Taxodium distichum
Encyclopedia
Taxodium distichum is a species of conifer
Pinophyta
The conifers, division Pinophyta, also known as division Coniferophyta or Coniferae, are one of 13 or 14 division level taxa within the Kingdom Plantae. Pinophytes are gymnosperms. They are cone-bearing seed plants with vascular tissue; all extant conifers are woody plants, the great majority being...

 native to the southeastern United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

.

Characteristics

It is a large tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

, reaching 25–40 m (rarely to 44 m) tall and a trunk diameter of 2–3 m, rarely to 5 m. The bark
Bark
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines and shrubs. Bark refers to all the tissues outside of the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner...

 is gray-brown to red-brown, shallowly vertically fissured, with a stringy texture. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are borne on deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 branchlets that are spirally arranged on the stem but twisted at the base to lie in two horizontal ranks, 1-2 cm long and 1-2 mm broad; unlike most other species in the family Cupressaceae, it is deciduous, losing the leaves in the winter months, hence the name 'bald'. It is monoecious. Male and female strobili mature in about 12 months; they are produced from buds formed in the late fall, with pollination in early winter. The seed cones
Conifer cone
A cone is an organ on plants in the division Pinophyta that contains the reproductive structures. The familiar woody cone is the female cone, which produces seeds. The male cones, which produce pollen, are usually herbaceous and much less conspicuous even at full maturity...

 are green maturing gray-brown, globular, 2-3.5 cm in diameter. They have from 20–30 spirally arranged four-sided scales, each bearing one or two (rarely three) trianglular seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s. The number of seeds per cone ranges from 20–40. The cones disintegrate when mature to release the large seeds. The seeds are 5-10 mm long, the largest of any species in the cypress family, and are produced every year but with heavy crops every three to five years. The seedlings have 3–9 (most often 6) cotyledon
Cotyledon
A cotyledon , is a significant part of the embryo within the seed of a plant. Upon germination, the cotyledon may become the embryonic first leaves of a seedling. The number of cotyledons present is one characteristic used by botanists to classify the flowering plants...

s.

The main trunks are surrounded by cypress knees.

The tallest known individual specimen, near Williamsburg, Virginia
Williamsburg, Virginia
Williamsburg is an independent city located on the Virginia Peninsula in the Hampton Roads metropolitan area of Virginia, USA. As of the 2010 Census, the city had an estimated population of 14,068. It is bordered by James City County and York County, and is an independent city...

, is 44.11 m tall, and the stoutest known, in the Cat Island National Wildlife Refuge
Cat Island National Wildlife Refuge
Cat Island National Wildlife Refuge was established on October 27, 2000 as the 526th refuge in the United States National Wildlife Refuge System. It is located near the town of St. Francisville, Louisiana, which is north of Baton Rouge...

 near Baton Rouge, Louisiana
Baton Rouge, Louisiana
Baton Rouge is the capital of the U.S. state of Louisiana. It is located in East Baton Rouge Parish and is the second-largest city in the state.Baton Rouge is a major industrial, petrochemical, medical, and research center of the American South...

, has a diameter
Diameter at breast height
Diameter at breast height, or DBH, is a standard method of expressing the diameter of the trunk or bole of a standing tree. DBH is one of the most common dendrometric measurements....

 of 521 cm. The oldest known specimen, located in Bladen County, North Carolina
Bladen County, North Carolina
-Demographics:As of the census of 2000, there were 32,278 people, 12,897 households, and 8,937 families residing in the county. The population density was 37 people per square mile . There were 15,316 housing units at an average density of 18 per square mile...

, is over 1,620 years old making this the oldest living plant in Eastern North America .

Taxonomy

The closely related Taxodium ascendens
Taxodium ascendens
Taxodium ascendens, also known as Pond Cypress, is a deciduous conifer of the genus Taxodium, native to North America. Many botanists treat it as a variety of Bald Cypress, Taxodium distichum Taxodium ascendens, also known as Pond Cypress, is a deciduous conifer of the genus Taxodium, native to...

