Piebald
Encyclopedia
A piebald or pied animal is one that has a spotting pattern of large unpigmented, usually white, areas of hair, feathers, or scales and normally pigmented patches, generally black. The colour of the animal's skin underneath its coat is also pigmented under the dark patches and unpigmented under the white patches. This alternating colour pattern is irregular and asymmetrical. Animals with this pattern may include horse
Horse
The horse is one of two extant subspecies of Equus ferus, or the wild horse. It is a single-hooved mammal belonging to the taxonomic family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature into the large, single-toed animal of today...

s dog
Dog
The domestic dog is a domesticated form of the gray wolf, a member of the Canidae family of the order Carnivora. The term is used for both feral and pet varieties. The dog may have been the first animal to be domesticated, and has been the most widely kept working, hunting, and companion animal in...

s, bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s, cat
Cat
The cat , also known as the domestic cat or housecat to distinguish it from other felids and felines, is a small, usually furry, domesticated, carnivorous mammal that is valued by humans for its companionship and for its ability to hunt vermin and household pests...

s, pig
Pig
A pig is any of the animals in the genus Sus, within the Suidae family of even-toed ungulates. Pigs include the domestic pig, its ancestor the wild boar, and several other wild relatives...

s, and cattle
Cattle
Cattle are the most common type of large domesticated ungulates. They are a prominent modern member of the subfamily Bovinae, are the most widespread species of the genus Bos, and are most commonly classified collectively as Bos primigenius...

, as well as snakes such as the ball python
Ball python
Python regius is a nonvenomous python species found in Africa. This is the smallest of the African pythons and is popular in the pet trade. No subspecies are currently recognized. They are also known as royal pythons or ball pythons. The name ball python refers to the animal's tendency to curl into...

. Some animals also exhibit colouration of the irises of the eye that match the surrounding skin (blue eyes for pink skin, brown for dark). The underlying genetic cause is related to a condition known as leucism.

Etymology

The word "piebald" originates from a combination of "pie," from "magpie," and "bald", meaning "white patch" or spot. The reference is to the distinctive black-and white plumage of the magpie
Magpie
Magpies are passerine birds of the crow family, Corvidae.In Europe, "magpie" is often used by English speakers as a synonym for the European Magpie, as there are no other magpies in Europe outside Iberia...


In horses

In British English
British English
British English, or English , is the broad term used to distinguish the forms of the English language used in the United Kingdom from forms used elsewhere...

 piebald (black and white) and skewbald
Skewbald
Skewbald is a color pattern of horses. A skewbald horse has a coat made up of white patches on a non-black base coat, such as chestnut, bay, or any color besides black coat. Skewbald horses which are bay and white are sometimes called tricoloured...

(white and any colour other than black) are together known as coloured. In North American English
North American English
North American English is the variety of the English language of North America, including that of the United States and Canada. Because of their shared histories and the similarities between the pronunciation, vocabulary and accent of American English and Canadian English, the two spoken languages...

, the term for this colouring pattern is pinto
Pinto horse
A pinto horse has a coat color that consists of large patches of white and any other color. The distinction between "pinto" and "solid" can be tenuous, as so-called "solid" horses frequently have areas of white hair. Various cultures throughout history appear to have selectively bred for pinto...

, with the specialized term "paint"
American Paint Horse
The American Paint Horse is a breed of horse that combines both the conformational characteristics of a western stock horse with a pinto spotting pattern of white and dark coat colors. Developed from a base of spotted horses with Quarter Horse and Thoroughbred bloodlines, the American Paint Horse...

 referring specifically to a breed of horse with American Quarter Horse
American Quarter Horse
The American Quarter Horse is an American breed of horse that excels at sprinting short distances. Its name came from its ability to outdistance other breeds of horses in races of a quarter mile or less; some individuals have been clocked at speeds up to 55 mph...

 or Thoroughbred
Thoroughbred
The Thoroughbred is a horse breed best known for its use in horse racing. Although the word thoroughbred is sometimes used to refer to any breed of purebred horse, it technically refers only to the Thoroughbred breed...

 bloodlines in addition to being spotted, whereas pinto refers to a spotted horse of any breed. In American usage, horse enthusiasts usually do not use the term "piebald," but rather describe the colour shade of a pinto literally with terms such as "black and white" for a piebald, "brown and white," or "bay and white," for skewbalds, or color-specific modifers such as "bay pinto", "sorrel pinto," "buckskin pinto," and such.

Genetically
Equine coat color genetics
Equine coat color genetics determine a horse's coat color. There are many different coat colors possible, but all colors are produced by the action of only a few different genes. The simplest genetic default color of all domesticated horses can be described as either "red" or "non-red", depending...

, a piebald horse begins with a black
Black (horse)
Black is a hair coat color of horses in which the entire hair coat is black. Black is a relatively uncommon coat color, and novices frequently mistake dark chestnuts or bays for black. However, some breeds of horses, such as the Friesian horse, Murgese and Ariegeois are almost exclusively black...

 base coat colour, and then the horse also has an allele
Allele
An allele is one of two or more forms of a gene or a genetic locus . "Allel" is an abbreviation of allelomorph. Sometimes, different alleles can result in different observable phenotypic traits, such as different pigmentation...

 for one of three basic spotting patterns overlaying the base colour. The most common coloured spotting pattern is called tobiano
Tobiano
Tobiano is a spotted color pattern commonly seen in Pinto horses, produced by a dominant gene. The tobiano gene produces white-haired, pink-skinned patches on a base coat color. The coloration is present from birth and does not change throughout the horse's lifetime, unless the horse also carries...

