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Squamata

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Squamata



 
 
Squamata, or the scaled reptiles, is the largest recent order
Order (biology)

In Biological classification used in biology, the order is a taxonomic rank between class and family . The superorder is a rank between class and order....
 of reptile
Reptile

Reptiles, or members of the class Reptilia, are air-breathing, cold-blooded vertebrates that have skin covered in scale as opposed to hair or feathers....
s, including lizard
Lizard

Lizards are a large and widespread group of squamate reptiles, with nearly 5,000 species, ranging across all continents except Antarctica as well as most oceanic island chains....
s and snake
Snake

Snakes are elongate legless carnivore reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears....
s. Members of the order are distinguished by their skins, which bear horny scales
Scale (zoology)

In most biology nomenclature, a scale is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran species, scales are plates on the surface of the insect wing, and provide coloration....
 or shields. They also possess movable quadrate bone
Quadrate bone

The quadrate bone is part of a skull in most tetrapods, including amphibians, sauropsids and early synapsids. In these animals it connects to the quadratojugal and squamosal in the skull, and forms part of the jaw joint ....
s, making it possible to move the upper jaw relative to the braincase. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey.






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Squamata, or the scaled reptiles, is the largest recent order
Order (biology)

In Biological classification used in biology, the order is a taxonomic rank between class and family . The superorder is a rank between class and order....
 of reptile
Reptile

Reptiles, or members of the class Reptilia, are air-breathing, cold-blooded vertebrates that have skin covered in scale as opposed to hair or feathers....
s, including lizard
Lizard

Lizards are a large and widespread group of squamate reptiles, with nearly 5,000 species, ranging across all continents except Antarctica as well as most oceanic island chains....
s and snake
Snake

Snakes are elongate legless carnivore reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears....
s. Members of the order are distinguished by their skins, which bear horny scales
Scale (zoology)

In most biology nomenclature, a scale is a small rigid plate that grows out of an animal's skin to provide protection. In lepidopteran species, scales are plates on the surface of the insect wing, and provide coloration....
 or shields. They also possess movable quadrate bone
Quadrate bone

The quadrate bone is part of a skull in most tetrapods, including amphibians, sauropsids and early synapsids. In these animals it connects to the quadratojugal and squamosal in the skull, and forms part of the jaw joint ....
s, making it possible to move the upper jaw relative to the braincase. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. They are the most variably-sized order of reptiles, ranging from the 16 mm
Millimetre

The millimetre is a Units of measurement of length in the metric system, equal to one thousandth of a metre, which is the current International System of Units SI base unit of length....
 (0.63 in
Inch

An inch is the name of a Units of measurement of length in a number of different systems, including Imperial units, and United States customary units....
) Jaragua Sphaero, Sphaerodactylus ariasae to the 8 m
Metre

The metre or meter is a Unit of measurement of length. It is the SI base unit of length in the metric system and in the International System of Units , used around the world for general and scientific purposes....
 (26 ft) green anaconda
Green Anaconda

Eunectes murinus is a non-venomous Boinae species found in South America. It is known as one of the largest of all snakes. Two subspecies are currently recognized, including the nominate subspecies described here....
, Eunectes murinus.

Classification

Classically, the order is divided into three suborders:

  • Lacertilia, the lizards;
  • Serpentes, the snakes;
  • Amphisbaenia
    Amphisbaenia

    The Amphisbaenia are a suborder of usually legless squamates closely related to lizards and snakes. As many species possess a pink body coloration and scales arranged in rings, they have a superficial resemblance to earthworms....
    , the worm lizards.


