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Population dynamics



 
 
Population dynamics is the branch of life sciences that studies short- and long-term changes in the size and age composition of population
Population

File:Population density.pngIn biology, a population is the collection of inter-breeding organisms of a particular species; in sociology, a collection of human beings....
s, and the biological
Biology

Biology is a branch of the natural sciences concerned with the study of living organisms and their interaction with each other and their environment ....
 and environmental
Environment (biophysical)

The biophysical environment is the symbiosis between the physics environment and the biological life forms within the environment, and include all variables that comprise the Earth's biosphere....
 processes influencing those changes. Population dynamics deals with the way populations are affected by birth
Birth rate

Crude birth rate is the natality or childbirths per 1,000 people per year.It can be represented by number of childbirths in that year, and p is the current population....
 and death rates, and by immigration
Immigration

While the movement of people has thought throughout history at various levels, modern immigration tourism are considered non-immigrants . Immigration that violates the immigration laws of the destination country is termed illegal immigration or undocumented immigration....
 and emigration
Emigration

Emigration is the act of leaving one's native country or region to Settler in another. It is the same as immigration but from the perspective of the country of origin....
, and studies topics such as aging populations or population decline
Population decline

Population decline is the reduction over time in a region's census. It can be caused for several reasons; notable ones include sub-replacement fertility , heavy emigration, disease, famine, and war....
.

lation dynamics is traditionally been the dominant branch of mathematical biology
Mathematical biology

Mathematical biology/Theoretical biology includes at least four major subfields: Biological mathematical modeling, Relational biology/Complex systems biology , Bioinformatics and Computational biomodeling/biocomputing, and is an interdisciplinary research field of academic study with a wide range of applications in Bio...
, which has a history of more than 210 years, although more recently the scope of mathematical biology
Mathematical biology

Mathematical biology/Theoretical biology includes at least four major subfields: Biological mathematical modeling, Relational biology/Complex systems biology , Bioinformatics and Computational biomodeling/biocomputing, and is an interdisciplinary research field of academic study with a wide range of applications in Bio...
 has greatly expanded.






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Population dynamics is the branch of life sciences that studies short- and long-term changes in the size and age composition of population
Population

File:Population density.pngIn biology, a population is the collection of inter-breeding organisms of a particular species; in sociology, a collection of human beings....
s, and the biological
Biology

Biology is a branch of the natural sciences concerned with the study of living organisms and their interaction with each other and their environment ....
 and environmental
Environment (biophysical)

The biophysical environment is the symbiosis between the physics environment and the biological life forms within the environment, and include all variables that comprise the Earth's biosphere....
 processes influencing those changes. Population dynamics deals with the way populations are affected by birth
Birth rate

Crude birth rate is the natality or childbirths per 1,000 people per year.It can be represented by number of childbirths in that year, and p is the current population....
 and death rates, and by immigration
Immigration

While the movement of people has thought throughout history at various levels, modern immigration tourism are considered non-immigrants . Immigration that violates the immigration laws of the destination country is termed illegal immigration or undocumented immigration....
 and emigration
Emigration

Emigration is the act of leaving one's native country or region to Settler in another. It is the same as immigration but from the perspective of the country of origin....
, and studies topics such as aging populations or population decline
Population decline

Population decline is the reduction over time in a region's census. It can be caused for several reasons; notable ones include sub-replacement fertility , heavy emigration, disease, famine, and war....
.

History

Population dynamics is traditionally been the dominant branch of mathematical biology
Mathematical biology

Mathematical biology/Theoretical biology includes at least four major subfields: Biological mathematical modeling, Relational biology/Complex systems biology , Bioinformatics and Computational biomodeling/biocomputing, and is an interdisciplinary research field of academic study with a wide range of applications in Bio...
, which has a history of more than 210 years, although more recently the scope of mathematical biology
Mathematical biology

Mathematical biology/Theoretical biology includes at least four major subfields: Biological mathematical modeling, Relational biology/Complex systems biology , Bioinformatics and Computational biomodeling/biocomputing, and is an interdisciplinary research field of academic study with a wide range of applications in Bio...
 has greatly expanded. The first principle of population dynamics is widely regarded as the exponential law of Malthus, as modelled by the Malthusian growth model
Malthusian growth model

The Malthusian growth model, sometimes called the simple exponential growth model, is essentially exponential growth based on a constant rate of compound interest....
. The early period was dominated by demographic
Demography

