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Parasitic wasp

 

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Parasitic wasp



 
 
The term Parasitoid wasp refers to a large evolutionary grade
Evolutionary grade

In alpha taxonomy, a grade refers to a level of morphology and/or physiological complexity. Organisms may be grouped by the grade of organisation they display without making any implications about their phylogenetic relationship....
 of hymenoptera
Hymenoptera

Hymenoptera is one of the larger order s of insects, comprising the sawfly, wasps, bees, and ants. The name refers to the membranous wings of the insects, and is derived from the Ancient Greek language wikt:???? : membrane and wikt:pte??? : wing....
n superfamilies, mainly in the Apocrita
Apocrita

Apocrita is a suborder of insects in the order Hymenoptera.The Apocrita includes wasps, bees and ants, and consists of many families. It includes the most advanced Hymenoptera and is distinguished from the Symphyta by the narrow "waist" formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the...
. They are primarily parasitoids of other animals, mostly other arthropod
Arthropod

Arthropods are animals belonging to the Scientific classification Arthropoda , and include the insects, arachnids, crustaceans, and others....
s. Many of them, such as the family Braconidae
Braconidae

Braconidae is a family of parasitoid wasps and one of the richest family of insects. From the approximate 12,000 described species , it is extrapolation that between 40,000 and 50,000 species exist worldwide....
, are considered beneficial to humans because they control
Biological pest control

Biological control of pests in agriculture is a method of pest control that relies on predation, parasitism, herbivory, or other natural mechanisms....
 populations of agricultural pests. Some of these wasps help pest control in a rather sophisticated manner. Certain types of plants have compounds that work in part with the saliva of caterpillars.






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The term Parasitoid wasp refers to a large evolutionary grade
Evolutionary grade

In alpha taxonomy, a grade refers to a level of morphology and/or physiological complexity. Organisms may be grouped by the grade of organisation they display without making any implications about their phylogenetic relationship....
 of hymenoptera
Hymenoptera

Hymenoptera is one of the larger order s of insects, comprising the sawfly, wasps, bees, and ants. The name refers to the membranous wings of the insects, and is derived from the Ancient Greek language wikt:???? : membrane and wikt:pte??? : wing....
n superfamilies, mainly in the Apocrita
Apocrita

Apocrita is a suborder of insects in the order Hymenoptera.The Apocrita includes wasps, bees and ants, and consists of many families. It includes the most advanced Hymenoptera and is distinguished from the Symphyta by the narrow "waist" formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the...
. They are primarily parasitoids of other animals, mostly other arthropod
Arthropod

Arthropods are animals belonging to the Scientific classification Arthropoda , and include the insects, arachnids, crustaceans, and others....
s. Many of them, such as the family Braconidae
Braconidae

Braconidae is a family of parasitoid wasps and one of the richest family of insects. From the approximate 12,000 described species , it is extrapolation that between 40,000 and 50,000 species exist worldwide....
, are considered beneficial to humans because they control
Biological pest control

Biological control of pests in agriculture is a method of pest control that relies on predation, parasitism, herbivory, or other natural mechanisms....
 populations of agricultural pests. Some of these wasps help pest control in a rather sophisticated manner. Certain types of plants have compounds that work in part with the saliva of caterpillars. When the saliva of the caterpillar and the juices of the plant mix, a fragrance is emitted that certain parasitoid wasps are very attracted to. The parasitoid wasps then kill the caterpillars and often use the carcasses to lay eggs within. This is a form of mutualism between the plant and the wasp and works only when the fragrance from the plant and the saliva combine.

Taxonomy and systematics

Historically, the classification system of Hymenoptera included two divisions or infraorders within the suborder Apocrita
Apocrita

Apocrita is a suborder of insects in the order Hymenoptera.The Apocrita includes wasps, bees and ants, and consists of many families. It includes the most advanced Hymenoptera and is distinguished from the Symphyta by the narrow "waist" formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the...
: one of these, the "Parasitica" containing the parasitoid wasp. However, the use of the name Parasitica (or its alternative, "Terebrantia") has been phased out in recent years, as it is a paraphyletic grouping, and most modern classifications explicitly reject the use of any groups that are not monophyletic.