Of these, the lizards form a paraphyletic
Paraphyly

In phylogenetics, a group of organisms is said to be paraphyletic if the group contains its most recent common ancestor Common descent but does not contain all the descendants of that ancestor....
 group. In newer classifications the name Sauria
Sauria

Sauria is a clade of reptiles that includes all living diapsids, as well as their common ancestor and all its extinct descendants. The ancestral saurian was probably a small lizard-like creature living in the Permian Period....
 is used for reptiles and birds in general, and the Squamata are divided differently:

  • Suborder Iguania
    Iguania

    Iguania is the suborder of Squamata that contains the iguanas, chameleons, agamids, and "New World lizards" such as anoles and Phrynosomatidae....
     (the agamid
    Agamidae

    Agamids, lizards of the family Agamidae, include more than 300 species in Africa, Asia, Australia, and a few in Southern Europe. phylogenetics they may be sister to the Iguanidae, and have a similar appearance....
    s, chameleon
    Chameleon

    Chameleons are a distinctive and highly specialized clade of lizards. They are distinguished by their parrot-like zygodactylous feet, their separately mobile and stereoscopic eyes, their very long, highly modified, and rapidly extrudable tongues, their swaying gait, and the possession by many of a prehensile tail, crests or horns on their...
    s, iguanid
    Iguanidae

    The Iguanidae is a family of lizards, composed of iguanas and related species....
    s. and other New World
    New World

    The New World is one of the names used for the non-Eurasian/non-African parts of the Earth, specifically the Americas and Australasia. When the term originated in the late 15th century, the Americas were new to the Europeans, who previously thought of the world as consisting only of Europe, Asia, and Africa ....
     lizards)
  • Suborder Scleroglossa
    • Infraorder Gekkota
      Gekkota

      Gekkota is an infraorder in the suborder Scleroglossa, comprising all geckos and the legless Pygopodidae....
       (the gecko
      Gecko

      Geckos are small to average sized lizards belonging to the family Gekkonidae which are found in warm climates throughout the world. Geckos are unique among lizards in their vocalizations, making chirping sounds in social interactions with other geckos....
      s)
    • Infraorder Anguimorpha
      Anguimorpha

      Anguimorphs of the infraorder Anguimorpha include the Anguidae , monitor lizards, mosasaurs, and Heloderma . The infraorder was named by F?rbringer in 1900 to include all autarchoglossans closer to Varanus and Anguis than Scincus....
       (the monitors
      Monitor lizard

      Monitor lizards or biawak are members of the family Varanidae, a group of carnivorous lizard which includes the heaviest living lizard, the Komodo dragon, with the crocodile monitor being the longest in the world....
      , Gila monster
      Gila monster

      The Gila monster , Heloderma suspectum, is a species of venom lizard native to the southwestern United States and northern Mexico . A heavy, slow-moving lizard, up to long, the Gila monster is the only venomous lizard native to the United States, and one of only two known species of venomous lizards in North America, the other being its...
      , alligator lizard
      Alligator lizard

      Alligator Lizard may refer to one of the following:Species of the genus Elgaria:*Northern Alligator Lizard*Southern Alligator LizardSpecies of the genus Gerrhonotus....
      s, galliwasps, slow-worms and others)
    • Infraorder Scincomorpha
      Scincomorpha

      Scincomorpha is an infraorder of saurians....
       (skink
      Skink

      Skinks are the most diverse group of lizards. They comprise the family Scincidae which shares the superfamily or infraorder Scincomorpha with several other lizard families, including Lacertidae ....
      s, whiptail lizards and common European lizards)
  • Suborder Serpentes (the snake
    Snake

    Snakes are elongate legless carnivore reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears....
    s)
    • Infraorder Alethinophidia
      Alethinophidia

      The Alethinophidia are an infraorder of snakes that includes all snakes other than blind snakes and thread snakes. Currently, 15 families are recognized, including 9 subfamilies and 316 genera....
       (viper
      Viperidae

      The Viperidae are a family of venomous snakes found all over the world, except in Australia and Madagascar. All have relatively long hinged fangs that permit deep penetration and injection of venom....
      s, boas, cobra
      Naja

      Naja is a genus of venom elapid snakes. They are the most recognized, and most widespread group of snakes commonly known as cobras, though there are several other genera which also make use of that common name....
      s, etc.)
    • Infraorder Scolecophidia
      Scolecophidia

      The Scolecophidia are an infraorder of snakes. They range in size from 10-100 cm in length, but may only be as small as 2 mm. All are fossorial....
       (blind snakes)
  • Suborder Amphisbaenia
    Amphisbaenia

    The Amphisbaenia are a suborder of usually legless squamates closely related to lizards and snakes. As many species possess a pink body coloration and scales arranged in rings, they have a superficial resemblance to earthworms....


