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The Extended Phenotype

 
The Extended Phenotype

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The Extended Phenotype



 
 
The Extended Phenotype (subtitled "The Gene as the Unit of Selection", and later, "The Long Reach of the Gene") is a 1982 book by Richard Dawkins
Richard Dawkins

Clinton Richard Dawkins, Royal Society#Fellowship, Royal Society of Literature is a United Kingdom ethology, evolutionary biology and popular science author....
. A revised edition was published in 1999 with an afterword by the philosopher Daniel Dennett
Daniel Dennett

Daniel Clement Dennett is a prominent United States Philosophy whose research centers on philosophy of mind, philosophy of science and philosophy of biology, particularly as those fields relate to evolutionary biology and cognitive science....
. Dawkins considers the concept of the Extended Phenotype to be his principal contribution to evolutionary theory
Evolution

In biology, evolution is change in the heritability trait of a population of organisms from one generation to the next. These changes are caused by a combination of three main processes: variation, reproduction, and selection....
.

Dawkins starts from the ideas of his 1976 book The Selfish Gene
The Selfish Gene

The Selfish Gene is a book on evolution by Richard Dawkins, published in 1976 in literature. It builds upon the principal theory of George C....
, which portrayed the organism
Organism

In biology, an organism is any life thing . In at least some form, all organisms are capable of response to stimulus , reproduction, growth and developmental biology, and maintenance of homeostasis as a stable whole....
 as a survival machine constructed by its gene
Gene

A gene is the basic unit of heredity in a living organism. All living things depend on genes. Genes hold the information to build and maintain their cell and pass genetic trait to offspring....
s to maximise their chances of replicating.






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The Extended Phenotype (subtitled "The Gene as the Unit of Selection", and later, "The Long Reach of the Gene") is a 1982 book by Richard Dawkins
Richard Dawkins

Clinton Richard Dawkins, Royal Society#Fellowship, Royal Society of Literature is a United Kingdom ethology, evolutionary biology and popular science author....
. A revised edition was published in 1999 with an afterword by the philosopher Daniel Dennett
Daniel Dennett

Daniel Clement Dennett is a prominent United States Philosophy whose research centers on philosophy of mind, philosophy of science and philosophy of biology, particularly as those fields relate to evolutionary biology and cognitive science....
. Dawkins considers the concept of the Extended Phenotype to be his principal contribution to evolutionary theory
Evolution

In biology, evolution is change in the heritability trait of a population of organisms from one generation to the next. These changes are caused by a combination of three main processes: variation, reproduction, and selection....
.

Dawkins starts from the ideas of his 1976 book The Selfish Gene
The Selfish Gene

The Selfish Gene is a book on evolution by Richard Dawkins, published in 1976 in literature. It builds upon the principal theory of George C....
, which portrayed the organism
Organism

In biology, an organism is any life thing . In at least some form, all organisms are capable of response to stimulus , reproduction, growth and developmental biology, and maintenance of homeostasis as a stable whole....
 as a survival machine constructed by its gene
Gene

A gene is the basic unit of heredity in a living organism. All living things depend on genes. Genes hold the information to build and maintain their cell and pass genetic trait to offspring....
s to maximise their chances of replicating. In a much more technical presentation than the earlier book, Dawkins devotes a significant portion of this work to an attempt to rebut criticism of The Selfish Gene.

Genes synthesize only proteins


Termite Cathedral Dsc03570
In the main portion of the book, Dawkins argues that the only thing that genes control directly is the synthesis of protein
Protein

Proteins are organic compounds made of amino acids arranged in a linear chain and joined together by peptide bonds between the carboxyl and amino groups of adjacent amino acid Residue ....
s. He points to the arbitrariness of restricting the idea of the phenotype
Phenotype

A phenotype is any observable characteristic or trait_ of an organism: such as its morphology , development, biochemical or physiological properties, or behavior....
 to apply only to the phenotypic expression of an organism's genes in its own body.

Genes affect the organism’s environment


Dawkins develops this idea by pointing to the effect that a gene may have on an organism's environment through that organism's behaviour, citing as examples caddis houses
Trichoptera

Caddisflies, sedge-flies or rail flies are small moth-like insects having two pairs of hairy membranous insect wing. They are closely related to Lepidoptera which have scales on their wings, and the two orders together form the superorder Amphiesmenoptera....
 and beaver dams
Beaver

Beavers are two primarily nocturnal, semi-aquatic species of rodent, one native to North America and one to Eurasia. They are known for building dams, canals, and lodges ....
. He then goes further to point to first animal morphology
Morphology (biology)

The term morphology in biology refers to form, structure and configuration of an organism. This includes aspects of the outward appearance as well as the form and structure of the internal parts like bones and organs....
 and ultimately animal behaviour, which can seem advantageous not to the animal itself, but rather to a parasite which afflicts it. Dawkins summarizes these ideas in what he terms the Central Theorem of the Extended Phenotype:

Gene-centred view of life


In conducting this argument, Dawkins aims to strengthen the case for a gene-centric
Gene-centered view of evolution

The gene-centered view of evolution, gene selection theory or selfish gene theory holds that natural selection acts through differential survival of competing genes, increasing the frequency of those alleles whose Phenotype effects successfully promote their own propagation....
 view of life, to the point where it is recognized that the organism itself needs to be explained. This is the challenge which he takes up in the final chapter entitled "Rediscovering the Organism."

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