Torsion (gastropod)
Overview
Torsion is an anatomical event which takes place during the very early part of the life of snail
Snail
Snail is a common name applied to most of the members of the molluscan class Gastropoda that have coiled shells in the adult stage. When the word is used in its most general sense, it includes sea snails, land snails and freshwater snails. The word snail without any qualifier is however more often...

s and slug
Slug
Slug is a common name that is normally applied to any gastropod mollusc that lacks a shell, has a very reduced shell, or has a small internal shell...

s of all kinds. In other words, torsion is a gastropod synapomorphy
Synapomorphy
In cladistics, a synapomorphy or synapomorphic character is a trait that is shared by two or more taxa and their most recent common ancestor, whose ancestor in turn does not possess the trait. A synapomorphy is thus an apomorphy visible in multiple taxa, where the trait in question originates in...

 which occurs in all gastropods during larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

l development. Torsion is the rotation of the visceral mass
Viscus
In anatomy, a viscus is an internal organ, and viscera is the plural form. The viscera, when removed from a butchered animal, are known collectively as offal...

, mantle
Mantle (mollusc)
The mantle is a significant part of the anatomy of molluscs: it is the dorsal body wall which covers the visceral mass and usually protrudes in the form of flaps well beyond the visceral mass itself.In many, but by no means all, species of molluscs, the epidermis of the mantle secretes...

 and shell
Gastropod shell
The gastropod shell is a shell which is part of the body of a gastropod or snail, one kind of mollusc. The gastropod shell is an external skeleton or exoskeleton, which serves not only for muscle attachment, but also for protection from predators and from mechanical damage...

 180˚ with respect to the head and foot of the gastropod. This brings the mantle cavity and anus to an anterior position above the head.

In some groups of gastropods (Opisthobranchia
Opisthobranchia
Opisthobranchs are a large and diverse group of specialized complex marine gastropods previously united under Opisthobranchia within the Heterobranchia, but no longer considered to represent a monophyletic grouping...

) there is a degree of secondary detorsion or rotation towards the original position, this may be only partial detorsion or full detorsion.

The torsion or twisting of the visceral mass of larval gastropods is not the same thing as the spiral coiling of the shell
Gastropod shell
The gastropod shell is a shell which is part of the body of a gastropod or snail, one kind of mollusc. The gastropod shell is an external skeleton or exoskeleton, which serves not only for muscle attachment, but also for protection from predators and from mechanical damage...

 of many shelled gastropods.
There are two different developmental stages which cause torsion.
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