The relationships between these suborders is not yet certain, though recent research suggests that several families may form a hypothetical venom clade which encompasses a majority (nearly 60%) of Squamate species. Named Toxicofera
Toxicofera

Toxicofera , is a hypothesis clade which represents about 4600 species of extant taxon squamates It encompasses all Venom reptile species, as well as numerous related non-venomous species....
, it combines the following groups from traditional classification
Scientific classification

Biological classification or scientific classification in biology, is a method by which biologists group and categorize species of organisms....
:

  • Suborder Serpentes (snakes)
  • Suborder Iguania
    Iguania

    Iguania is the suborder of Squamata that contains the iguanas, chameleons, agamids, and "New World lizards" such as anoles and Phrynosomatidae....
     (agamids, chameleons, iguanids, etc.)
    • Infraorder Anguimorpha
      Anguimorpha

      Anguimorphs of the infraorder Anguimorpha include the Anguidae , monitor lizards, mosasaurs, and Heloderma . The infraorder was named by F?rbringer in 1900 to include all autarchoglossans closer to Varanus and Anguis than Scincus....
      , consisting of:
      • Family Varanidae (monitor lizards, including the Komodo dragon)
      • Family Anguidae
        Anguidae

        The Anguidae is a large and diverse family of lizards native to the northern hemisphere. The group includes the slowworms, glass lizards, and alligator lizards, among others....
         (alligator lizards, glass lizards, etc.)
      • Family Helodermatidae (Gila monster and Mexican beaded lizard)