(Pondcypress) is treated by some botanists as a distinct species, while others classify it as merely a variety of Bald-cypress, as Taxodium distichum var. imbricatum (Nutt.) Croom. It differs in shorter leaves borne on erect shoots, and in ecology, being largely confined to low nutrient blackwater
Blackwater river
A blackwater river is a river with a deep, slow-moving channel that flows through forested swamps and wetlands. As vegetation decays in the water, tannins are leached out, resulting in transparent, acidic water that is darkly stained, resembling tea or coffee. Most major blackwater rivers are in...

 habitats. A few authors also treat Taxodium mucronatum
Taxodium mucronatum
Taxodium mucronatum, also known as Montezuma Cypress, Sabino, or Ahuehuete is a species of Taxodium native to much of Mexico , and also the Rio Grande Valley in southernmost Texas, USA as well as Huehuetenango Department in Guatemala...

as a variety of Bald-cypress, as T. distichum var. mexicanum Gordon, thereby considering the genus as comprising only one species.

Range

The native range extends from Delaware Bay
Delaware Bay
Delaware Bay is a major estuary outlet of the Delaware River on the Northeast seaboard of the United States whose fresh water mixes for many miles with the waters of the Atlantic Ocean. It is in area. The bay is bordered by the State of New Jersey and the State of Delaware...

 south to Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

 and west to Texas
East Texas
East Texas is a distinct geographic and ecological area in the U.S. state of Texas.According to the Handbook of Texas, the East Texas area "may be separated from the rest of Texas roughly by a line extending from the Red River in north central Lamar County southwestward to east central Limestone...

 and southeastern Oklahoma
Oklahoma
Oklahoma is a state located in the South Central region of the United States of America. With an estimated 3,751,351 residents as of the 2010 census and a land area of 68,667 square miles , Oklahoma is the 28th most populous and 20th-largest state...

-(Little Dixie
Little Dixie (Oklahoma)
Little Dixie is the name given to the southeast region of Oklahoma, which is heavily influenced by southern "Dixie" culture, as it was settled chiefly by Southerners seeking a start in new lands following the American Civil War...

 region, Oklahoma), and also inland up the Mississippi
Mississippi River
The Mississippi River is the largest river system in North America. Flowing entirely in the United States, this river rises in western Minnesota and meanders slowly southwards for to the Mississippi River Delta at the Gulf of Mexico. With its many tributaries, the Mississippi's watershed drains...

 and Ohio River
Ohio River
The Ohio River is the largest tributary, by volume, of the Mississippi River. At the confluence, the Ohio is even bigger than the Mississippi and, thus, is hydrologically the main stream of the whole river system, including the Allegheny River further upstream...

s north to southern Illinois
Illinois
Illinois is the fifth-most populous state of the United States of America, and is often noted for being a microcosm of the entire country. With Chicago in the northeast, small industrial cities and great agricultural productivity in central and northern Illinois, and natural resources like coal,...

 and Indiana
Indiana
Indiana is a US state, admitted to the United States as the 19th on December 11, 1816. It is located in the Midwestern United States and Great Lakes Region. With 6,483,802 residents, the state is ranked 15th in population and 16th in population density. Indiana is ranked 38th in land area and is...

. Large planted specimens are seen as far north as Pittsburgh . Ancient Bald-cypress forest
Forest
A forest, also referred to as a wood or the woods, is an area with a high density of trees. As with cities, depending where you are in the world, what is considered a forest may vary significantly in size and have various classification according to how and what of the forest is composed...

s, with some trees more than 1,700 years old, once dominated swamp
Swamp
A swamp is a wetland with some flooding of large areas of land by shallow bodies of water. A swamp generally has a large number of hammocks, or dry-land protrusions, covered by aquatic vegetation, or vegetation that tolerates periodical inundation. The two main types of swamp are "true" or swamp...

s in the southeast US. The largest remaining old-growth stands of Bald-cypress are at Corkscrew Swamp Sanctuary
Corkscrew Swamp Sanctuary
Corkscrew Swamp Sanctuary is a National Audubon Society sanctuary located in southwest Florida, north of Naples, Florida and east of Bonita Springs, in the United States. The sanctuary was established to protect one of the largest remaining stands of Bald Cypress and Pond Cypress Corkscrew Swamp...

, near Naples
Naples, Florida
Naples is a city in Collier County, Florida, United States. As of July 1, 2007, the U.S. Census Bureau estimated the city's population at 21,653. Naples is a principal city of the Naples–Marco Island Metropolitan Statistical Area, which had an estimated total population of 315,839 on July 1, 2007...

, Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

. and in the Three Sisters tract along eastern North Carolina's Black River
Black River (North Carolina)
The Black River is a tributary of the Cape Fear River, approximately 50 mi long, in southeastern North Carolina in the United States....

. The Corkscrew trees are around 500 years of age and some exceed 40 m in height. The Black River trees were cored in 1986 by University of Arkansas
University of Arkansas
The University of Arkansas is a public, co-educational, land-grant, space-grant, research university. It is classified by the Carnegie Foundation as a research university with very high research activity. It is the flagship campus of the University of Arkansas System and is located in...

  dendrologists with dates ranging back to 364 AD. In the northern and more inland part of its range from Delaware and Maryland to Williamsburg, Virginia, it is found in groups growing in swamps and is accompanied by other hardwoods. In the southern parts of its range from extreme southeastern Virginia, Virginia Beach south to Florida and west to Texas, bald cypress can be found growing with loblolly pine
Loblolly Pine
Pinus taeda is one of several pines native to the Southeastern United States, from central Texas east to Florida, and north to Delaware. It is particularly dominant in the eastern half of North Carolina, where there are huge expanses consisting solely of Loblolly Pine trees...

, live oak
Live oak
Live oak , also known as the southern live oak, is a normally evergreen oak tree native to the southeastern United States...

 and it may be heavily covered in spanish moss
Spanish Moss
Spanish moss is a flowering plant that grows upon larger trees, commonly the Southern Live Oak or Bald Cypress in the southeastern United States....

. A place to observe this in the far northern part of its range is at First Landing State Park
First Landing State Park
First Landing State Park offers recreational opportunities at Cape Henry in the independent city of Virginia Beach, Virginia....

, Virginia Beach, Virginia where you will see bald cypress growing with live oak, loblolly pine, spanish moss and other trees at their farthest north and farthest south ranges. From eastern North Carolina
Eastern North Carolina
Eastern North Carolina is the region encompassing the eastern tier of North Carolina. It is known geographically as the state's Coastal Plain region. Primary subregions of Eastern North Carolina include the Fayetteville Metropolitan Area, the Lower Cape Fear , the Sandhills, the Inner Banks and...

 down throughout Florida, bald cypress may be accompanied in forests by sabal minor (dwarf palmetto).

It is native to humid climates where annual precipitation ranges from about 760 mm (in Texas) to 1630 mm (along the Gulf Coast).

Although it grows best in warm climates, the natural northern limit of the species is not due to a lack of cold tolerance, but to specific reproductive requirements; further north, regeneration is prevented by ice
Ice
Ice is water frozen into the solid state. Usually ice is the phase known as ice Ih, which is the most abundant of the varying solid phases on the Earth's surface. It can appear transparent or opaque bluish-white color, depending on the presence of impurities or air inclusions...

 damage to seedlings. Larger trees are able to tolerate much lower temperatures and lower humidities.

Soils and Topography

Most Bald-cypress grow on flat ground on alluvial
Alluvium
Alluvium is loose, unconsolidated soil or sediments, eroded, deposited, and reshaped by water in some form in a non-marine setting. Alluvium is typically made up of a variety of materials, including fine particles of silt and clay and larger particles of sand and gravel...

 soils, usually at elevations of less than 50 m above sea level, although some stands may occur at elevations of 500 m in Texas.

Bald-cypress occurs mainly along riparian (riverside) wetlands normally subject to periodic flooding by silt-rich 'brownwater' rivers, unlike the related Taxodium ascendens
Taxodium ascendens
Taxodium ascendens, also known as Pond Cypress, is a deciduous conifer of the genus Taxodium, native to North America. Many botanists treat it as a variety of Bald Cypress, Taxodium distichum Taxodium ascendens, also known as Pond Cypress, is a deciduous conifer of the genus Taxodium, native to...

, which occurs in silt-poor blackwater river
Blackwater river
A blackwater river is a river with a deep, slow-moving channel that flows through forested swamps and wetlands. As vegetation decays in the water, tannins are leached out, resulting in transparent, acidic water that is darkly stained, resembling tea or coffee. Most major blackwater rivers are in...

s and pond
Pond
A pond is a body of standing water, either natural or man-made, that is usually smaller than a lake. A wide variety of man-made bodies of water are classified as ponds, including water gardens, water features and koi ponds; all designed for aesthetic ornamentation as landscape or architectural...

s. Bald-cypress tolerates minor salinity, but does not grow in brackish or saline coastal waters.

