Parastacidae
Encyclopedia
Parastacidae is the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 of freshwater
Freshwater
Fresh water is naturally occurring water on the Earth's surface in ice sheets, ice caps, glaciers, bogs, ponds, lakes, rivers and streams, and underground as groundwater in aquifers and underground streams. Fresh water is generally characterized by having low concentrations of dissolved salts and...

 crayfish
Crayfish
Crayfish, crawfish, or crawdads – members of the superfamilies Astacoidea and Parastacoidea – are freshwater crustaceans resembling small lobsters, to which they are related...

 found in the southern hemisphere
Southern Hemisphere
The Southern Hemisphere is the part of Earth that lies south of the equator. The word hemisphere literally means 'half ball' or "half sphere"...

. The family is a classic Gondwana
Gondwana
In paleogeography, Gondwana , originally Gondwanaland, was the southernmost of two supercontinents that later became parts of the Pangaea supercontinent. It existed from approximately 510 to 180 million years ago . Gondwana is believed to have sutured between ca. 570 and 510 Mya,...

-distributed taxon, with extant members in South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

, Madagascar
Madagascar
The Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...

, Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

, New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

 and New Guinea
New Guinea
New Guinea is the world's second largest island, after Greenland, covering a land area of 786,000 km2. Located in the southwest Pacific Ocean, it lies geographically to the east of the Malay Archipelago, with which it is sometimes included as part of a greater Indo-Australian Archipelago...

, and extinct taxa
Taxon
|thumb|270px|[[African elephants]] form a widely-accepted taxon, the [[genus]] LoxodontaA taxon is a group of organisms, which a taxonomist adjudges to be a unit. Usually a taxon is given a name and a rank, although neither is a requirement...

 also in Antarctica.

Three genera are to be found in Chile
Chile
Chile ,officially the Republic of Chile , is a country in South America occupying a long, narrow coastal strip between the Andes mountains to the east and the Pacific Ocean to the west. It borders Peru to the north, Bolivia to the northeast, Argentina to the east, and the Drake Passage in the far...

, Virilastacus, Samastacus and Parastacus, the last of which also occurs disjunctly in southern Brazil
Brazil
Brazil , officially the Federative Republic of Brazil , is the largest country in South America. It is the world's fifth largest country, both by geographical area and by population with over 192 million people...

.

There are no crayfish native to continental Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

, but six species on Madagascar
Madagascar
The Republic of Madagascar is an island country located in the Indian Ocean off the southeastern coast of Africa...

, all of the genus Astacoides
Astacoides
Astacoides is a genus of freshwater crayfish endemic to Madagascar. The first specimens were brought to Europe in 1839, and seven species are now recognised, most of which are on the IUCN Red List. They are large and slow-growing, and are threatened by habitat loss and overexploitation by local...

.
Australasia
Australasia
Australasia is a region of Oceania comprising Australia, New Zealand, the island of New Guinea, and neighbouring islands in the Pacific Ocean. The term was coined by Charles de Brosses in Histoire des navigations aux terres australes...

 is particularly rich in crayfish. The small genus Paranephrops
Paranephrops
Paranephrops is a genus of freshwater crayfish species endemic to New Zealand. They are known as ' in New Zealand, derived from the Māori language...

is endemic
Endemic (ecology)
Endemism is the ecological state of being unique to a defined geographic location, such as an island, nation or other defined zone, or habitat type; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, all species of lemur are endemic to the...

 to New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

. (Huxley claims that Paranephrops also occurs in Fiji
Fiji
Fiji , officially the Republic of Fiji , is an island nation in Melanesia in the South Pacific Ocean about northeast of New Zealand's North Island...

.) Two genera, Astacopsis and Parastacoides are endemic to Tasmania
Tasmania
Tasmania is an Australian island and state. It is south of the continent, separated by Bass Strait. The state includes the island of Tasmania—the 26th largest island in the world—and the surrounding islands. The state has a population of 507,626 , of whom almost half reside in the greater Hobart...

, while a further two are found on either side of the Bass Strait
Bass Strait
Bass Strait is a sea strait separating Tasmania from the south of the Australian mainland, specifically the state of Victoria.-Extent:The International Hydrographic Organization defines the limits of the Bass Strait as follows:...

 - Geocharax and Engaeus
Engaeus
Engaeus is a genus of freshwater crayfish found in Australia. Fifteen of the 35 species in the genus occur in Tasmania, where they are known as the Tasmanian land crayfish...

. The greatest diversity, however, is found on the Australian mainland
Mainland
Mainland is a name given to a large landmass in a region , or to the largest of a group of islands in an archipelago. Sometimes its residents are called "Mainlanders"...

. Three genera are endemic and have restricted distributions (Engaewa
Engaewa
Engaewa is a genus of burrowing crayfish belonging to the family Parastacidae. There are five recognised species, all endemic to the high-rainfall zone of south-west Western Australia, from Dunsborough to Albany. They are all small , and found only in freshwater swamps and seepages...

, Gramastacus and Tenuibranchiurus), while two are more widespread and contain nearly ninety species between them: Euastacus
Euastacus
Euastacus is a genus of freshwater crayfish known as "spiny crayfish". They are found in the south-east of the Australian mainland, along with another genus of crayfish, Cherax...

, around the Australian coast from Melbourne
Melbourne
Melbourne is the capital and most populous city in the state of Victoria, and the second most populous city in Australia. The Melbourne City Centre is the hub of the greater metropolitan area and the Census statistical division—of which "Melbourne" is the common name. As of June 2009, the greater...

 to Brisbane
Brisbane
Brisbane is the capital and most populous city in the Australian state of Queensland and the third most populous city in Australia. Brisbane's metropolitan area has a population of over 2 million, and the South East Queensland urban conurbation, centred around Brisbane, encompasses a population of...

, and Cherax
Cherax
Cherax is the largest and most widespread genus of fully and partially aquatic crayfish in the Southern Hemisphere. Its members may be found in lakes, rivers and streams across most of Australia and New Guinea. In Australia the many species of Cherax are commonly known as yabbies...

across Australia and New Guinea. One fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

 species, Aenigmastacus crandalli
Aenigmastacus
Aenigmastacus crandalli is a species of fossil freshwater crayfish. It was found in Eocene lake deposits in British Columbia, and was described in 2011. It is the first member of the Gondwana-distributed family Parastacidae to be found in the Northern Hemisphere, and is the only species in the...

, is known from Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

.
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