Persian Leopard
Encyclopedia
The Persian leopard also called Caucasian leopard, is the largest leopard
Leopard
The leopard , Panthera pardus, is a member of the Felidae family and the smallest of the four "big cats" in the genus Panthera, the other three being the tiger, lion, and jaguar. The leopard was once distributed across eastern and southern Asia and Africa, from Siberia to South Africa, but its...

 subspecies, and is native to eastern Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

, the Caucasus
Caucasus
The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

 mountains, northern Iran
Iran
Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...

, southern Turkmenistan
Turkmenistan
Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states...

, and parts of western Afghanistan
Afghanistan
Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...

. It is endangered
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

 throughout its range with fewer than 871–1,290 mature individuals and a declining population trend.

Characteristics

The Persian leopard is large, weighing up to 60 kg (132.3 lb), and light in color.

Biometric data collected from 25 female and male individuals in various provinces of Iran indicates average body length of 259 cm (102 in). A young male from northern Iran weighed 64 kg (141.1 lb).

Distribution and habitat

There has been a huge decline in the former range of the leopard in the Caucasus, which is estimated to have once covered the whole region, except for steppe areas. The severe economic crisis following major political and social changes in 1992 in the former Soviet Union
Soviet Union
The Soviet Union , officially the Union of Soviet Socialist Republics , was a constitutionally socialist state that existed in Eurasia between 1922 and 1991....

, together with a weakening of formerly effective protection systems resulted in a sharp rise in hunting of wild ungulates, persecution of leopards and fragmentation of the ranges of all wildlife. As of 2008, of the estimated 871-1,290 mature leopards
  • about 200–300 survive in Afghanistan
    Afghanistan
    Afghanistan , officially the Islamic Republic of Afghanistan, is a landlocked country located in the centre of Asia, forming South Asia, Central Asia and the Middle East. With a population of about 29 million, it has an area of , making it the 42nd most populous and 41st largest nation in the world...

    , where their status is poorly known;
  • fewer than 10–13 survive in Armenia
    Armenia
    Armenia , officially the Republic of Armenia , is a landlocked mountainous country in the Caucasus region of Eurasia...

    ;
  • fewer than 10–13 survive in Azerbaijan
    Azerbaijan
    Azerbaijan , officially the Republic of Azerbaijan is the largest country in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded by the Caspian Sea to the east, Russia to the north, Georgia to the northwest, Armenia to the west, and Iran to...

    ;
  • fewer than 5 survive in Georgia
    Georgia (country)
    Georgia is a sovereign state in the Caucasus region of Eurasia. Located at the crossroads of Western Asia and Eastern Europe, it is bounded to the west by the Black Sea, to the north by Russia, to the southwest by Turkey, to the south by Armenia, and to the southeast by Azerbaijan. The capital of...

    ;
  • about 550–850 live in Iran
    Iran
    Iran , officially the Islamic Republic of Iran , is a country in Southern and Western Asia. The name "Iran" has been in use natively since the Sassanian era and came into use internationally in 1935, before which the country was known to the Western world as Persia...

    , which is the leopard stronghold in Southwest Asia
    Southwest Asia
    Western Asia, West Asia, Southwest Asia or Southwestern Asia are terms that describe the westernmost portion of Asia. The terms are partly coterminous with the Middle East, which describes a geographical position in relation to Western Europe rather than its location within Asia...

    ;
  • about 3–4 survive in Nagorno-Karabakh
    Nagorno-Karabakh
    Nagorno-Karabakh is a landlocked region in the South Caucasus, lying between Lower Karabakh and Zangezur and covering the southeastern range of the Lesser Caucasus mountains...

    ;
  • fewer than 10 survive in the Russian North Caucasus
    North Caucasus economic region
    North Caucasus economic region tr.: Severo-Kavkazsky ekonomichesky rayon) is one of twelve economic regions of Russia.In this area, descending northward from the principal chain of the Caucasus Mountains to a level plain, are found rich deposits of oil, natural gas, and coal. The major cities are...

    ;
  • fewer than 5 survive in Turkey
    Turkey
    Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

    ;
  • about 78–90 live in Turkmenistan
    Turkmenistan
    Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states...

    .


Their habitat consists of subalpine meadows, broadleaved forests and rugged ravines from 600 – in the Greater Caucasus
Greater Caucasus
Greater Caucasus , sometimes translated as "Caucasus Major", "Big Caucasus" or "Large Caucasus") is the major mountain range of the Caucasus Mountains....

