Subnivean
Encyclopedia
Subnivean refers to a zone that is in or under the snow layer. From the Latin for "under" (sub) and "snow" (nives). This space can form when latent heat from the ground melts a thin layer of snow above it, leaving, in places, a layer of air between the ground and the additional snow cover. The space is typically 1–4 cm in height.

Subnivean animal
Animal
Animals are a major group of multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and...

s include small mammals such as mice
Mouse
A mouse is a small mammal belonging to the order of rodents. The best known mouse species is the common house mouse . It is also a popular pet. In some places, certain kinds of field mice are also common. This rodent is eaten by large birds such as hawks and eagles...

, vole
Vole
A vole is a small rodent resembling a mouse but with a stouter body, a shorter hairy tail, a slightly rounder head, smaller ears and eyes, and differently formed molars . There are approximately 155 species of voles. They are sometimes known as meadow mice or field mice in North America...

s, shrew
Shrew
A shrew or shrew mouse is a small molelike mammal classified in the order Soricomorpha. True shrews are also not to be confused with West Indies shrews, treeshrews, otter shrews, or elephant shrews, which belong to different families or orders.Although its external appearance is generally that of...

s, and lemming
Lemming
Lemmings are small rodents, usually found in or near the Arctic, in tundra biomes. They are subniveal animals, and together with voles and muskrats, they make up the subfamily Arvicolinae , which forms part of the largest mammal radiation by far, the superfamily Muroidea, which also includes rats,...

s that must rely on winter snow cover for survival. These mammals move under the snow for protection from heat loss and some predators. In winter regions that do not have permafrost
Permafrost
In geology, permafrost, cryotic soil or permafrost soil is soil at or below the freezing point of water for two or more years. Ice is not always present, as may be in the case of nonporous bedrock, but it frequently occurs and it may be in amounts exceeding the potential hydraulic saturation of...

, the subnivean zone maintains a temperature of close to 32 °F (0 °C) regardless of the temperature above the snow cover, once the snow cover has reached a depth of six inches (15 cm) or more. The sinuous tunnels left by these small mammals can be seen from above when the snow melts to the final inch or so.

Some winter predators such as fox
Fox
Fox is a common name for many species of omnivorous mammals belonging to the Canidae family. Foxes are small to medium-sized canids , characterized by possessing a long narrow snout, and a bushy tail .Members of about 37 species are referred to as foxes, of which only 12 species actually belong to...

es and large owl
Owl
Owls are a group of birds that belong to the order Strigiformes, constituting 200 bird of prey species. Most are solitary and nocturnal, with some exceptions . Owls hunt mostly small mammals, insects, and other birds, although a few species specialize in hunting fish...

s can hear their prey through the snow and pounce from above. Ermine can enter and hunt below the snowpack. Snowmobile
Snowmobile
A snowmobile, also known in some places as a snowmachine, or sled,is a land vehicle for winter travel on snow. Designed to be operated on snow and ice, they require no road or trail. Design variations enable some machines to operate in deep snow or forests; most are used on open terrain, including...

s and ATV
All-terrain vehicle
An all-terrain vehicle , also known as a quad, quad bike, three wheeler, or four wheeler, is defined by the American National Standards Institute as a vehicle that travels on low pressure tires, with a seat that is straddled by the operator, along with handlebars for steering control...

s can collapse the subnivean space. Skis and snow shoes are less likely to collapse subnivean space if the snowpack is deep enough.

Larger animals also utilize subnivean space. In the Arctic, ringed seals
Pinniped
Pinnipeds or fin-footed mammals are a widely distributed and diverse group of semiaquatic marine mammals comprising the families Odobenidae , Otariidae , and Phocidae .-Overview: Pinnipeds are typically sleek-bodied and barrel-shaped...

 have closed spaces under the snow and above openings in the ice. In addition to resting and sleeping there, the female seals give birth to their pups on the ice. Female polar bear
Polar Bear
The polar bear is a bear native largely within the Arctic Circle encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the world's largest land carnivore and also the largest bear, together with the omnivorous Kodiak Bear, which is approximately the same size...

s also den in snow caves to give birth to their young. Both types of dens are protected from exterior temperatures. Formation of these large spaces is from the animals' activity, not ground heat.
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