Sailfin molly
Encyclopedia
The sailfin molly, Poecilia latipinna, is a species of fish, of the genus Poecilia
Poecilia
Poecilia is a genus of euryhaline brackish water fish in family Poeciliidae of order Cyprinodontiformes. The type species is P. vivipara. Live-bearers, the Poecilia species are collectively known as mollies, with the exception of Endler's livebearer and the famous guppy...

. They inhabit fresh
Freshwater
Fresh water is naturally occurring water on the Earth's surface in ice sheets, ice caps, glaciers, bogs, ponds, lakes, rivers and streams, and underground as groundwater in aquifers and underground streams. Fresh water is generally characterized by having low concentrations of dissolved salts and...

, brackish
Brackish water
Brackish water is water that has more salinity than fresh water, but not as much as seawater. It may result from mixing of seawater with fresh water, as in estuaries, or it may occur in brackish fossil aquifers. The word comes from the Middle Dutch root "brak," meaning "salty"...

, and coast
Coast
A coastline or seashore is the area where land meets the sea or ocean. A precise line that can be called a coastline cannot be determined due to the dynamic nature of tides. The term "coastal zone" can be used instead, which is a spatial zone where interaction of the sea and land processes occurs...

al waters from North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

 to Texas
Texas
Texas is the second largest U.S. state by both area and population, and the largest state by area in the contiguous United States.The name, based on the Caddo word "Tejas" meaning "friends" or "allies", was applied by the Spanish to the Caddo themselves and to the region of their settlement in...

 and the Yucatán Peninsula
Yucatán Peninsula
The Yucatán Peninsula, in southeastern Mexico, separates the Caribbean Sea from the Gulf of Mexico, with the northern coastline on the Yucatán Channel...

 of Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

.

Taxonomy

The sailfin molly was originally described in 1821 as Mollienesia latipinna by the naturalist Charles Alexandre Lesueur
Charles Alexandre Lesueur
Charles Alexandre Lesueur was a French naturalist, artist and explorer.Pictured here is the oil portrait by Charles Willson Peale of Charles-Alexandre Lesueur...

. Lesueur based his description upon specimens from freshwater ponds in the vicinity of New Orleans, Louisiana
New Orleans, Louisiana
New Orleans is a major United States port and the largest city and metropolitan area in the state of Louisiana. The New Orleans metropolitan area has a population of 1,235,650 as of 2009, the 46th largest in the USA. The New Orleans – Metairie – Bogalusa combined statistical area has a population...

. However, Lesueur described other collections of the sailfin molly as Mollienesia multilineata in 1821, the same year in which he described M. latipinna. This conflict created confusion and eventually necessitated a ruling by the International Commission on Zoological Nomenclature
International Commission on Zoological Nomenclature
The International Commission on Zoological Nomenclature is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 28 members from 20 countries, mainly practicing zoological taxonomists...

 (ICZN). In 1959, the ICZN placed precedence on the name Mollienesia latipinna Lesueur 1821. In a landmark definitive work on Poeciliid fishes, Donn Rosen and Reeve Bailey (1959) noted the priority of Poecilia by Marcus Elieser Bloch
Marcus Elieser Bloch
Marcus Elieser Bloch was a German medical doctor and naturalist. He is generally considered one of the most important ichthyologists of the 18th century.- Life :...

 and Johann Gottlob Schneider
Johann Gottlob Schneider
Johann Gottlob Theaenus Schneider was a German classicist and naturalist.-Biography:Schneider was born at Collm in Saxony...

 (1801) with regards to Mollienesia by Lesueur (1821), thereby relegating Mollienesia to the synonymy of Poecilia.

Common names

Sailfin molly (English), Breitflossenkärpfling (German
German language
German is a West Germanic language, related to and classified alongside English and Dutch. With an estimated 90 – 98 million native speakers, German is one of the world's major languages and is the most widely-spoken first language in the European Union....

), seilfinnemolly (Norwegian
Norwegian language
Norwegian is a North Germanic language spoken primarily in Norway, where it is the official language. Together with Swedish and Danish, Norwegian forms a continuum of more or less mutually intelligible local and regional variants .These Scandinavian languages together with the Faroese language...

), zeilvinkarper (Dutch
Dutch language
Dutch is a West Germanic language and the native language of the majority of the population of the Netherlands, Belgium, and Suriname, the three member states of the Dutch Language Union. Most speakers live in the European Union, where it is a first language for about 23 million and a second...