, and rocky slopes, mountain steppes, and sparse juniper forests in the Lesser Caucasus
Lesser Caucasus
Lesser Caucasus is one of the two main mountain ranges of Caucasus mountains, of length about 600 km....

 and Iran. Only some small and isolated populations remain in the whole ecoregion
Ecoregion
An ecoregion , sometimes called a bioregion, is an ecologically and geographically defined area that is smaller than an ecozone and larger than an ecosystem. Ecoregions cover relatively large areas of land or water, and contain characteristic, geographically distinct assemblages of natural...

. Suitable habitat in each range country is limited and most often situated in remote border areas. Local populations depend on immigration from source populations in the south, mainly in Iran.

The Persian leopard is a flag-ship species. Its presence is a sign of the health of the entire ecosystem.

In Armenia

In Armenia, people and leopards co-existed since the early Holocene
Holocene
The Holocene is a geological epoch which began at the end of the Pleistocene and continues to the present. The Holocene is part of the Quaternary period. Its name comes from the Greek words and , meaning "entirely recent"...

. By the mid-20th century leopards were relatively common in the country's mountains. Today, the leopard stronghold is the rugged and cliffy terrain of Khosrov Reserve, located south-east of Yerevan
Yerevan
Yerevan is the capital and largest city of Armenia and one of the world's oldest continuously-inhabited cities. Situated along the Hrazdan River, Yerevan is the administrative, cultural, and industrial center of the country...

 on the south-western slopes of the Geghama mountains
Geghama mountains
The Geghama mountains , are mountains in Armenia. is a tableland-type watershed basin of Sevan Lake from east, inflows of rivers Araks and Razdan from north and west, Azat and Vedi rivers from south-west and Arpachai river from south...

, where between October 2000 to July 2002 tracks of no more than 10 individuals were found in an area of 780 km² (301.2 sq mi). Leopards were known to live on the Meghri
Meghri
Meghri is a city in southern Armenia, located in the Syunik province, near the border with Iran. The city's economy is based on the food industry, and contains a bread-baking factory, canneries and a winery. Meghri has a significantly milder climate than the rest of the cities in Armenia, and...

 Ridge in the extreme south of Armenia, where only one individual was camera-trapped
Camera trap
A camera trap is a remotely activated camera that is equipped with a motion sensor or an infrared sensor, or uses a light beam as a trigger. Camera trapping is a method for capturing wild animals on film when researchers are not present, and has been used in ecological research for decades...

 between August 2006 to April 2007, and no signs of other leopards were found during track surveys conducted over an area of 296.9 km² (114.6 sq mi). The local prey base could support 4–10 individuals, but poaching
Poaching
Poaching is the illegal taking of wild plants or animals contrary to local and international conservation and wildlife management laws. Violations of hunting laws and regulations are normally punishable by law and, collectively, such violations are known as poaching.It may be illegal and in...

 and disturbance caused by livestock breeding, gathering of edible plants and mushrooms, deforestation and human-induced wild fires are so high that they exceed the tolerance limits of leopards.

In Azerbaijan

Leopards are present in the Talysh Mountains
Talysh Mountains
Talysh Mountains is a mountain chain in northwestern Iran, and southeastern Republic of Azerbaijan in the northwest section of the Elburz Mountains, extending southeastward from the Lankaran Lowland in southeast Azerbaijan to the lower part of the Sefid Rud in northwest Iran...

 in the farsoutheast, where their habitat is continuous with that on the Iranian side of Talysh Mountains. They also survived in northwest Azerbaijan in the Akhar-Bakhar section of Ilisu State Reserve
Ilisu State Reserve
Ilisu State Reserve was established on the area of 93 km² in 1987. It aims to protect natural complexes of southern slopes of Major Caucasus, to preserve rare and endangered flora and fauna, to restore forests and prevent erosion of soil and flood. The reserve accounts for 500 plant species with...

 in the foothills of the Greater Caucasus until recently, but current numbers are extremely low.

Despite occasional sightings, it was not clear whether leopards had been extinct in Azerbaijan by the late 1990s, until a specimen was camera-trapped
Camera trap
A camera trap is a remotely activated camera that is equipped with a motion sensor or an infrared sensor, or uses a light beam as a trigger. Camera trapping is a method for capturing wild animals on film when researchers are not present, and has been used in ecological research for decades...

 in March 2007 in the Hirkan State Reserve/Hirkan National Park
Hirkan National Park
Hirkan National Park — is a national park of Azerbaijan. It was established by the decree of the President of Azerbaijan Ilham Aliyev, in an area in Lankaran Rayon and Astara Rayon administrative districts on February 9, 2004 on the basis of the former "Hirkan State Reserve" which it superseded,...

.