List of Families

Tropical worm lizards[[Darwin's worm lizard]] (Amphisbaena darwinii) -
[[Bipedidae]]
[[Richard Taylor|Taylor]], 1951
Bipes worm lizards[[Mexican mole lizard]] (Bipes biporus) -
[[Rhineuridae]]
[[Paulo Vanzolini|Vanzolini]], 1951
North American worm lizards[[Rhineuridae|North American worm lizard]] (Rhineura floridana)
Amphisbaenia 1
[[Trogonophidae]]
[[John Edward Gray|Gray]], 1865
Palearctic worm lizards[[Checkerboard worm lizard]] (Trogonophis wiegmanni) -
[[Diploglossa]]
FamilyCommon NamesExample SpeciesExample Photo
[[Anguidae]]
[[Nicolaus Michael Oppel|Oppel]], 1811
Glass lizards[[Anguis fragilis|Slow worm]] (Anguis fragilis)
Anguidae
[[Anniellidae]]
[[John Edward Gray|Gray]], 1852
American legless lizards[[American legless lizard|California legless lizard]] (Anniella pulchra)
Anniella Pulchra
[[Xenosauridae]]
[[Edward Drinker Cope|Cope]], 1866
Knob-scaled lizards[[Chinese crocodile lizard|Chinese crocodile lizard]] (Shinisaurus crocodilurus)
Chin Krokodilschwanzechse 01
[[Gekkota]]
FamilyCommon NamesExample SpeciesExample Photo
[[Dibamidae]]
[[George Albert Boulenger|Boulenger]], 1884
Blind lizards[[Dibamus nicobaricum]] -
[[Gekkonidae]]
[[John Edward Gray|Gray]], 1825
Geckos[[Underwoodisaurus milii|Thick-tailed gecko]] (Underwoodisaurus milii)
Underwoodisaurus Milii
[[Pygopodidae]]
[[George Albert Boulenger|Boulenger]], 1884
Legless lizards[[Lialis burtonis|Burton's snake lizard]] (Lialis burtonis) -
[[Iguania]]
FamilyCommon NamesExample SpeciesExample Photo
[[Agamidae]]
[[Johann Baptist von Spix|Spix]], 1825
Agamas [[Eastern bearded dragon]] (Pogona barbata)
Bearded Dragon04
[[Chamaeleonidae]]
[[John Edward Gray|Gray]], 1825
Chameleons[[Veiled chameleon]] (Chamaeleo calyptratus)
Chamaelio Calyptratus
[[Corytophanidae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Casquehead lizards[[Plumed basilisk|Plumed basilisk]] (Basiliscus plumifrons)
Plumedbasiliskcele4 Edit
[[Crotaphytidae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Collared and leopard lizards[[Common collared lizard]] (Crotaphytus collaris)
Collared Lizard in Zion National Park
[[Hoplocercidae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Wood lizards or clubtails[[Club-tail iguana]] (Hoplocercus spinosus) -
[[Iguanidae]]Iguanas[[Marine iguana]] (Amblyrhynchus cristatus)
Marineiguana03
[[Leiosauridae]]
[[D.R. Frost|Frost]] et al., 2001
- [[Darwin's iguana]] (Diplolaemus darwinii) -
[[Opluridae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Madagascan iguanas [[Chalarodon]] (Chalarodon madagascariensis) -
[[Phrynosomatidae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Earless, spiny, tree, side-blotched and horned lizards[[Greater earless lizard|Greater earless lizard]] (Cophosaurus texanus)
Reptile Tx Usa
[[Polychrotidae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Anoles[[Carolina anole|Carolina anole]] (Anolis carolinensis)
[[Tropiduridae]]
[[D.R. Frost|Frost]] & [[R. Etheridge|Etheridge]], 1989
Neotropical ground lizards([[Microlophus peruvianus]])
Mperuvianus
[[Platynota]]
FamilyCommon NamesExample SpeciesExample Photo
[[Helodermatidae]]Gila monsters[[Gila monster|Gila monster]] (Heloderma suspectum)
Gila
[[Lanthanotidae]]Earless monitor[[Lanthanotidae|Earless monitor]] (Lanthanotus borneensis) -
[[Varanidae]]Monitor lizards[[Perentie]] (Varanus giganteus)
Perentie Lizard Perth Zoo Smc Spet 2005
[[Scincomorpha]]
FamilyCommon NamesExample SpeciesExample Photo
[[Cordylidae]]Spinytail lizards [[Girdle-tailed lizard]] (Cordylus warreni)
Cordylus Breyeri1
[[Gerrhosauridae]]Plated lizards[[Sudan plated lizard|Sudan plated lizard]] (Gerrhosaurus major)
[[Gymnophthalmidae]]Spectacled lizards - -
[[Lacertidae]]
[[Nicolaus Michael Oppel|Oppel]], 1811
Wall or true lizards[[Ocellated lizard]] (Lacerta lepida)
Perleidechse 20
[[Scincidae]]
[[Nicolaus Michael Oppel|Oppel]], 1811
Skinks[[Western blue-tongued lizard|Western blue-tongued skink]] (Tiliqua occipitalis)
Tiliqua Occipitalis
[[Teiidae]]Tegus or whiptails[[Blue tegu]] (Tupinambis teguixin)
Goldteju Tupinambis Teguixin
[[Xantusiidae]]Night lizards[[Granite night lizard]] (Xantusia henshawi)
Xantusia Henshawi
[[Alethinophidia]]
FamilyCommon NamesExample SpeciesExample Photo
[[Acrochordidae]]
[[Charles Lucien Bonaparte|Bonaparte]], 1831
File snakes[[Acrochordus granulatus|Marine file snake]] (Acrochordus granulatus)
Wart Snake 1
[[Aniliidae]]
[[Leonhard Hess Stejneger|Stejneger]], 1907
Coral pipe snakes[[Anilius scytale|Burrowing false coral]] (Anilius scytale) 
[[Anomochilidae]]
Cundall, Wallach and [[Douglas Athon Rossman|Rossman]], 1993.
Dwarf pipe snakes[[Anomochilus leonardi|Leonard's pipe snake]], (Anomochilus leonardi) 
[[Atractaspididae]]
[[Albert C. L. G. Günther|Günther]], 1858
Mole vipersBibron's burrowing asp (Atractaspis bibroni)  
Boidae
Gray
John Edward Gray

