Atlas rises in the west at the Atlantic Ocean and stretches in an eastern direction to the Moroccan-Algerian border. At the Atlantic and to the southwest the range drops abruptly...

, Morocco.
V. nikolskii Vedmederja, Grubant & Rudajewa, 1986 0 Nikolsky's viper Central Ukraine
Ukraine
Ukraine is a country in Eastern Europe. It has an area of 603,628 km², making it the second largest contiguous country on the European continent, after Russia...

.
V. palaestinae
Vipera palaestinae
Vipera palaestinae is a venomous viper species found in the geographical region of Palestine, more specifically, an area spanning western Syria, northwestern Jordan, northern and central Israel, the West Bank and Lebanon. It is considered a leading cause of snakebite within its range...

Werner, 1938 0 Palestine viper Syria, Jordan
Jordan
Jordan , officially the Hashemite Kingdom of Jordan , Al-Mamlaka al-Urduniyya al-Hashemiyya) is a kingdom on the East Bank of the River Jordan. The country borders Saudi Arabia to the east and south-east, Iraq to the north-east, Syria to the north and the West Bank and Israel to the west, sharing...

, Israel
Israel
The State of Israel is a parliamentary republic located in the Middle East, along the eastern shore of the Mediterranean Sea...

 and Lebanon.
V. pontica
Vipera pontica
Vipera pontica is a venomous viper species found in Turkey and Georgia. No subspecies are currently recognized.-Description:Probably does not grow to more than 60 cm in length. The largest type specimen was 277 mm.Holotype: NHMG 5021....

Billing, Nilson & Sattler, 1990 0 Pontic adder Known only from the Coruh valley in Artvin Province
Artvin Province
Artvin Province is a province in Turkey, on the Black Sea coast in the north-eastern corner of the country, on the border with Georgia.The provincial capital is the city of Artvin.-Geography:...

, northeastern Turkey.
V. raddei
Vipera raddei
Vipera raddei is a venomous viper species found in Turkey, Iran, Armenia, Azerbaijan, and possibly also Iraq. No subspecies are currently recognized.-Description:Adult males grow to a maximum length of 99 cm...

Boettger
Oskar Boettger
Oskar Boettger was a German zoologist who was a native of Frankfurt am Main. He was an uncle of the noted malacologist Caesar Rudolf Boettger ....

, 1890
0 Rock viper Eastern Turkey, northwestern Iran, Armenia, Azerbaijan, and probably Iraq
Iraq
Iraq ; officially the Republic of Iraq is a country in Western Asia spanning most of the northwestern end of the Zagros mountain range, the eastern part of the Syrian Desert and the northern part of the Arabian Desert....

.
V. seoanei
Vipera seoanei
Vipera seoanei is a venomous viper species found in extreme southwestern France and the northern regions of Spain and Portugal. Two subspecies are currently recognized, including the nominate race described here.-Description:...

Lataste
Fernand Lataste
Fernand Lataste was a French zoologist who for example named the fat-tailed gerbil in 1880. The Lataste's Viper Vipera latastei, a viper from Spain, Portugal and North Africa, was named in his honor....

, 1879
1 Baskian viper Extreme southwestern France and the northern regions of Spain and Portugal.
V. ursinii
Vipera ursinii
Vipera ursinii is a venomous viper and a very widespread species, found from southeastern France all the way to China . No subspecies are currently recognized...

(Bonaparte
Charles Lucien Bonaparte
Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano was a French naturalist and ornithologist.-Biography:...

, 1835)
0 Meadow viper Southeastern France, eastern Austria
Austria
Austria , officially the Republic of Austria , is a landlocked country of roughly 8.4 million people in Central Europe. It is bordered by the Czech Republic and Germany to the north, Slovakia and Hungary to the east, Slovenia and Italy to the south, and Switzerland and Liechtenstein to the...