In Georgia

Since 1954, leopards were thought to be extinct in Georgia — killed by hunters. In April 2001, an adult female was shot on the border to Kabardino-Balkaria
Kabardino-Balkaria
The Kabardino-Balkar Republic , or Kabardino-Balkaria , is a federal subject of Russia located in the North Caucasus. Population: -Geography:The republic is situated in the North Caucasus mountains, with plains in the northern part....

, her two cubs captured and taken to the Novosibirsk
Novosibirsk
Novosibirsk is the third-largest city in Russia, after Moscow and Saint Petersburg, and the largest city of Siberia, with a population of 1,473,737 . It is the administrative center of Novosibirsk Oblast as well as of the Siberian Federal District...

 Zoo in Russia. In the winter of 2003, zoologists found footprints of a leopard in Vashlovani Reserve
Vashlovani National Park
Vashlovani National Park is a national park located in the eastern part of Georgia, and was established in 1935 to preserve its unique shallow forests. In April 2003 the Reserve's area was expanded to 84.80 km² and Vashlovani National Park was made...

 in southeastern Georgia and later camera-trapped
Camera trap
A camera trap is a remotely activated camera that is equipped with a motion sensor or an infrared sensor, or uses a light beam as a trigger. Camera trapping is a method for capturing wild animals on film when researchers are not present, and has been used in ecological research for decades...

 one young male individual several times. Leopard signs have also been found at two localities in Tusheti
Tusheti
Tusheti is a historic region in northeast Georgia.-Geography:Located on the northern slopes of the Greater Caucasus Mountains, Tusheti is bordered by the Russian republics of Chechnya and Dagestan to the north and east, respectively; and by the Georgian historic provinces Kakheti and...

, the headwaters of the Andi Koisu and Assa rivers, bordering Dagestan
Dagestan
The Republic of Dagestan is a federal subject of Russia, located in the North Caucasus region. Its capital and the largest city is Makhachkala, located at the center of Dagestan on the Caspian Sea...

.

Over the last 60 years, there have been several sightings of leopards around the Tbilisi
Tbilisi
Tbilisi is the capital and the largest city of Georgia, lying on the banks of the Mt'k'vari River. The name is derived from an early Georgian form T'pilisi and it was officially known as Tiflis until 1936...

 area and in the Shida Kartli
Shida Kartli
Shida Kartli is a region in Georgia. It consists of the following districts: Gori, Kaspi, Kareli, Java, Khashuri.The northern part of the region, namely Java, and northern territories of Kareli and Gori, is controlled by the authorities of the self-proclaimed republic of South Ossetia since...

 province to the northwest of the capital. Leopards live primarily in dense forests, although several have been spotted in the lowland plains in the southeastern region of Kakheti
Kakheti
Kakheti is a historical province in Eastern Georgia inhabited by Kakhetians who speak a local dialect of Georgian. It is bordered by the small mountainous province of Tusheti and the Greater Caucasus mountain range to the north, Russian Federation to the Northeast, Azerbaijan to the Southeast, and...

 in 2004.

In Iran

Leopards are mainly found in the Alborz
Alborz
Alborz , also written as Alburz, Elburz or Elborz, is a mountain range in northern Iran stretching from the borders of Azerbaijan and Armenia in the northwest to the southern end of the Caspian Sea, and ending in the east at the borders of Turkmenistan and Afghanistan...

 and the Zagros mountain ranges. These ranges cover a vast area starting from the borders with Turkey, Azerbaijan and Armenia, extending to the Caspian litoral region and on to Turkmenistan and western parts of Afghanistan in the Alborz
Alborz
Alborz , also written as Alburz, Elburz or Elborz, is a mountain range in northern Iran stretching from the borders of Azerbaijan and Armenia in the northwest to the southern end of the Caspian Sea, and ending in the east at the borders of Turkmenistan and Afghanistan...

 range. Along the Zagros range, leopard habitats extend to the south of Iran, close to the Persian Gulf
Persian Gulf
The Persian Gulf, in Southwest Asia, is an extension of the Indian Ocean located between Iran and the Arabian Peninsula.The Persian Gulf was the focus of the 1980–1988 Iran-Iraq War, in which each side attacked the other's oil tankers...

.

Seventy four protected and non-protected areas were identified across the country in recent years as sites where leopards are present. However, 69% of them are located in the Hircanian forests and Alborz mountain range in the northern part of Iran. This area is regarded as one of the most important habitats for leopards in the country. Even though studies revealed that leopards in Iran inhabit a wide range of temperatures ranging from -23 to +49 ºC, they are most often found in the habitats with temperature of 13 to 18 ºC, 0 to 20 days of ice cover per year and rain fall of more than 200 mm/year.