John Edward Gray was a United Kingdom zoology. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....
, 1825
BoasAmazon tree boa
Amazon tree boa

Corallus hortulanus is a non-venomous Boinae species found in South America. No subspecies are currently recognized....
 (Corallus hortulanus)
Corallushortulanus
Bolyeriidae
Hoffstetter, 1946
Round Island boasRound Island burrowing boa (Bolyeria multocarinata) 
Colubridae
Oppel
Nicolaus Michael Oppel

Nicolaus Michael Oppel was a Germany natural history. He was a student of, and worked as an assistant to, Andr? Marie Constant Dum?ril at the Mus?um national d'Histoire naturelle in Paris, France, cataloging and classifying species of reptile....
, 1811
ColubridsGrass snake
Grass Snake

The Grass Snake , sometimes called the Ringed Snake or Water Snake is a European non-venom snake.The Grass Snake is typically dark green or brown in colour with a characteristic yellow collar behind the head, which explains the alternative name ringed snake....
 (Natrix natrix)
Natrix Natrix (marek Szczepanek)
Cylindrophiidae
Fitzinger
Leopold Fitzinger

Leopold Joseph Franz Johann Fitzinger was an Austrian zoologist.Fitzinger was born in Vienna and studied botany at the university of Vienna under Nikolaus Joseph von Jacquin....
, 1843
Asian pipe snakesRed-tailed pipe snake
Red-tailed Pipe Snake

Cylindrophis ruffus is a snake species found in Southeast Asia. No subspecies are currently recognized....
 (Cylindrophis ruffus)
Cylindrophis Rufus
Elapidae
Elapidae

The Elapidae, or elapids, are a family of venom snakes found in tropical and subtropical regions around the world, including the Indian Ocean and the Pacific....

Boie
Friedrich Boie

Friedrich Boie was a Germany scientist and brother of Heinrich Boie. He was born at Meldorf in Holstein and died at Kiel.Boie was the author of Bemerkungen ?ber Merrem's Versuch eines Systems der Amphibien ....
, 1827
Cobras, coral snakes, mambas, kraits, sea snakes, sea kraits, Australian elapidsKing cobra
King Cobra

The King Cobra is the world's longest venom snake, with a length that can be as large as 5.7 m . This species is widespread throughout Southeast Asia and parts of India, but found mostly in forested areas....
 (Ophiophagus hannah)
Ophiophagus Hannah2
Loxocemidae
Loxocemidae

The Loxocemidae are a monotypic Family of snakes created for the monotypic genus Loxocemus that contains the species L. bicolor found in Mexico....

Cope
Edward Drinker Cope

Edward Drinker Cope was an United States paleontology and comparative anatomy, as well as a noted herpetology and ichthyology.Born to a wealthy Society of Friends family, Cope quickly distinguished himself as a child prodigy interested in science; he published his first scientific paper in 1859....
, 1861
Mexican burrowing snakesMexican burrowing snake (Loxocemus bicolor)
Loxocemus Bicolor
Pythonidae
Fitzinger
Leopold Fitzinger

Leopold Joseph Franz Johann Fitzinger was an Austrian zoologist.Fitzinger was born in Vienna and studied botany at the university of Vienna under Nikolaus Joseph von Jacquin....
, 1826
PythonsBall python
Ball python

Python regius is a non-venomous Pythonidae species found in Africa. This is the smallest of the African pythons and is popular in the pet trade....
 (Python regius)
Tropidophiidae
Brongersma, 1951
Dwarf boasNorthern eyelash boa
Trachyboa

Trachyboa is a genus of Tropidophiidaes found in Central America and South America. They are largely Terrestrial animal fish-eating snakes that inhabit tropical lowlands....
 (Trachyboa boulengeri)
 
Uropeltidae
Müller
Johannes Peter Müller

Johannes Peter M?ller , was a Germany physiologist, comparative anatomy, and ichthyology not only known for his discoveries but also for his ability to synthesize knowledge....
, 1832
Shield-tailed snakes, short-tailed snakesCuvier's shieldtail (Uropeltis ceylanica)
Silybura Shortii
Viperidae
Viperidae

The Viperidae are a family of venomous snakes found all over the world, except in Australia and Madagascar. All have relatively long hinged fangs that permit deep penetration and injection of venom....