Presently, it is not clear what the eventual taxonomic fate of these groups will be. A number of clade
Clade

A clade is a term used in modern alpha taxonomy, the scientific classification of living and fossil organisms, to describe a monophyletic group, defined as a group consisting of a single common ancestor and all its descendants.The term "monophyletic group" is used in this article in the conventional sense of "an a...
s seem to stand out in newer studies, and these may be treated as unranked taxa or at the ranks of infraorder and division.

Superfamilies

The traditional superfamilies united in the "Parasitica" are:
  • Superfamily Ceraphronoidea
    Ceraphronoidea

    Ceraphronoidea is a small Hymenopteran superfamily that includes only two families, and a total of some 800 species, though a great many species are still undescribed....
  • Superfamily Chalcidoidea
  • Superfamily Cynipoidea
    Cynipoidea

    Cynipoidea is a moderate-sized Hymenopteran superfamily that presently includes five modern families and three extinct families, though others have been recognized in the past....
  • Superfamily Evanioidea
    Evanioidea

    Evanioidea is a small Hymenopteran superfamily that includes three families, two of which are much more closely related to one another than they are to the remaining family, Evaniidae....
  • Superfamily Ichneumonoidea
  • Superfamily Megalyroidea
  • Superfamily Mymarommatoidea (sometimes included in Serphitoidea)
  • Superfamily Platygastroidea
    Platygastroidea

    The Hymenopteran superfamily Platygastroidea has, in the past, often been treated as a lineage within the superfamily Proctotrupoidea, but most classifications since 1977 have recognized it as an independent group, composed of two families, the Platygastridae and the Scelionidae, with a combined diversity of some 4000 described species....
  • Superfamily Proctotrupoidea
    Proctotrupoidea

    The Hymenopteran superfamily Proctotrupoidea is a somewhat confusing assemblage of taxa, with new families being added with surprising frequency, and very little to unify them all into a single natural group....
  • Superfamily Serphitoidea (fossil
    Fossil

    Fossils are the preserved remains or trace fossil of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous Rock formations and sedimentary rock layers is known as the fossil record....
    )
  • Superfamily Stephanoidea
  • Superfamily Trigonaloidea


Other than the "Parasitica", there are a few Apocrita
Apocrita

Apocrita is a suborder of insects in the order Hymenoptera.The Apocrita includes wasps, bees and ants, and consists of many families. It includes the most advanced Hymenoptera and is distinguished from the Symphyta by the narrow "waist" formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the...
 that are also sometimes called "parasitic wasps": most of the members of the superfamily Chrysidoidea
Chrysidoidea

The superfamily Chrysidoidea or Bethyloidea is a very large cosmopolitan group of parasitoid or cleptoparasite wasps, with three large, common families and four tiny, rare families....
, as well as most of the families superfamily Vespoidea
Vespoidea

Vespoidea is a Scientific classification of Order Hymenoptera of Class Insecta, although older Taxonomy schemes may vary in this categorization, particularly in their recognition of a now-obsolete superfamily Scolioidea....
 such as Bradynobaenidae, Mutillidae
Mutillidae

Mutillidae, or velvet ants, are a family of wasps whose wingless females resemble ants, though only distantly related. The "velvet ant" name refers to their hair, which may be red, black, white, silvery or golden....
, Rhopalosomatidae
Rhopalosomatidae

Rhopalosomatidae is a family of Hymenoptera. It contains 37 extant species in four genera. Two fossil genera are known....
, Sapygidae
Sapygidae

The Sapygidae are a family of solitary aculeate wasps. There does not seem to be a common English name, but Club-horned wasps seems as good a name as any, though various other groups of wasps also have clubbed or thickened antennae....
, Scoliidae
Scoliidae

Scoliidae, the scoliid wasps, is a small family represented by 6 genera and about 20 species in North America, but they occur worldwide, with a total of around 300 species....
, Sierolomorphidae
Sierolomorphidae

The Sierolomorphidae are a family of about 10 known species of wasps found in Tropical America and Asia. They are rare and very little is known of their biology....
, Tiphiidae
Tiphiidae

Tiphiidae is a family of large solitary wasps whose larvae are almost universally parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea....
, and a few species of Pompilidae. Most of these groups are ectoparasitoids. Among the sawflies
Sawfly

Sawfly is the common name for is an insect in the order Hymenoptera, suborder Symphyta.Sawflies are distinguishable from most other Hymenoptera by the broad connection between the abdomen and the thorax , and the caterpillar-like larvae ....
, which are not Apocrita, there is only one small parasitic family, Orussidae
Orussidae