In Bamu National Park located northeast of Shiraz
Shiraz
Shiraz may refer to:* Shiraz, Iran, a city in Iran* Shiraz County, an administrative subdivision of Iran* Vosketap, Armenia, formerly called ShirazPeople:* Hovhannes Shiraz, Armenian poet* Ara Shiraz, Armenian sculptor...

 in Fars Province, camera trap
Camera trap
A camera trap is a remotely activated camera that is equipped with a motion sensor or an infrared sensor, or uses a light beam as a trigger. Camera trapping is a method for capturing wild animals on film when researchers are not present, and has been used in ecological research for decades...

ping carried out from autumn 2007 to spring 2008 revealed seven individuals in a sampling area of 321.12 km² (124 sq mi).

In the North Caucasus regions of Russia

Signs of leopard presence have been found in the upper Andiyskoe and Avarskoye Koisu rivers in Dagestan
Dagestan
The Republic of Dagestan is a federal subject of Russia, located in the North Caucasus region. Its capital and the largest city is Makhachkala, located at the center of Dagestan on the Caspian Sea...

. According to local reports leopards may also occur in Ingushetia
Ingushetia
The Republic of Ingushetia is a federal subject of Russia , located in the North Caucasus region with its capital at Magas. In terms of area, the republic is the smallest of Russia's federal subjects except for the two federal cities, Moscow and Saint Petersburg...

, Ossetia
Ossetia
Ossetia Ossetic: Ир, Ирыстон Ir, Iryston; Russian: Осетия, Osetiya; Georgian: ოსეთი, Oset'i) is an ethnolinguistic region located on both sides of the Greater Caucasus Mountains, largely inhabited by the Ossetians. The Ossetian language is part of the Eastern Iranian branch of the Indo-European...

 and Chechnya
Chechnya
The Chechen Republic , commonly referred to as Chechnya , also spelled Chechnia or Chechenia, sometimes referred to as Ichkeria , is a federal subject of Russia . It is located in the southeastern part of Europe in the Northern Caucasus mountains. The capital of the republic is the city of Grozny...

. Leopards no longer occur in the Western Caucasus
Western Caucasus
The Western Caucasus is a western region of the Caucasus in Southern Russia, extending from the Black Sea to Mount Elbrus.-World Heritage Site:...

. In 2009, a leopard reintroduction centre was created in Sochi National Park
Sochi National Park
Sochi National Park is a national park in Western Caucasus, near the city of Sochi, in Southern Russia. It is Russia's second oldest national park, established on May 5, 1983.-Geography:...

, where two male leopards from Turkmenistan
Turkmenistan
Turkmenistan , formerly also known as Turkmenia is one of the Turkic states in Central Asia. Until 1991, it was a constituent republic of the Soviet Union, the Turkmen Soviet Socialist Republic . Turkmenistan is one of the six independent Turkic states...

 are being kept since September 2009, and two females from Iran since May 2010. Their descendants will be released into the wild in the Caucasus Biosphere Reserve.

Ecology and behaviour

Leopards' diet varies depending on the habitat of their territory. Their principal prey is the most abundant ungulate
Ungulate
Ungulates are several groups of mammals, most of which use the tips of their toes, usually hoofed, to sustain their whole body weight while moving. They make up several orders of mammals, of which six to eight survive...

 such as Bezoar goat
Bezoar Ibex
The Bezoar Ibex is a vulnerable wild goat, native to Afghanistan, Armenia, Azerbaijan, Georgia, Iran, Russia and Turkey. It has been extirpated from Lebanon. The Bezoar goat is found in the mountains of Asia Minor and across the Middle East...

, roe deer
Roe Deer
The European Roe Deer , also known as the Western Roe Deer, chevreuil or just Roe Deer, is a Eurasian species of deer. It is relatively small, reddish and grey-brown, and well-adapted to cold environments. Roe Deer are widespread in Western Europe, from the Mediterranean to Scandinavia, and from...

, Goitered gazelle
Goitered Gazelle
The Goitered, Black-tailed or Persian gazelle is a gazelle found in the north part of Azerbaijan, in a large area of central Asia, including part of Iran and southern west Pakistan in the western end of the range, as well as the Gobi desert...

, West Caucasian tur
West Caucasian Tur
The West Caucasian tur is a mountain dwelling goat antelope found only in the western half of the Caucasus Mountain range....

, mouflons, urial
Urial
The Urial , also known as the Arkars or Shapo, is a subspecies group of the wild sheep Ovis orientalis. Noticeable features are the reddish-brown long fur that fades during winter; males are characterized by a black ruff stretching from the neck to the chest and large horns. It is found in western...