, and is a dominant gene. Tobiano creates spots that are large and rounded, usually with a somewhat vertical orientation, with white that usually crosses the back of the horse, white on the legs, with the head mostly dark. Three less common spotting gene
Gene
A gene is a molecular unit of heredity of a living organism. It is a name given to some stretches of DNA and RNA that code for a type of protein or for an RNA chain that has a function in the organism. Living beings depend on genes, as they specify all proteins and functional RNA chains...

s are the sabino, frame and splash overo
Overo
Overo refers to several genetically unrelated pinto coloration patterns of white-over-dark body markings in horses, and is a term used by the American Paint Horse Association to classify a set of pinto patterns that are not Tobiano...

 genes, which create various patterns that are mostly dark, with jagged spotting, often with a horizontal orientation, white on the head. The frame variant has dark or minimally marked legs. The sabino
Sabino horse
Sabino is a group of white spotting patterns in horses that affect the skin and hair. A wide variety of irregular color patterns are accepted as sabino. In the strictest sense, "sabino" refers to the white patterns produced by the Sabino 1 gene, for which there is a DNA test...

 pattern can be very minimal, usually adding white that runs up the legs onto the belly or flanks, with "lacy" or roaning
Roan (horse)
Roan is a horse coat color pattern characterized by an even mixture of colored and white hairs on the body, while the head and "points"—lower legs, mane and tail—are more solid-colored. The roan pattern is dominantly-inherited, and is found in many horse breeds...

 at the edge of the white, plus white on the head that either extends past the eye, over the chin, or both. The genetics of overo and sabino are not yet fully understood, but they can appear in the offspring of two solid-coloured parents, whereas a tobiano must always have at least one tobiano parent.

In other animals

Many other animal species may also be "pied" or piebald. Snake
Snake
Snakes are elongate, legless, carnivorous reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears. Like all squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales...

s, especially ball python
Ball python
Python regius is a nonvenomous python species found in Africa. This is the smallest of the African pythons and is popular in the pet trade. No subspecies are currently recognized. They are also known as royal pythons or ball pythons. The name ball python refers to the animal's tendency to curl into...

s, may also exhibit seemingly varying patches of completely pigmentless scales along with patches of pigmented scales. The term was used in The Conservationist
The Conservationist
The Conservationist is a 1974 novel by 1991 Nobel laureate Nadine Gordimer. The book was a joint winner of the Man Booker Prize for fiction.-Plot summary:...

by Nadine Gordimer
Nadine Gordimer
Nadine Gordimer is a South African writer and political activist. She was awarded the 1991 Nobel Prize in Literature when she was recognised as a woman "who through her magnificent epic writing has – in the words of Alfred Nobel – been of very great benefit to humanity".Her writing has long dealt...

 when referring to kingfisher
Kingfisher
Kingfishers are a group of small to medium sized brightly coloured birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species being found in the Old World and Australia...

s. The various types of magpie
Magpie (disambiguation)
-Birds:* Magpie, members of four genera of the bird family Corvidae** Australian Magpie, Gymnorhina tibicen, crow-sized black and white bird in the family Artamidae** Azure-winged Magpie, Cyanopica cyana...

 are so called because of their pied plumage. The bald eagle
Bald Eagle
The Bald Eagle is a bird of prey found in North America. It is the national bird and symbol of the United States of America. This sea eagle has two known sub-species and forms a species pair with the White-tailed Eagle...

 derives its name from the word "piebald" in reference to the contrast of its white head and tail with dark body.

See also

  • Equine coat color
    Equine coat color
    Horses exhibit a diverse array of coat colors and distinctive markings. A specialized vocabulary has evolved to describe them.While most horses remain the same color throughout life, a few, over the course of several years, will develop a different coat color from that with which they were born...

  • Equine coat color genetics
    Equine coat color genetics
    Equine coat color genetics determine a horse's coat color. There are many different coat colors possible, but all colors are produced by the action of only a few different genes. The simplest genetic default color of all domesticated horses can be described as either "red" or "non-red", depending...

  • Skewbald
    Skewbald
    Skewbald is a color pattern of horses. A skewbald horse has a coat made up of white patches on a non-black base coat, such as chestnut, bay, or any color besides black coat. Skewbald horses which are bay and white are sometimes called tricoloured...

  • Tricoloured (horse)
  • Leucistic
    Leucistic
    Leucism is a condition characterized by reduced pigmentation in animals and humans. Unlike albinism, it is caused by a reduction in all types of skin pigment, not just melanin.- Details :...

  • Pinto horse
    Pinto horse
    A pinto horse has a coat color that consists of large patches of white and any other color. The distinction between "pinto" and "solid" can be tenuous, as so-called "solid" horses frequently have areas of white hair. Various cultures throughout history appear to have selectively bred for pinto...

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