), molinezja szerokopłetwa (Polish
Polish language
Polish is a language of the Lechitic subgroup of West Slavic languages, used throughout Poland and by Polish minorities in other countries...

), bubuntis (Tagalog
Tagalog language
Tagalog is an Austronesian language spoken as a first language by a third of the population of the Philippines and as a second language by most of the rest. It is the first language of the Philippine region IV and of Metro Manila...

), and molliénésie á voilure or simply "molly voile" (French
French language
French is a Romance language spoken as a first language in France, the Romandy region in Switzerland, Wallonia and Brussels in Belgium, Monaco, the regions of Quebec and Acadia in Canada, and by various communities elsewhere. Second-language speakers of French are distributed throughout many parts...

).

There is some confusion with the Yucatan molly, P. velifera. While most names that contain a "sail" element refer to the present species, the German Segelkärpfling, the Latin velifera and possibly others are used for the Yucatan molly. The French terms are used for both species indiscriminately. http://widgets.opera.com/ru/category/web-developer/4/y

Geographical distribution

The sailfin molly is found in fresh, brackish, and coastal saltwater in coastal lowland habitats from North Carolina to Texas and the Yucatán Peninsula
Yucatán Peninsula
The Yucatán Peninsula, in southeastern Mexico, separates the Caribbean Sea from the Gulf of Mexico, with the northern coastline on the Yucatán Channel...

 of Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

. Preferring marsh
Marsh
In geography, a marsh, or morass, is a type of wetland that is subject to frequent or continuous flood. Typically the water is shallow and features grasses, rushes, reeds, typhas, sedges, other herbaceous plants, and moss....

es, lowland streams, swamp
Swamp
A swamp is a wetland with some flooding of large areas of land by shallow bodies of water. A swamp generally has a large number of hammocks, or dry-land protrusions, covered by aquatic vegetation, or vegetation that tolerates periodical inundation. The two main types of swamp are "true" or swamp...

s, and estuaries, the sailfin molly is very common in peninsular Florida
Florida
Florida is a state in the southeastern United States, located on the nation's Atlantic and Gulf coasts. It is bordered to the west by the Gulf of Mexico, to the north by Alabama and Georgia and to the east by the Atlantic Ocean. With a population of 18,801,310 as measured by the 2010 census, it...

. Non-indigenous populations are established in New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

, in the western U.S.
Western United States
.The Western United States, commonly referred to as the American West or simply "the West," traditionally refers to the region comprising the westernmost states of the United States. Because the U.S. expanded westward after its founding, the meaning of the West has evolved over time...

 and in Hawaii. Sailfin mollies introduced to California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 have caused a decline in populations of the federally protected and endangered Desert Pupfish (Cyprinodon macularius
Cyprinodon macularius
Cyprinodon macularius is a rare species of fish in the family Cyprinodontidae and is known by the common name desert pupfish. It is a federally listed endangered species in the United States....

).

Habitat

Sailfin mollies are most commonly observed as the shallow surface waters along the edges of marsh
Marsh
In geography, a marsh, or morass, is a type of wetland that is subject to frequent or continuous flood. Typically the water is shallow and features grasses, rushes, reeds, typhas, sedges, other herbaceous plants, and moss....

es, lowland streams, ponds, swamp
Swamp
A swamp is a wetland with some flooding of large areas of land by shallow bodies of water. A swamp generally has a large number of hammocks, or dry-land protrusions, covered by aquatic vegetation, or vegetation that tolerates periodical inundation. The two main types of swamp are "true" or swamp...

s, estuaries
Estuary
An estuary is a partly enclosed coastal body of water with one or more rivers or streams flowing into it, and with a free connection to the open sea....

 and even ephemeral
Ephemeral
Ephemeral things are transitory, existing only briefly. Typically the term is used to describe objects found in nature, although it can describe a wide range of things....

 water bodies such as roadside ditches. Small to large aggregations
Shoaling and schooling
In biology, any group of fish that stay together for social reasons are said to be shoaling , and if, in addition, the group is swimming in the same direction in a coordinated manner, they are said to be schooling . In common usage, the terms are sometimes used rather loosely...

 of the species are most commonly found under floating vegetation or near structures in the water, minimizing their chances of being observed by potential predators.