 (extinct), Hungary, central Italy, Croatia, Bosnia-Herzegovina, northern and northeastern Albania, Romania, northern Bulgaria, Greece, Turkey, northwestern Iran, Armenia, Azerbaijan, Georgia, Russia and across the Khazakstan, Kirgizia and eastern Uzbekistan
Uzbekistan
Uzbekistan , officially the Republic of Uzbekistan is a doubly landlocked country in Central Asia and one of the six independent Turkic states. It shares borders with Kazakhstan to the west and to the north, Kyrgyzstan and Tajikistan to the east, and Afghanistan and Turkmenistan to the south....

 steppes to China (Xinjiang
Xinjiang
Xinjiang is an autonomous region of the People's Republic of China. It is the largest Chinese administrative division and spans over 1.6 million km2...

).
V. wagneri
Vipera wagneri
Vipera wagneri is a venomous viper species found in eastern Turkey and northwest Iran. No subspecies are currently recognized.-Description: Grows to a maximum length of 70-95 cm....

Nilson & Andrén, 1984 0 Ocellated mountain viper The mountains of eastern Turkey and adjacent northwest Iran.
V. xanthina
Vipera xanthina
Vipera xanthina is a venomous viper species found in northeastern Greece and Turkey, as well as certain islands in the Aegean Sea. No subspecies are currently recognized.-Description:...

(Gray
John Edward Gray
John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

, 1849)
0 Rock viper Extreme northeastern Greece, the Greek islands of Simi
SIMI
Simi may refer to:*Symi, a Greek island*Simi Valley, California*Students Islamic Movement of India - a students' social Islamic terrorist organization - now banned - in India.*Society of the Irish Motor Industry...

, Kos
Kos
Kos or Cos is a Greek island in the south Sporades group of the Dodecanese, next to the Gulf of Gökova/Cos. It measures by , and is from the coast of Bodrum, Turkey and the ancient region of Caria. Administratively the island forms a separate municipality within the Kos peripheral unit, which is...

, Kalimnos, Leros
Leros
Leros is a Greek island and municipality in the Dodecanese in the southern Aegean Sea. It lies 317 km from Athens's port of Piraeus, from which it can be reached by an 11-hour ferry ride . Leros is part of the Kalymnos peripheral unit...

, Lipsos, Patmos
Patmos
Patmos is a small Greek island in the Aegean Sea. One of the northernmost islands of the Dodecanese complex, it has a population of 2,984 and an area of . The highest point is Profitis Ilias, 269 meters above sea level. The Municipality of Patmos, which includes the offshore islands of Arkoi ,...

, Samos
Samos Island
Samos is a Greek island in the eastern Aegean Sea, south of Chios, north of Patmos and the Dodecanese, and off the coast of Asia Minor, from which it is separated by the -wide Mycale Strait. It is also a separate regional unit of the North Aegean region, and the only municipality of the regional...

, Chios
Chios
Chios is the fifth largest of the Greek islands, situated in the Aegean Sea, seven kilometres off the Asia Minor coast. The island is separated from Turkey by the Chios Strait. The island is noted for its strong merchant shipping community, its unique mastic gum and its medieval villages...

 and Lesbos, European Turkey
European Turkey
European Turkey may refer to:*Eastern Thrace, the European portion of Turkey*Rumelia, the historical Ottoman territories in Europe*the suggested Accession of Turkey to the European Union...

, the western half of Anatolia
Anatolia
Anatolia is a geographic and historical term denoting the westernmost protrusion of Asia, comprising the majority of the Republic of Turkey...

 (inland eastward to Kayseri
Kayseri
Kayseri is a large and industrialized city in Central Anatolia, Turkey. It is the seat of Kayseri Province. The city of Kayseri, as defined by the boundaries of Kayseri Metropolitan Municipality, is structurally composed of five metropolitan districts, the two core districts of Kocasinan and...

), and islands (e.g. Chalki
Chalki
Halki is a Greek island and municipality in the Dodecanese archipelago in the Aegean Sea, some west of Rhodes. With the area of , it is the smallest inhabited island of Dodecanese. It is part of the Rhodes peripheral unit. It has permanent population of 330 , concentrated in the only village...

, Kastellórizon [Meis Adasi]) of the Turkish mainland shelf.
  • ) Not including the nominate subspecies.

T) Type species
Type species
In biological nomenclature, a type species is both a concept and a practical system which is used in the classification and nomenclature of animals and plants. The value of a "type species" lies in the fact that it makes clear what is meant by a particular genus name. A type species is the species...

.

External links

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