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Solanum
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- "Horsenettle" and variants redirect here. If used for a particular species, this name usually applies to the Carolina Horsenettle (S. carolinense).
- For the "Solanum Virus", see The Zombie Survival Guide.
Solanum, the nightshades, horsenettles and relatives, is a large and diverse genus of annual and perennial plants. They grow as forbs, vines, sub-shrubs, shrubs, and small trees, and often have attractive fruit and flowers.

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Encyclopedia
- "Horsenettle" and variants redirect here. If used for a particular species, this name usually applies to the Carolina Horsenettle (S. carolinense).
- For the "Solanum Virus", see The Zombie Survival Guide.
Solanum, the nightshades, horsenettles and relatives, is a large and diverse genus of annual and perennial plants. They grow as forbs, vines, sub-shrubs, shrubs, and small trees, and often have attractive fruit and flowers. Many formerly independent genera like Lycopersicon (the tomatoes) or Cyphomandra are included in Solanum as subgenera or sections today. Thus, the genus nowadays contains roughly 1,500-2,000 species.
In particular, the green parts, including unripe fruit, are often poisonous to humans (albeit not necessarily to other animals), many bear edible fruits, leaves, or tubers. Several species are cultivated, including three globally important food crops:
- Tomato, S. lycopersicum
- Potato, S. tuberosum
- Eggplant, S. melongena
Other species are significant food crops regionally, such as Ethiopian Eggplant and gilo (S. aethiopicum), naranjilla or lulo (S. quitoense), Turkey Berry (S. torvum), or the "bush tomatoes" (several Australian species).
While most medical relevance of Solanum is due to poisonings which are not uncommon and may be fatal, several species are locally used in folk medicine, particularly by native peoples who have long employed them. Giant Devil's-fig (S. chrysotrichum) has been shown to be an effective treatment for seborrhoeic dermatitis in a scientific study.
Solanum species are used as food plants by the larvae of some Lepidoptera species (butterflies and moths) - see list of Lepidoptera that feed on Solanum.
Systematics
The genus was established by Carl Linnaeus in 1753. Its subdivision has always been problematic, but slowly some sort of consensus is being achieved.
The following list is a provisional lineup of the genus' traditional subdivisions, together with some notable species. Many of the subgenera and sections might not be valid; they are used here provisionally as the phylogeny of this genus is not fully resolved yet and many species have not been reevaluated.
Cladistic analyses of DNA sequence data suggests that the present subdivisions and rankings are largely invalid. Far more subgenera would seem to warrant recognition, with Leptostemonum being the only one that can at present be clearly subdivided into sections. Notably, it includes as a major lineage several members of the traditional sections Cyphomandropsis and the old genus Cyphomandra.
Subgenus Bassovia
Section Allophylla
Solanum granuloso-leprosum
Section Cyphomandropsis
- Solanum glaucophyllum Desf. – Waxy-leaved Nightshade
Section Pachyphylla
Solanum betaceum – Tamarillo Solanum exiguum Solanum roseum
Subgenus Leptostemonum
Section Acanthophora
Section Anisantherum
Section Campanulata
Section Crinitum
Section Croatianum
Section Erythrotrichum
Solanum robustum H.L.Wendl. – Shrubby Nightshade
Section Graciliflorum
Section Herposolanum
Solanum wendlandii Hook.f. – Giant Potatocreeper
Section Irenosolanum
- Solanum incompletum Dunal – Popolo ku mai (Hawaii)
- Solanum nelsonii – Nelson's Horsenettle, Akia (Hawaii)
- Solanum sandwicense Hook. & Arn. – Hawaiian Horsenettle, Popoloaiakeakua (Oahu, Kauai)
Section Ischyracanthum
Section Lasiocarpa
Section Melongena
Solanum aculeastrum – Soda Apple, Sodaapple Nightshade, Goat Apple, Poison Apple, "bitter-apple" Solanum campechiense – Redberry Nightshade Solanum carolinense – Carolina Horsenettle, Radical Weed, Sand Brier, Devil's Tomato, "bull nettle", "tread-softly", "apple of Sodom", "wild tomato" Solanum citrullifolium – Watermelon Nightshade Solanum dimidiatum Raf. – Torrey's Nightshade Solanum elaeagnifolium – Silver-leaved Nightshade, Prairie Berry, Silverleaf Nettle, White Horsenettle, Silver Nightshade, "bull-nettle", "trompillo" (Spanish); Silver-leaf bitter-apple, satansbos (South Africa) Solanum heterodoxum Dunal – Melon-leaved Nightshade Solanum incanum L. Solanum linnaeanum – Devil's Apple, "apple of Sodom" Solanum macrocarpon L. Solanum marginatum L.f. – White-margined Nightshade Solanum melongena – Eggplant, Aubergine (including S. ovigerum) Solanum rostratum Dunal – Buffalo Bur, Texas Thistle Solanum sisymbriifolium – Sticky Nightshade, fire-and-ice Solanum virginianum L.
