Stegopelta
Encyclopedia
Stegopelta is a genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of armored dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

. It is based on a partial skeleton from the latest Albian
Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous epoch/series. Its approximate time range is 112.0 ± 1.0 Ma to 99.6 ± 0.9 Ma...

-earliest Cenomanian
Cenomanian
The Cenomanian is, in the ICS' geological timescale the oldest or earliest age of the Late Cretaceous epoch or the lowest stage of the Upper Cretaceous series. An age is a unit of geochronology: it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding...

-age Lower and Upper Cretaceous Belle Fourche Member of the Frontier Formation
Frontier Formation
The Frontier Formation is a sedimentary geological formation whose strata date back to the Late Cretaceous. The formation's extents are: northwest Colorado, southeast Idaho, southern Montana, northern Utah, and western Wyoming. It occurs in many sedimentary basins and uplift areas.The formation...

 of Fremont County, Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

.

History

In 1905, Samuel Wendell Williston
Samuel Wendell Williston
Samuel Wendell Williston was an American educator and paleontologist who was the first to propose that birds developed flight cursorially , rather than arboreally . He was also an entomologist, specialising in Diptera.-Early life:Williston was born in Boston, Massachusetts to Samuel Williston and...

 described FMNH UR88, a partial armored dinosaur skeleton
Skeleton
The skeleton is the body part that forms the supporting structure of an organism. There are two different skeletal types: the exoskeleton, which is the stable outer shell of an organism, and the endoskeleton, which forms the support structure inside the body.In a figurative sense, skeleton can...

 consisting of a maxilla
Maxilla
The maxilla is a fusion of two bones along the palatal fissure that form the upper jaw. This is similar to the mandible , which is also a fusion of two halves at the mental symphysis. Sometimes The maxilla (plural: maxillae) is a fusion of two bones along the palatal fissure that form the upper...

 fragment, seven cervical and two dorsal vertebrae, part of a sacrum
Sacrum
In vertebrate anatomy the sacrum is a large, triangular bone at the base of the spine and at the upper and back part of the pelvic cavity, where it is inserted like a wedge between the two hip bones. Its upper part connects with the last lumbar vertebra, and bottom part with the coccyx...

 and both ilia
Ilium (bone)
The ilium is the uppermost and largest bone of the pelvis, and appears in most vertebrates including mammals and birds, but not bony fish. All reptiles have an ilium except snakes, although some snake species have a tiny bone which is considered to be an ilium.The name comes from the Latin ,...

, caudal vertebrae, parts of the scapula
Scapula
In anatomy, the scapula , omo, or shoulder blade, is the bone that connects the humerus with the clavicle ....

e, both humeral heads
Humerus
The humerus is a long bone in the arm or forelimb that runs from the shoulder to the elbow....

, portions of an ulna
Ulna
The ulna is one of the two long bones in the forearm, the other being the radius. It is prismatic in form and runs parallel to the radius, which is shorter and smaller. In anatomical position The ulna is one of the two long bones in the forearm, the other being the radius. It is prismatic in form...

 and both radii
Radius
In classical geometry, a radius of a circle or sphere is any line segment from its center to its perimeter. By extension, the radius of a circle or sphere is the length of any such segment, which is half the diameter. If the object does not have an obvious center, the term may refer to its...

, a metacarpal, partial tibia
Tibia
The tibia , shinbone, or shankbone is the larger and stronger of the two bones in the leg below the knee in vertebrates , and connects the knee with the ankle bones....

, metatarsal
Metatarsus
The metatarsus or metatarsal bones are a group of five long bones in the foot located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the medial side : the first, second, third, fourth, and fifth...

