Thermoception
Encyclopedia
Thermoception or thermoreception is the sense
Sense
Senses are physiological capacities of organisms that provide inputs for perception. The senses and their operation, classification, and theory are overlapping topics studied by a variety of fields, most notably neuroscience, cognitive psychology , and philosophy of perception...

 by which an organism
Organism
In biology, an organism is any contiguous living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homoeostasis as a stable whole.An organism may either be unicellular or, as in the case of humans, comprise...

 perceives temperature
Temperature
Temperature is a physical property of matter that quantitatively expresses the common notions of hot and cold. Objects of low temperature are cold, while various degrees of higher temperatures are referred to as warm or hot...

. The details of how temperature receptors work are still being investigated. Ciliopathy
Ciliopathy
A ciliopathy is a genetic disorder of the cellular cilia or the cilia anchoring structures, the basal bodies, or of ciliary function.Although ciliopathies are usually considered to involve proteins that localize to the primary cilia or centrosomes, it is possible for ciliopathies to be associated...

 is associated with decreased ability to sense heat, thus cilia may aid in the process. Transient receptor potential channels (TRP channels) are believed to play a role in many species in sensation of hot, cold, and pain. Mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

s have at least two types of sensor: those that detect heat (i.e. temperatures above body temperature) and those that detect cold (i.e. temperatures below body temperature).

A particularly specialized form of thermoception is used by Crotalinae
Crotalinae
The Crotalinae, commonly known as "pit vipers" or crotaline snakes, are a subfamily of venomous vipers found in Asia and the Americas. They are distinguished by the presence of a heat-sensing pit organ located between the eye and the nostril on either side of the head...

 (pit viper) and Boidae (boa) snakes, which can effectively see the infrared radiation emitted
Infrared sensing in snakes
The ability to sense infrared thermal radiation evolved independently in several different families of snakes. Essentially, it allows these animals to “see” radiant heat at wavelengths between 5 and 30 μm to a degree of accuracy such that a blind rattlesnake can target vulnerable body parts of the...

 by hot objects. The snake's face has a pair of holes, or pits, lined with temperature sensors. The sensors indirectly detect infrared radiation by its heating effect on the skin inside the pit. They can work out which part of the pit is hottest, and therefore the direction of the heat source, which could be a warm-blooded prey animal. By combining information from both pits, the snake can also estimate the distance of the object.

The common vampire bat
Vampire bat
Vampire bats are bats whose food source is blood, a dietary trait called hematophagy. There are three bat species that feed solely on blood: the Common Vampire Bat , the Hairy-legged Vampire Bat , and the White-winged Vampire Bat .All three species are native to the Americas, ranging from Mexico to...

 may also have specialized infrared sensors on its nose (see ). A nucleus has been found in the brain of vampire bats that has a similar position and has similar histology
Histology
Histology is the study of the microscopic anatomy of cells and tissues of plants and animals. It is performed by examining cells and tissues commonly by sectioning and staining; followed by examination under a light microscope or electron microscope...

 to the infrared nucleus of infrared sensitive snakes.

Other animals with specialized heat detectors are forest fire seeking beetles (Melanophilia acuminata), which lay their eggs in conifers freshly killed by forest fires. Darkly pigmented butterflies Pachliopta aristolochiae
Pachliopta aristolochiae
The Common Rose is a swallowtail butterfly belonging to the Pachliopta subgenus, the Roses, of the genus Atrophaneura or Red-bodied Swallowtails. It is a common butterfly which is extensively distributed across South and South East Asia.-Range:It is widely distributed in Asia...

 and Troides rhadamathus use specialized heat detectors to avoid damage while basking. The blood sucking bugs
Hemiptera
Hemiptera is an order of insects most often known as the true bugs , comprising around 50,000–80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, and others...

 Triatoma infestans
Triatoma infestans
Triatoma infestans, commonly called vinchuca and also known as "barber bug", is a blood-sucking bug and the most important vector of Trypanosoma cruzi which can lead to Chagas disease...

may also have a specialised thermoception organ.
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