Oppel
Nicolaus Michael Oppel

Nicolaus Michael Oppel was a Germany natural history. He was a student of, and worked as an assistant to, Andr? Marie Constant Dum?ril at the Mus?um national d'Histoire naturelle in Paris, France, cataloging and classifying species of reptile....
, 1811
Vipers, pitvipers, rattlesnakesEuropean asp
Vipera aspis

Vipera aspis is a venomous snake Viperinae species found in southwestern Europe. Bites from this species can be more severe than from the European adder, Vipera berus; not only can they be very painful, but according to Stemmler , about 4% of all untreated bites are fatal....
 (Vipera aspis)
Vipera Aspis Aspis 1
Xenopeltidae
Xenopeltidae

The Xenopeltidae are a monotypic Family of snakes created to the genus Xenopeltis, which is found in Southeast Asia. Its members are known for their highly iridescent scales....

Bonaparte
Charles Lucien Bonaparte

Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano was a France natural history and ornithology. He was the son of Lucien Bonaparte and Alexandrine de Bleschamp, and nephew of Emperor Napoleon I of France....
, 1845
Sunbeam snakesSunbeam snake
Xenopeltis unicolor

Xenopeltis unicolor is a non-venomous Xenopeltidae species found in Southeast Asia. This is a primitive snake known for its highly iridescent scales....
 (Xenopeltis unicolor)
Scolecophidia
Scolecophidia

The Scolecophidia are an infraorder of snakes. They range in size from 10-100 cm in length, but may only be as small as 2 mm. All are fossorial....
FamilyCommon NamesExample SpeciesExample Photo
Anomalepidae
Taylor
Edward Harrison Taylor

Edward Harrison Taylor was an United States herpetologist from Kansas.He was born in Maysville, Missouri and studied at the University of Kansas in Lawrence, Kansas, graduating with a Bachelor's degree in 1912....
, 1939
Dawn blind snakesDawn blind snake (Liotyphlops beui) 
Leptotyphlopidae
Leptotyphlopidae

The Leptotyphlopidae are a family of snakes found in North America and South America, Africa, and Asia. All are fossorial and adapted to burrowing, feeding on ants and termites....

Stejneger
Leonhard Hess Stejneger

Leonhard Hess Stejneger was a zoologist.Stejneger was born in Bergen, Norway, Norway. His father was Peter Stamer Steineger who was educated in private schools and was a merchant and auditor and his mother was Ingeborg...
, 1892
Slender blind snakesTexas blind snake
Texas Blind Snake

Leptotyphlops dulcis is a Leptotyphlopidae species found in the southwestern United States and northern Mexico. Three subspecies are currently recognized, including the nominate subspecies described here....
 (Leptotyphlops dulcis)
Leptotyphlops Dulcis
Typhlopidae
Typhlopidae

The Typhlopidae are a family of blind snakes. They are found mostly in the tropical regions of Africa, Asia and the Americas. The rostral scale overhangs the mouth to form a shovel like burrowing structure....

Merrem
Blasius Merrem

Blasius Merrem was a Germany natural history.Merrem was born at Bremen , and studied at the university of Göttingen under Johann Friedrich Blumenbach....
, 1820
Blind snakesBlack blind snake (Typhlops reticulatus) 


Evolution

Squamates are a monophyletic group that is a sister group to the tuatara
Tuatara

The tuatara is a reptile endemism to New Zealand which, though it resembles most lizards, is actually part of a distinct lineage, order Sphenodontia....
. The squamates and tuatara together are a sister group to crocodiles and birds, the extanct archosaurs. Squamate fossils first appear in the early Jurassic
Jurassic

The Jurassic is a geologic period that extends from about annum to  Ma, that is, from the end of the Triassic to the beginning of the Cretaceous....
, but a mitochondrial phylogeny suggests that they evolved in the late Permian
Permian