Ecology

The seeds remain viable for less than one year, and are dispersed in two ways. One is by water; the seeds float and move on water until flooding recedes or the cone is deposited on shore. The second is by wildlife; squirrel
Squirrel
Squirrels belong to a large family of small or medium-sized rodents called the Sciuridae. The family includes tree squirrels, ground squirrels, chipmunks, marmots , flying squirrels, and prairie dogs. Squirrels are indigenous to the Americas, Eurasia, and Africa and have been introduced to Australia...

s eat seeds but often drop some scales from the cones they harvest. Seeds do not germinate under water and rarely germinate on well drained soils; seedlings normally become established on soil that is continuously saturated, but not flooded, for one to three months. After germination, seedlings must grow quickly to escape floodwaters; they often reach a height of 20–75 cm (up to 100 cm in fertilized nursery
Nursery (horticulture)
A nursery is a place where plants are propagated and grown to usable size. They include retail nurseries which sell to the general public, wholesale nurseries which sell only to businesses such as other nurseries and to commercial gardeners, and private nurseries which supply the needs of...

 conditions) in their first year. Seedlings die if inundated for more than about two to four weeks. Natural regeneration is therefore prevented on sites that are always flooded during the growing season. Although vigorous saplings and stump sprouts can produce viable seed, most specimens do not produce seed until they are about 30 years old. In good conditions, Bald-cypress grows fairly fast when young, then more slowly with age. Trees have been measured to reach 3 m in five years, 21 m tall in 41 years, and 36 m in 96 years; height growth has largely ceased by the time the trees are 200 years old. Some individuals can live over 1,000 years. It may be difficult to determine the age of an old tree because of frequent missing or false rings of stemwood caused by variable and stressful growing environments.

Bald-cypress growing in swamps have a peculiarity of growth called cypress knee
Cypress knee
In the biology of trees, a cypress knee is a distinctive structure in a root of a cypress tree of any of various species of the subfamily Taxodioideae. Their function is unknown, but they are generally seen in swamps...

s. These are woody projections sent above the ground or water that are part of the root
Root
In vascular plants, the root is the organ of a plant that typically lies below the surface of the soil. This is not always the case, however, since a root can also be aerial or aerating . Furthermore, a stem normally occurring below ground is not exceptional either...

 system. Their function was once thought to be to provide oxygen
Oxygen
Oxygen is the element with atomic number 8 and represented by the symbol O. Its name derives from the Greek roots ὀξύς and -γενής , because at the time of naming, it was mistakenly thought that all acids required oxygen in their composition...

 to the roots, which grow in the low dissolved oxygen (DO) waters typical of a swamp (as in mangrove
Mangrove
Mangroves are various kinds of trees up to medium height and shrubs that grow in saline coastal sediment habitats in the tropics and subtropics – mainly between latitudes N and S...

s). However, there is little evidence for this; in fact, roots of swamp-dwelling specimens whose knees are removed do not decrease in oxygen content and the trees continue to thrive. Another more likely function is that of structural support and stabilization. Bald-cypress growing on flood-prone sites tend to form buttressed bases, but trees grown on drier sites may lack this feature. Buttressed bases and a strong, intertwined root system allows them to resist very strong winds; even hurricanes rarely overturn them.

Many agents damage Bald-cypress trees. The main lethal damaging agent is the fungus
Fungus
A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...

 Stereum taxodii, which causes a brown pocket rot known as "pecky cypress". It attacks the heartwood of living trees, usually from the crown down to the roots. A few other fungi attack the sapwood and the heartwood of the tree, but they do not usually cause serious damage. Insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s like the cypress flea beetle and the baldcypress leafroller (Archips goyerana) (closely related to the fruit tree leafroller
Archips argyrospila
The Fruit-Tree Leafroller Moth is a moth of the Tortricidae family. It is found in most of the United States and southern Canada....

) can seriously damage Bald Cypress trees by destroying leaves, cones or the bark of tree. Coypu
Coypu
The coypu , , also known as the river rat, and nutria, is a large, herbivorous, semiaquatic rodent and the only member of the family Myocastoridae. Originally native to subtropical and temperate South America, it has since been introduced to North America, Europe, Asia, and Africa, primarily by...

 also clip and unroot young Bald-cypress seedlings, sometimes killing a whole plantation in a short amount of time.