The sailfin molly is a tolerant
Physiological tolerance
Physiological tolerance or drug tolerance is commonly encountered in pharmacology, when a subject's reaction to a drug is reduced at a later time even though the dose or concentration at the effect site is the same. This means that larger doses are required to achieve the same effect...

 species, as it can exploit the thin film of oxygen-rich surface water with its upturned mouth, so is able to survive oxygen-depleted habitats. A euryhaline
Euryhaline
Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the molly which can live in fresh, brackish, or salt water. The European shore crab is an example of a euryhaline invertebrate that can live in salt and brackish water...

 species, the sailfin molly may be found in a variety of saline
Salinity
Salinity is the saltiness or dissolved salt content of a body of water. It is a general term used to describe the levels of different salts such as sodium chloride, magnesium and calcium sulfates, and bicarbonates...

 environments, tolerating salinities as high as 87 ppt
PPT
- Organizations :* Parti progressiste tchadien, a political party active in Chad between 1947 and 1973* Patria Para Todos, a left-wing political party in Venezuela* Permanent Peoples' Tribunal, an international opinion tribunal founded in Bologna, 1979...

 and breeding in brackish water
Brackish water
Brackish water is water that has more salinity than fresh water, but not as much as seawater. It may result from mixing of seawater with fresh water, as in estuaries, or it may occur in brackish fossil aquifers. The word comes from the Middle Dutch root "brak," meaning "salty"...

s.

Adults thrive best in isolated pools or organically enriched waterways where few other fish occur (Minckley 1973).

Distinctive features

The body of the sailfin molly is essentially oblong
Rectangle
In Euclidean plane geometry, a rectangle is any quadrilateral with four right angles. The term "oblong" is occasionally used to refer to a non-square rectangle...

. The head is small and dorsally flattened, with a small, upturned mouth. The caudal peduncle is broad and the caudal fin is large, rounded, and sometimes tipped with black. The pelvic fins originate at a point anterior to the dorsal fin. In mature males, the dorsal fin is greatly enlarged and colourful (it is this feature that gives the species its common name) and the caudal fin is similarly colourful; these conspicuous secondary sexual features play a role in female mate choices. Females tend to be larger and more plainly coloured, a difference characteristic to the Poeciliidae. (See: Sexual selection
Sexual selection
Sexual selection, a concept introduced by Charles Darwin in his 1859 book On the Origin of Species, is a significant element of his theory of natural selection...

)

It is a smaller fish than the Yucatan molly (P. velifera
Poecilia velifera
Poecilia velifera, the Yucatan molly, lives in coastal waters of the Yucatan peninsula. These live-bearer fish are particularly well known for both the extreme size variation among males, and the sexual dimorphism between males and females in both body shape and behavior.It is outwardly similar to...

), though that species often does not grow to full length if bred in an aquarium. The dorsal fins are the most distinctive character: Those of the sailfin molly have less than 15 fin rays, counting where the fin meets the back, whereas the Yucatan molly has 18-19 (intermediate numbers may indicate hybrids which should be avoided). If a male spreads his dorsal fins in display, in this species it forms a trapezoid
Trapezoid
In Euclidean geometry, a convex quadrilateral with one pair of parallel sides is referred to as a trapezoid in American English and as a trapezium in English outside North America. A trapezoid with vertices ABCD is denoted...

, with the posterior edge being shortest. The height of the dorsal fin, measured at the posterior edge, is a bit less than the height of the tail.

Colouration

The body is generally light grey
Grey
Grey or gray is an achromatic or neutral color.Complementary colors are defined to mix to grey, either additively or subtractively, and many color models place complements opposite each other in a color wheel. To produce grey in RGB displays, the R, G, and B primary light sources are combined in...

, although breeding males may be greenish-blue. Several rows of spots occur along the sides, back, and dorsal fin. Often, these spots blend together, forming stripes. Aquarists have developed many colour variations in this species (variation occurs naturally in the wild), with melanistic
Melanism
Melanism is an undue development of dark-colored pigment in the skin or its appendages, and the opposite of albinism. It is also the medical term for black jaundice.The word is deduced from the , meaning black pigment....

, leucistic, albino
Albinism
Albinism is a congenital disorder characterized by the complete or partial absence of pigment in the skin, hair and eyes due to absence or defect of an enzyme involved in the production of melanin...

, and speckled forms known.

Sailfin and black mollies

There exists an entirely melanistic
Melanism
Melanism is an undue development of dark-colored pigment in the skin or its appendages, and the opposite of albinism. It is also the medical term for black jaundice.The word is deduced from the , meaning black pigment....

 form called the midnight molly, or nondescriptly, "black molly". The latter originally refers to melanistic breeds of Poecilia sphenops
Poecilia sphenops
Poecilia sphenops is a species of fish, of the genus Poecilia, known under the common name molly; to distinguish it from its congeners, it is sometimes called short-finned molly or common molly. They inhabit fresh water streams and coastal brackish and marine waters of Mexico...