, and wild boar. They also prey on smaller wildlife such as Crested porcupine
Crested Porcupine
The crested porcupine is a species of rodent in the Hystricidae family.It is extant in mainland Italy, Sicily, North Africa and sub-Saharan Africa.-Physical Attributes:...

 and Cape hare
Cape Hare
The Cape, Common or Brown Hare is a hare natively found throughout Africa, and has spread to many parts of the Europe, Middle East and Asia. The Cape Hare is a nocturnal herbivore.They are fast...

, and occasionally attack livestock and herd dogs.

Studies revealed that presence of leopards in Iran is highly correlated with presence of wild goat
Wild Goat
The wild goat is a widespread species of goat, with a distribution ranging from Europe and Asia Minor to central Asia and the Middle East. It is the ancestor of the domestic goat.-Social structure:...

 and wild sheep. Opportunistic predation on smaller prey species is also probable. Attack of a leopard individual on onager
Onager
The Onager is a large member of the genus Equus of the family Equidae native to the deserts of Syria, Iran, Pakistan, India, Israel and Tibet...

 was also recorded.

Threats

Persian leopards are threatened by poaching
Poaching
Poaching is the illegal taking of wild plants or animals contrary to local and international conservation and wildlife management laws. Violations of hunting laws and regulations are normally punishable by law and, collectively, such violations are known as poaching.It may be illegal and in...

, depletion of their prey base due to poaching, human disturbance such as presence of military and training of troops in border areas, habitat loss due to deforestation, fire, agricultural expansion, overgrazing, and infrastructure development.

In Iran, primary threats are habitat disturbances followed by illegal hunting and excess of livestock in the leopard habitats. The leopards' chances for survival outside protected areas appear very slim. Intensive dry condition in wide areas of leopard habitats in recent years is affecting leopard main prey species such as wild goat and wild sheep.

Taxonomic history

The Russian explorer Satunin first described the Caucasian leopard P.p. ciscaucasica in 1914 on the basis of a specimen from the Kuban
Kuban
Kuban is a geographic region of Southern Russia surrounding the Kuban River, on the Black Sea between the Don Steppe, Volga Delta and the Caucasus...

 region of North Caucasus. The British zoologist Pocock
Reginald Innes Pocock
Reginald Innes Pocock F.R.S. was a British zoologist.Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's School, Oxford. He received tutoring in zoology from Sir Edward Poulton, and was allowed...

 described specimens from different areas of Persia as P. p. saxicolor in 1927, recognizing the similarity to P.p. ciscaucasica. Today, these names are considered synonyms.

Further reading

  • Khorozyan, I., Malkhasyan, A., Asmaryan, S. (2005) The Persian Leopard Prowls Its Way to Survival. Endangered Species Update 22 (2): 51-60 pdf
  • Kiabi, B. H., Dareshouri, B. F., Ghaemi, R. A., Jahanshahi, M. (2002) Population status of the Persian Leopard (Panthera pardus saxicolor Pocock, 1927) in Iran. Zoology in the Middle East 26: 41-47. abstract
  • Aghili, A. (2005) Leopard Survey in Caucasus Ecoregion (Northwest) of Iran. Leopard Conservation Society, Centre for Sustainable Development (CENESTA), Iran Department of Environment, Natural Environment & Biodiversity Office
  • Sanei, A. (2007). Analysis of leopard (Panthera pardus) status in Iran (No.1). Sepehr Publication Center (In Persian), Tehran, 298 p. ISBN:978-964-6123-74-8.
  • Woodroffe, R. (2000)Predators and people: using human densities to interpret declines of large carnivores. Animal Conservation (2000) 3: 165–173
  • Zulfiqar, A. (2001) Leopard in Pakistan’s North West Frontier Province. Cat News 35: 9–10.
  • Shakula, V. (2004) First record of leopard in Kazakhstan
    Kazakhstan
    Kazakhstan , officially the Republic of Kazakhstan, is a transcontinental country in Central Asia and Eastern Europe. Ranked as the ninth largest country in the world, it is also the world's largest landlocked country; its territory of is greater than Western Europe...

    . Cat News 41: 11–12
  • Gasparyan, K.M. and F.S. Agadjanyan. 1974. The panther in Armenia. Biological Journal of Armenia 27: 84–87
  • Janashvili, A. (1984) "Leopard". Georgian Soviet Encyclopedia. Vol. 11. Tbilisi, Pp. 567
  • Zakaria, M. and Sanei, A. (2011). Conservation and management prospects of the Persian and Malayan leopards. Asia Life Sciences Supplment 7: 1-5.

External links

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