The PermianThe term "Permian" was introduced into geology in 1841 by Sir Roderick Murchison, president of the Geological Society of London, who identified typical strata in extensive Russian explorations undertaken with Edouard de Verneuil; Murchison asserted in 1841 that he named his "Permian system" after the ancient kingdom...
. The evolutionary relationships within the squamates are not yet completely worked out, with the relationship of snakes to other groups being most problematic. From morphological data, Iguanid lizards have been thought to have diverged from other squamates very early, but recent molecular phylogenies
Molecular phylogeny

Molecular phylogenetics, also known as molecular systematics, is the use of the structure of molecules to gain information on an organism's evolutionary relationships....
, both from mitochondrial
Mitochondrial DNA

Mitochondrial DNA is the DNA located in organelles called mitochondrion. Most other DNA present in eukaryotic organisms is found in the cell nucleus....
 and nuclear DNA
DNA

Deoxyribonucleic acid is a nucleic acid that contains the genetics instructions used in the development and functioning of all known living organisms and some viruses....
, do not support this early divergence. Because snakes have a faster molecular clock
Molecular clock

The molecular clock is a technique in molecular evolution to relate the time that two species speciation to the number of molecular differences measured between the species' DNA sequences or proteins....
 than other squamates, and there are few early snake and snake ancestor fossils, it is difficult to resolve the relationship between snakes and other squamate groups.

Reproduction

The male members of the group Squamata have a hemipenis
Hemipenis

A hemipenis is one of a pair of intromittent organs of male squamatas .Hemipenes are usually held inverted, within the body, and are everted for reproduction via erectile tissue like that in the human penis....
. Hemipenes are usually held inverted, within the body, and are everted for reproduction via erectile tissue
Erectile tissue

Erectile tissue is tissue in the body that can become erect, usually by becoming engorged with blood....
 like that in the human penis
Penis

The penis is an external sex organ of certain biologically male organisms, in both vertebrates and invertebrates.The penis is a reproductive organ, technically an intromittent organ, and for Eutheria, additionally serves as the external organ of urination....
. Only one is used at a time, and some evidence indicates males alternate use between copulations. The hemipenis itself has a variety of shapes, depending on species. Often the hemipenis bears spines or hooks, in order to anchor the male within the female. Some species even have forked hemipenes (each hemipenis has two tips). Due to being everted and inverted, hemipenes do not have a completely enclosed channel for the conduction of sperm
Sperm

The term sperm is derived from the Greek word sperma and refers to the male reproductive Cell . In the types of sexual reproduction known as anisogamy and oogamy, there is a marked difference in the size of the gametes with the smaller one being termed the "male" or sperm cell....
, but rather a seminal groove which seals as the erectile tissue expands. This is also the only reptile group in which can be found both viviparous and ovoviviparous
Ovoviviparity

Ovoviviparous, also known as oviviparous, animals develop within Egg s that remain within the mother's body up until they hatch or are about to hatch....
 species, as well as the usual oviparous reptiles. Some species, like the Komodo dragon
Komodo dragon

The Komodo dragon is a species of lizard that inhabits the islands of Komodo , Rinca, Flores, and Gili Motang in Indonesia. A member of the monitor lizard family , it is the Largest organisms#Reptiles , growing to an average length of and weighing around ....
, can actually reproduce asexual
Asexual reproduction

Asexual reproduction is reproduction which does not involve meiosis, ploidy reduction, or fertilization. Only one parent is involved in asexual reproduction....
ly and undergo parthenogenesis
Parthenogenesis

Parthenogenesis is an asexual form of reproduction found in females where growth and development of embryos or seeds occurs without fertilization by a male....
.

Venom

Venom is modified saliva
Saliva

Saliva is the watery and usually frothy substance produced in the mouths of humans and most other animals. Saliva is produced in and secreted from the salivary glands....
, delivered through fang
Snake venom

Snake venom is highly modified saliva that is produced by special glands of certain species of snakes. The gland which secretes the zootoxin is a modification of the parotid gland of other vertebrates, and is usually situated on each side of the head below and behind the eye, invested in a muscular sheath....
s. The fangs of 'advanced' venomous snakes like viperids and elapids are hollow in order to inject venom more effectively, while the fangs of rear-fanged snakes such as the Boomslang merely have a groove on the posterior edge to channel venom into the wound. Snake venoms are often prey specific, its role in self-defense is secondary. Venom, like all salivary secretions, is a pre-digestant which initiates the breakdown of food into soluble compounds allowing for proper digestion and even "non-venomous" snake bites (like any animal bite) will cause tissue damage.