. Midnight mollies actually originated from hybrids between that and the present species.

As hybridisation, like in most Poecilia
Poecilia
Poecilia is a genus of euryhaline brackish water fish in family Poeciliidae of order Cyprinodontiformes. The type species is P. vivipara. Live-bearers, the Poecilia species are collectively known as mollies, with the exception of Endler's livebearer and the famous guppy...

, is easy between these two species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 and due to the more spectacular appearance of P. latipinna, such sailfin-black molly hybrids, with males' conspicuous, large, yellow-rimmed dorsal fins, are often seen.
Due to genomic recombination
Genetic recombination
Genetic recombination is a process by which a molecule of nucleic acid is broken and then joined to a different one. Recombination can occur between similar molecules of DNA, as in homologous recombination, or dissimilar molecules, as in non-homologous end joining. Recombination is a common method...

, F1 hybrid
F1 hybrid
F1 hybrid is a term used in genetics and selective breeding. F1 stands for Filial 1, the first filial generation seeds/plants or animal offspring resulting from a cross mating of distinctly different parental types....

s often display novel and bizarre fin shapes. This can include a grotesquely elongated gonopodium in males; such hybrids are usually unable to breed. Otherwise, hybrids can be bred among themselves, or with higher rate of success with their parent species. They often have a somewhat decreased lifespan, but not as much (in healthy fish) as the deformed "balloon" molly breeds of P. sphenops. Like the sailfin molly, they require a bit more attention of the fishkeeper than the extremely hardy "true" black mollies.
The female has a fan-shaped anal fin and male has a pointed anal fin.

Size, age, and growth

The natural lifespan of sailfin mollies is short, particularly in the case of the males, which may live less than a year after achieving sexual maturity. Depending upon environmental conditions, sailfin mollies may become reproductive in less than a year. Sailfin mollies are small fish. At one year of age, males typically range in size from 15–68 mm SL
Fish measurement
Fish measurement refers to the measuring of the length of individual fish and of various parts of their anatomy. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.-Overall length:...

, while mature females are likely to be 19–53 mm SL. The size of adult males is directly correlated with population density. The greater the population, the smaller the average size of males. The maximum recorded size for this species is 150 mm TL
Fish measurement
Fish measurement refers to the measuring of the length of individual fish and of various parts of their anatomy. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.-Overall length:...

.

Diet

Sailfin mollies feed primarily upon algae
Algae
Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

 and other plant materials, although they will consume a number of aquatic invertebrate
Invertebrate
An invertebrate is an animal without a backbone. The group includes 97% of all animal species – all animals except those in the chordate subphylum Vertebrata .Invertebrates form a paraphyletic group...

s, including the larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e of mosquito
Mosquito
Mosquitoes are members of a family of nematocerid flies: the Culicidae . The word Mosquito is from the Spanish and Portuguese for little fly...

es.

Reproduction

Fertilisation is internal
Internal fertilization
In mammals, internal fertilization is done through copulation, which involves the insertion of the penis into the vagina. Some other higher vertebrate animals reproduce internally, but their fertilization is cloacal.The union of spermatozoa of the parent organism. At some point, the growing egg or...

, and is accomplished by means of highly modified fin elements within the anal fin of males that form a structure known as the gonopodium. Sailfin mollies produce broods of 10-140 live young, depending upon maturity and size, and females may store sperm long after the demise of their relatively short-lived mates. The gestation
Gestation
Gestation is the carrying of an embryo or fetus inside a female viviparous animal. Mammals during pregnancy can have one or more gestations at the same time ....

 period for this species is about three to four weeks, depending upon temperature, and a single female may give birth on multiple occasions throughout the year. Although sex ratios of the broods are balanced, adult populations tend to be largely female, as males appear to suffer higher rates of mortality due to a greater susceptibility to predators and disease as a consequence of their brighter colours and a life devoted to frenzied breeding. There is no parental care exhibited by this species. A ratio of three females to one male is preferred, as with all live bearers, because the females are harassed by males to the point of exhaustion, and having more females gives the others a rest.

Predators

Sailfin mollies are members of the lower end of the food chain
Food chain
A food web depicts feeding connections in an ecological community. Ecologists can broadly lump all life forms into one of two categories called trophic levels: 1) the autotrophs, and 2) the heterotrophs...