Section Micracantha
- Solanum jamaicense Mill. – Jamaican Nightshade
- Solanum lanceifolium Jacq. – Lance-leaved Nightshade
- Solanum tampicense Dunal – Wetland Nightshade
Section Monodolichopus
Section Nycterium
Section Oliganthes
Solanum aethiopicum – Ethiopian Eggplant, Nakati, Mock Tomato, Ethiopian Nightshade; including S. gilo (Scarlet Eggplant, Gilo or jiló) Solanum centrale – Australian Desert Raisin, Bush Raisin, Bush Sultana, "bush tomato", akatjurra (Alyawarre), kampurarpa (Pitjantjatjara), merne akatyerre (Arrernte), kutjera Solanum cleistogamum – "bush tomato", merne mwanyerne (Arrernte) Solanum ellipticum – Potato Bush, "bush tomato" Solanum pyracanthon Lam. – Porcupine Tomato, Devil's Thorn Solanum quadriloculatum – "bush tomato", "wild tomato" (Australia)
Section Persicariae
- Solanum bahamense – Bahama Nightshade, Canker Berry, berengena de playa
- Solanum ensifolium – Erubia
Section Polytrichum
Section Pugiunculifera
Section Somalanum
Section Torva
Subgenus Lyciosolanum
Subgenus Solanum sensu stricto
Section Afrosolanum
Section Anarrhichomenum
Section Archaesolanum
Solanum aviculare – Poroporo (New Zealand), Kangaroo Apple (Australia)
Section Basarthrum
Section Benderianum
Section Brevantherum
- Solanum bullatum
- Solanum erianthum D.Don – Potato Tree, "Mullein Nightshade"
- Solanum mauritianum – Woolly Nightshade, Ear-leaved Nightshade, Flannel Weed, Bugweed, Tobacco Weed, Kerosene Plant, "wild tobacco" (Austrialia)
Section Dulcamara
Solanum crispum – Chilean Potato Vine, Chilean Nightshade, Chilean Potato Tree Solanum dulcamara – Bittersweet Solanum imbaburense Solanum laxum Spreng. – Jasmine Nightshade Solanum leiophyllum Solanum seaforthianum Andrews – Brazilian Nightshade Solanum triquetrum Cav. – Texas Nightshade Solanum wallacei – Wallace's Nightshade, Catalina Nightshade, Clokey's Nightshade, "wild tomato" (including S. clokeyi) Solanum xanti – Purple Nightshade, San Diego Nightshade
Section Herpystichum
Section Holophylla
Solanum diphyllum L. – Twin-leaved nightshade Solanum pseudocapsicum – Jerusalem Cherry, Madeira Winter Cherry, "winter cherry" (including S. capsicastrum) Solanum pseudoquina (including S. inaequale Vell.)
Section Juglandifolia
Section Lemurisolanum
Section Lycopersicoides
- Solanum lycopersicoides Dunal – Peruvian Wolfpeach
Section Lycopersicon
Solanum arcanum Peralta – "wild tomato" Solanum cheesmaniae (L.Riley) Fosberg Solanum galapagense S.C.Darwin & Peralta Solanum huaylasense Peralta Solanum lycopersicum – Tomato Solanum peruvianum L. – Peruvian Nightshade, "wild tomato" Solanum pimpinellifolium – Currant Tomato
Section Macronesiotes
Section Normania
Section Petota
Section Pteroidea
Section Quadrangulare
Section Regmandra
Section Solanum
- Solanum adscendens – Sonoita Nightshade
- Solanum americanum – American Nightshade, American Black Nightshade, Glossy Nightshade
- Solanum chenopodioides Lam. – Goosefoot Nightshade, Slender Nightshade (including S. gracilius)
- Solanum douglasii Dunal – Green-spotted Nightshade
- Solanum interius Rydb.
- Solanum nigrescens M.Martens & Galeotti – Divine Nightshade
- Solanum nigrum – European Black Nightshade, "black nightshade"
- Solanum pseudogracile Heiser – Glowing Nightshade
- Solanum ptychanthum – West Indian Nightshade, Eastern Black Nightshade
- Solanum retroflexum – Wonderberry, Sunberry
- Solanum sarrachoides – Hairy Nightshade
- Solanum scabrum Mill. – Garden Huckleberry
- Solanum triflorum Nutt. – Cut-leaved Nightshade
- Solanum villosum Mill. – Yellow Nightshade
Other notable species
Formerly placed here
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