Recent research suggests that the evolutionary origin of venom may exist deep in the squamate phylogeny, with 60% of squamates placed in this hypothetical group called Toxicofera
Toxicofera

Toxicofera , is a hypothesis clade which represents about 4600 species of extant taxon squamates It encompasses all Venom reptile species, as well as numerous related non-venomous species....
. Venom has been known in the families Heloderma
Heloderma

Heloderma, the only genus of the family Helodermatidae, consists of venom lizards native to the southwestern United States, Mexico and as far south as Guatemala....
tidae, Elapidae
Elapidae

The Elapidae, or elapids, are a family of venom snakes found in tropical and subtropical regions around the world, including the Indian Ocean and the Pacific....
, Viperidae
Viperidae

The Viperidae are a family of venomous snakes found all over the world, except in Australia and Madagascar. All have relatively long hinged fangs that permit deep penetration and injection of venom....
, and some members of the Colubridae. However, all snake
Snake

Snakes are elongate legless carnivore reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears....
s, some agamid lizards and most monitor lizard
Monitor lizard

Monitor lizards or biawak are members of the family Varanidae, a group of carnivorous lizard which includes the heaviest living lizard, the Komodo dragon, with the crocodile monitor being the longest in the world....
s are now believed by some to have proteins very closely related to venom.

Humans and Squamates


Bites and fatalities

Snakebite Morbidity Map
In the US alone, more than 8,000 venomous snake bites are reported each year. It is estimated that 125,000 people a year die from venomous snake bites. In addition, large pet constrictors, like boas and pythons, have been known to kill their owners through strangulation on rare occasions.

Lizard bites, unlike venomous snake bites, are not fatal. The Komodo dragon
Komodo dragon

The Komodo dragon is a species of lizard that inhabits the islands of Komodo , Rinca, Flores, and Gili Motang in Indonesia. A member of the monitor lizard family , it is the Largest organisms#Reptiles , growing to an average length of and weighing around ....
 has been known to kill people due to its size. The two known venomous species of lizard, the Gila monster
Gila monster

The Gila monster , Heloderma suspectum, is a species of venom lizard native to the southwestern United States and northern Mexico . A heavy, slow-moving lizard, up to long, the Gila monster is the only venomous lizard native to the United States, and one of only two known species of venomous lizards in North America, the other being its...
 and Mexican beaded lizard have never caused a human death by envenomation.

Conservation

Even though they survived the worst changes in Earth
Earth

Earth is the third planet from the Sun. Earth is the largest of the terrestrial planets in the Solar System in diameter, mass and density. It is also referred to as the World and Wiktionary:Terra.Note that by International Astronomical Union convention, the term "Terra" is used for naming extensive land masses, rather...
's history, today many squamate species are endangered due to habitat loss, hunting and poaching, the pet trade, alien species being introduced to their habitat (which puts native creatures at risk through unfair competition & predation), and many other unnecessary reasons. Because of this, some are in fact extinct with Africa having the most extinct species of squamates
List of extinct animals of Africa

The list of extinct animals in Africa features the animals that have become extinct on the Africa and its islands, like Madagascar, Mauritius, Rodrigues, R?union, Seychelles, Saint Helena, Cape Verde, etc....
. However, breeding programs and wildlife parks are trying to save many endangered reptiles from extinction. Many zoos & breeders educate people about the importance of snake
Snake

Snakes are elongate legless carnivore reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears....
s and lizard
Lizard

Lizards are a large and widespread group of squamate reptiles, with nearly 5,000 species, ranging across all continents except Antarctica as well as most oceanic island chains....
s.

Cited references


External links