Demography is the statistical study of all populations. It can be a very general science that can be applied to any kind of dynamic population, that is, one that changes over time or space ....
 studies such as the work of Benjamin Gompertz
Benjamin Gompertz

Benjamin Gompertz , was a autodidact mathematician, denied admission to university because he was Jewish. Nevertheless he was made Fellow of the Royal Society in 1819....
 and Pierre François Verhulst
Pierre François Verhulst

Pierre Fran?ois Verhulst was a mathematician and a doctor in number theory from the University of Ghent in 1825. Verhulst published in 1838 the logistic equation:...
 in the early 19th century, who refined and adjusted the Malthusian demographic model.

A more general model formulation was proposed by F.J. Richards in 1959, by which the models of Gompertz, Verhulst and also Ludwig von Bertalanffy
Ludwig von Bertalanffy

Karl Ludwig von Bertalanffy was an Austrian-born biology known as one of the founders of systems theory. Von Bertalanffy grew up in Austria and subsequently worked in Vienna, London, Canada and the USA....
 are covered as special cases of the general formulation. The computer game SimCity
SimCity

SimCity is a city-building game Construction and management simulation games personal computer game, first released in 1989 and designed by Will Wright ....
 and the MMORPG
MMORPG

A massively multiplayer online role-playing game is a genre of computer role-playing games in which a large number of player interact with one another in a virtual world....
 Ultima Online
Ultima Online

Ultima Online is a graphical massively multiplayer online role-playing game , released on September 25, 1997, by Origin Systems. It was instrumental to the development of the genre, and is still running today....
, among others, tried to simulate
Computer simulation

A computer simulation, a computer model or a computational model is a computer program, or network of computers, that attempts to simulation an abstract model of a particular system....
 some of these population dynamics.

Fisheries and wildlife management

In fisheries and wildlife management
Wildlife management

Wildlife management is the process of keeping certain wildlife populations, including endangered animals, at desirable levels determined by wildlife managers....
, population is affected by three dynamic rate functions.

  • Natality or birth rate
    Birth rate

    Crude birth rate is the natality or childbirths per 1,000 people per year.It can be represented by number of childbirths in that year, and p is the current population....
    , often recruitment, which means reaching a certain size or reproductive stage. Usually refers to the age a fish can be caught and counted in nets
  • Growth rate
    Growth rate

    Growth rate may refer to:*Exponential growth, a growth rate classification*Compound annual growth rate or CAGR, a measure of financial growth...
    , which measures the growth of individuals in size and length. More important in fisheries, where population is often measured in biomass.
  • Mortality
    Mortality rate

    Mortality rate is a measure of the number of deaths in some population, scaled to the size of that population, per unit time. Mortality rate is typically expressed in units of deaths per 1000 individuals per year; thus, a mortality rate of 9.5 in a population of 100,000 would mean 950 deaths per year in that entire population....
    , which includes harvest mortality and natural mortality. Natural mortality includes non-human predation, disease and old age.


If N1 is the number of individuals at time 1 then N1 = N0 + B - D + I - E where N0 is the number of individuals at time 0, B is the number of individuals born, D the number that died, I the number that immigrated, and E the number that emigrated between time 0 and time 1.

If we measures these rates over many time intervals, we can determine how a population's density changes over time. Immigration and emigration are present, but are usually not measured.

All of these are measured to determine the harvestable surplus, which is the number of individuals that can be harvested from a population without affecting long term stability, or average population size. The harvest within the harvestable surplus is considered compensatory mortality, where the harvest deaths are substituting for the deaths that would occur naturally. Harvest beyond that is additive mortality, harvest in addition to all the animals that would have died naturally. These terms are not the universal good and evil of population management, for example, in deer, the DNR are trying to reduce deer population size overall to an extent, since hunters have reduced buck competition and increased deer population unnaturally.

Intrinsic rate of increase

The rate at which a population increases in size, i.e. the change in population size over a particular period of time is known as the intrinsic rate of increase. The concept is commonly used in insect population biology to determine how environmental factors affect the rate at which pest populations increase.

See also


External links

  • . Greenboxes (Beta) is a repository for open-source population modelling and PVA code. Greenboxes allows users an easy way to share their code and to search for others shared code.
  • . An online compilation of state-ot-the-art basic tools for the analysis of population dynamics with emphasis on benthic invertebrates.