Cultivation and uses

It is a very popular ornamental tree, grown for its light, feathery foliage and orange-brown fall color, which can also range to a dull red. In cultivation it thrives on a wide range of soils including well-drained sites where it would not grow naturally due to the inability of the young seedlings to compete with other vegetation. Cultivation is successful far to the north of its native range, north to southern Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

. It is also commonly planted in Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

, Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 and elsewhere with temperate to subtropical climates. It does however require continental climate
Continental climate
Continental climate is a climate characterized by important annual variation in temperature due to the lack of significant bodies of water nearby...

s with hot summers for good growth; when planted in areas with cool summer oceanic climate
Oceanic climate
An oceanic climate, also called marine west coast climate, maritime climate, Cascadian climate and British climate for Köppen climate classification Cfb and subtropical highland for Köppen Cfb or Cwb, is a type of climate typically found along the west coasts at the middle latitudes of some of the...

s, growth is healthy but very slow (some in northeastern England have only reached 4–5 m tall in about 50 years), and cones are not produced.

Bald-cypress has been noted for its high merchantable yields. In virgin stands, yields from 112 to 196 m³/ha were common, and some stands might have exceeded 1000 m³/ha. Bald-cypress swamps are some of the world's most productive ecosystems.

The odorless wood of Bald-cypress, closely resembling that of Cupressus
Cupressus
The genus Cupressus is one of several genera within the family Cupressaceae that have the common name cypress; for the others, see cypress. It is considered a polyphyletic group...

, has long been valued for its water resistance thus called 'wood eternal'. Still-usable prehistoric wood is often found in swamps as far North as New Jersey, and occasionally as far as New England, although it is more common in the southeast. The somewhat-mineralized wood is mined from some swamps in the southeast, and is highly prized for specialty uses such as wood carvings. Pecky cypress, caused by the fungus Stereum taxodii is used for decorative wall paneling.

The Bald-cypress was designated the official state tree of Louisiana
Louisiana
Louisiana is a state located in the southern region of the United States of America. Its capital is Baton Rouge and largest city is New Orleans. Louisiana is the only state in the U.S. with political subdivisions termed parishes, which are local governments equivalent to counties...

 in 1963. It is considered by some to be a symbol of the southern swamps.

Cypress trees can be used in the making of shingles. Joshua D. Brown
Joshua Brown (Texas pioneer)
Joshua D. Brown was a Texas pioneer who in the 1850s became the first settler of Kerrville, the seat of Kerr County in the Texas Hill Country west of San Antonio....

, the first settler of Kerrville
Kerrville, Texas
Kerrville is a city in Kerr County, Texas, United States. The population was 20,425 at the 2000 census. In 2009, the population was 22,826...

, Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

, made his living producing shingles from cypress growing along the Guadalupe River
Guadalupe River (Texas)
The Guadalupe River runs from Kerr County, Texas to San Antonio Bay on the Gulf of Mexico. The river is a popular destination for rafters and canoers. Larger cities along the river include New Braunfels, Kerrville, Seguin, Gonzales, Cuero, and Victoria...

 of the Texas Hill Country
Texas Hill Country
The Texas Hill Country is a vernacular term applied to a region of Central Texas featuring tall rugged hills consisting of thin layers of soil atop limestone or granite. It also includes the Llano Uplift and the second largest granite monadnock in the United States, Enchanted Rock, which is located...

.

See also

  • Battle Creek Cypress Swamp
    Battle Creek Cypress Swamp
    Battle Creek Cypress SwampThe Battle Creek Cypress Swamp, Maryland, in springDesignationU.S. National Natural LandmarkLocationCalvert County, MarylandNearest CityPrince Frederick, MarylandAreaAbout 100 acresAbout 40 ha...

    , Maryland
  • Barley Barber Swamp
    Barley Barber Swamp
    The Barley Barber Swamp, once a vast Bald Cypress forest and waterway in the Greater Everglades watershed, is a parcel of land surrounded by Florida Power & Light's Martin County power plant. The swamp is located just over a mile east of Lake Okeechobee and due west of Indiantown, Florida...

    , Florida
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