, and armor
Armour (zoology)
Armour in animals is external or superficial protection against attack by predators, formed as part of the body , usually through the hardening of body tissues, outgrowths or secretions. It has therefore mostly developed in 'prey' species...

 including a shoulder
Shoulder
The human shoulder is made up of three bones: the clavicle , the scapula , and the humerus as well as associated muscles, ligaments and tendons. The articulations between the bones of the shoulder make up the shoulder joints. The major joint of the shoulder is the glenohumeral joint, which...

 spine and neck ring. Unfortunately, it was in poor condition, as it had eroded from a slope and been walked on by cattle
Cattle
Cattle are the most common type of large domesticated ungulates. They are a prominent modern member of the subfamily Bovinae, are the most widespread species of the genus Bos, and are most commonly classified collectively as Bos primigenius...

. Ankylosauria
Ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are first known to have appeared in the early Jurassic Period of...

ns being very poorly known, Williston compared his new genus to Stegosaurus
Stegosaurus
Stegosaurus is a genus of armored stegosaurid dinosaur. They lived during the Late Jurassic period , some 155 to 150 million years ago in what is now western North America. In 2006, a specimen of Stegosaurus was announced from Portugal, showing that they were present in Europe as well...

, and the armor to that of Glyptodon
Glyptodon
Glyptodon was a large, armored mammal of the family Glyptodontidae, a relative of armadillos that lived during the Pleistocene Epoch. It was roughly the same size and weight as a Volkswagen Beetle, though flatter in shape...

; like that mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

, Stegopelta had a fused section of armor (in its case over the pelvis
Pelvis
In human anatomy, the pelvis is the lower part of the trunk, between the abdomen and the lower limbs .The pelvis includes several structures:...

). Moodie redescribed it in 1910, and considered it to be close to, if not the same as, Ankylosaurus
Ankylosaurus
Ankylosaurus is a genus of ankylosaurid dinosaur, containing one species, A. magniventris...

.

The genus fell into obscurity. Walter Coombs synonymized it with the more famous but equally poorly-known Nodosaurus
Nodosaurus
Nodosaurus was a genus of herbivorous ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found in North America. Two incomplete specimens have been discovered in Wyoming and Kansas, and no skulls...

in his 1978 redescription of the Ankylosauria. It was reinstated as a valid genus by Ken Carpenter
Kenneth Carpenter
Kenneth Carpenter is a paleontologist. He is the museum director of the USU Eastern Prehistoric Museum and author or co-author of a number of books on dinosaurs and Mesozoic life...

 and James Kirkland (1998), who recognized it as having distinct vertebral and armor characteristics. Tracy Ford took this farther in 2000, assigning it to a new subfamily in Ankylosauridae
Ankylosauridae
An ankylosaurid is a member of the Ankylosauridae family of armored dinosaurs that evolved 125 million years ago and became extinct 65 million years ago during the Cretaceous-Tertiary extinction event...

 based on armor characteristics, which he called Stegopeltinae. Also included was Glyptodontopelta
Glyptodontopelta
Glyptodontopelta is a genus of dinosaur from the Late Cretaceous. It was an ankylosaur, an armored dinosaur....

. This has not been generally accepted, but most recent reviews have accepted Stegopelta as a distinct genus with uncertain affinities.

Paleobiology

Because it is so poorly known, at this point all that can be said about the habits and life of Stegopelta is that it was a slow quadruped
Quadruped
Quadrupedalism is a form of land animal locomotion using four limbs or legs. An animal or machine that usually moves in a quadrupedal manner is known as a quadruped, meaning "four feet"...

al herbivore
Herbivore
Herbivores are organisms that are anatomically and physiologically adapted to eat plant-based foods. Herbivory is a form of consumption in which an organism principally eats autotrophs such as plants, algae and photosynthesizing bacteria. More generally, organisms that feed on autotrophs in...

 that fed low to the ground and relied on its armor for defense.

Its armor included a fused region over the sacrum, and shoulder spines that may have been split, as seen in Edmontonia
Edmontonia
Edmontonia was an armoured dinosaur, a part of the nodosaur family from the Late Cretaceous Period. It is named after the Edmonton Formation , the unit of rock it was found in.-Description:...

.
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