. As such, they are prey for various animals, including aquatic insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s, other fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

es, reptile
Reptile
Reptiles are members of a class of air-breathing, ectothermic vertebrates which are characterized by laying shelled eggs , and having skin covered in scales and/or scutes. They are tetrapods, either having four limbs or being descended from four-limbed ancestors...

s, amphibian
Amphibian
Amphibians , are a class of vertebrate animals including animals such as toads, frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic tetrapods...

s, bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s and mammal
Mammal
Mammals are members of a class of air-breathing vertebrate animals characterised by the possession of endothermy, hair, three middle ear bones, and mammary glands functional in mothers with young...

s. Specific examples include: giant water bugs (Belostomatidae), largemouth bass
Largemouth bass
The largemouth bass is a species of black bass in the sunfish family native to North America . It is also known as widemouth bass, bigmouth, black bass, bucketmouth, Potter's fish, Florida bass, Florida largemouth, green bass, green trout, linesides, Oswego bass, southern largemouth...

 (Micropterus salmoides), American alligator
American Alligator
The American alligator , sometimes referred to colloquially as a gator, is a reptile endemic only to the Southeastern United States. It is one of the two living species of alligator, in the genus Alligator, within the family Alligatoridae...

 (Alligator mississippiensis), bullfrog
Bullfrog
The American bullfrog , often simply known as the bullfrog in Canada and the United States, is an aquatic frog, a member of the family Ranidae, or “true frogs”, native to much of North America. This is a frog of larger, permanent water bodies, swamps, ponds, and lakes, where it is usually found...

 (Rana catesbeiana), snowy egret
Snowy Egret
The Snowy Egret is a small white heron. It is the American counterpart to the very similar Old World Little Egret, which has established a foothold in the Bahamas....

 (Egretta thula), and raccoon
Raccoon
Procyon is a genus of nocturnal mammals, comprising three species commonly known as raccoons, in the family Procyonidae. The most familiar species, the common raccoon , is often known simply as "the" raccoon, as the two other raccoon species in the genus are native only to the tropics and are...

 (Procyon lotor).

Parasites

The trematode Saccocoelioides sogandaresi is a known parasite of the sailfin molly.

The ciliated parasite Ichthyophthirius
Ichthyophthirius
Ichthyophthirius multifiliis is a common disease of freshwater fish. It is caused by the protozoa Ichtyopthirius. Ich is one of the most common and persistent diseases.The protozoan is an ectoparasite...

, or more commonly known as white spot or Ich

Importance to humans

The sailfin molly, in its many colour varieties, is of considerable interest and value to aquarists
Aquarium
An aquarium is a vivarium consisting of at least one transparent side in which water-dwelling plants or animals are kept. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, marine mammals, turtles, and aquatic plants...

, and many artificially selected varieties are produced and sold in pet shops. Wild-type sailfin mollies are also bred as feeder fish
Feeder fish
Feeder fish is the generic name for certain types of inexpensive fish commonly fed as live prey to captive animals such as sharks, turtles and some other fish such as oscars.-Advantages of using feeder fish:...

 for larger carnivorous fish. Naturally occurring populations of sailfin mollies help to control mosquito
Mosquito
Mosquitoes are members of a family of nematocerid flies: the Culicidae . The word Mosquito is from the Spanish and Portuguese for little fly...

 populations by feeding on the larva
Larva
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle...

e and pupa
Pupa
A pupa is the life stage of some insects undergoing transformation. The pupal stage is found only in holometabolous insects, those that undergo a complete metamorphosis, going through four life stages; embryo, larva, pupa and imago...

e of these pests
Pest (animal)
A pest is an animal which is detrimental to humans or human concerns. It is a loosely defined term, often overlapping with the related terms vermin, weeds, parasites and pathogens...

. As noted above, they may not be suitable as a biological control agent
Biological pest control
Biological control of pests in agriculture is a method of controlling pests that relies on predation, parasitism, herbivory, or other natural mechanisms...

 in many cases, as they can easily become an invasive species
Invasive species
"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

, or even a pest themselves.

Conservation

This species is not listed as threatened or vulnerable by the World Conservation Union (IUCN)
World Conservation Union
The International Union for Conservation of Nature and Natural Resources is an international organization dedicated to finding "pragmatic solutions to our most pressing environment and development challenges." The organization publishes the IUCN Red List, compiling information from a network of...

.
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