Protobird
Encyclopedia
"Protobird" is an informal term that has been used by some paleontologists when discussing animals that, while technically classified as non-avian dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

s, possess many features normally associated with bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s. All protobirds are extinct.

Zhou and Farlow (2001), for example, used the term "protobird" for primitive members of the clade Avialae
Avialae
Avialae is a clade of dinosaurs containing their only living representatives, birds , and the most immediate extinct relatives of birds.-Competing definitions:...

. In this sense, protobirds would include animals like Confuciusornis
Confuciusornis
Confuciusornis is a genus of primitive crow-sized birds from the Early Cretaceous Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago...

, Sapeornis
Sapeornis
Sapeornis is a genus of primitive bird which lived during the Early Cretaceous . The genus contains only the species Sapeornis chaoyangensis which is known from fossils found in Jiufotang Formation rocks near Chaoyang, PRC...

, and the Enantiornithes
Enantiornithes
Enantiornithes is an extinct group of primitive birds. They were the most abundant and diverse avialans of the Mesozoic. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over 50 species of Enantiornithines have been named, but some...

. These animals were small, flying, feathered, and closely related to birds. The authors restricted the term "bird" to refer only to Aves, which they used to mean only modern ("crown group") birds.

Gregory S. Paul
Gregory S. Paul
Gregory Scott Paul is a freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dinosaurs and his detailed illustrations, both live and skeletal...

 used the term "protobird" in a wider sense in 1988, to refer to the extremely bird-like non-avian dinosaurs (Maniraptora
Maniraptora
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. Ornitholestes and the Alvarezsauroidea...

), including oviraptorosaurs
Oviraptorosauria
Oviraptorosaurs are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot - like skulls, with or without bony crests atop the head...

, troodontids
Troodontidae
Troodontidae is a family of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and scrappy and they have therefore been allied, at various times, with many dinosaurian lineages...

, and dromaeosaurids
Dromaeosauridae
Dromaeosauridae is a family of bird-like theropod dinosaurs. They were small- to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek dromeus meaning 'runner' and sauros meaning 'lizard'...

. Paul speculated that these forms were so bird-like they probably had feather
Feather
Feathers are one of the epidermal growths that form the distinctive outer covering, or plumage, on birds and some non-avian theropod dinosaurs. They are considered the most complex integumentary structures found in vertebrates, and indeed a premier example of a complex evolutionary novelty. They...

s, an idea later proven by fossil evidence.
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