The family Orussidae is the only Symphytan group which is parasitoid, thus giving them the common name parasitic wood wasps. They are an ancient group, well-represented in the fossil record, and are believed to represent a sort of "missing link" within the order Hymenoptera, as they are often considered to be the sister taxon to the Apocrita...
. This is the Apocrita's closest living relative. It thus appears that the ancestors of bee
Bee

Bees are flying insects closely related to wasps and ants. Bees are a monophyly lineage within the superfamily Apoidea, presently classified by the unranked taxon name Anthophila....
s and bumblebee
Bumblebee

A bumblebee is any member of the bee genus Bombus, in the family Apidae; there are over 250 known species primarily occurring in the Northern Hemisphere....
s were parasitic too.

Systematics

According to recent cladistic studies, there are a number of basal lineages among the Apocrita, as well as a diverse group of parasitic wasps that seems to form a major clade. If the Apocrita are divided into infraorders, about 6 of these must be recognized, and several of these warrant further subdivision, with a number of families being moved out of the Proctotrupoidea
Proctotrupoidea

The Hymenopteran superfamily Proctotrupoidea is a somewhat confusing assemblage of taxa, with new families being added with surprising frequency, and very little to unify them all into a single natural group....
:

Ic Ichneumonidae
* Superfamily Ichneumonoidea seems closer to the Aculeata
Aculeata

The name Aculeata is used to refer to a monophyletic lineage of Hymenoptera. The word "Aculeata" is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger ....
 than to other parasitic wasps. If the Aculeata are treated as a division, the Ichneumonoidea would form a basal superfamily in a new infraorder.
  • Superfamily Stephanoidea forms a clade or infraorder of its own.
  • Another clade or infraorder contains the following groups:
    • Superfamily Megalyroidea
    • Superfamily Trigonaloidea
    • a clade or division containing
      • Superfamily Ceraphronoidea
        Ceraphronoidea

        Ceraphronoidea is a small Hymenopteran superfamily that includes only two families, and a total of some 800 species, though a great many species are still undescribed....
      • Superfamily Evanioidea
        Evanioidea

        Evanioidea is a small Hymenopteran superfamily that includes three families, two of which are much more closely related to one another than they are to the remaining family, Evaniidae....
        .
  • A clade or infraorder containing families formerly in the Proctotrupoidea:
    • Family Maamingidae
    • a clade, division or superfamily containing
      • Family Austroniidae
      • Family Diapriidae
        Diapriidae

        Diapriidae is a family of insects belonging to the order Hymenoptera. These tiny wasps are typically parasitoids on the larvae and pupae of a wide range of insects, especially fly; a few are hyperparasitoids ....
      • Family Monomachidae.
  • Another clade or infraorder of families formerly in the Proctotrupoidea:
    • Family Proctorenyxidae
    • Family Roproniidae


  • The bulk of the parasitic wasps, a clade or infraorder containing two very distinct groups:
    • One clade or division containing:
      • Superfamily Platygastroidea
        Platygastroidea

        The Hymenopteran superfamily Platygastroidea has, in the past, often been treated as a lineage within the superfamily Proctotrupoidea, but most classifications since 1977 have recognized it as an independent group, composed of two families, the Platygastridae and the Scelionidae, with a combined diversity of some 4000 described species....
      • Superfamily Chalcidoidea
      • Superfamily Mymarommatoidea (sometimes included in Serphitoidea)
      • Superfamily Serphitoidea (fossil
        Fossil

        Fossils are the preserved remains or trace fossil of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossiliferous Rock formations and sedimentary rock layers is known as the fossil record....
        , tentatively placed here).
    • Another clade or division containing:
      • Superfamily Cynipoidea
        Cynipoidea

        Cynipoidea is a moderate-sized Hymenopteran superfamily that presently includes five modern families and three extinct families, though others have been recognized in the past....
      • Superfamily Proctotrupoidea
        Proctotrupoidea

        The Hymenopteran superfamily Proctotrupoidea is a somewhat confusing assemblage of taxa, with new families being added with surprising frequency, and very little to unify them all into a single natural group....
         sensu stricto


External links

  • at Texas Cooperative Extension. Reprinted from Drees, Bastiaan M. and John A. Jackman. A Field Guide to Common Texas Insects. Houston: Gulf Publishing Company, 1999. ISBN 0877192634.
  • : Apocrita
  • Pictorial overview.In easy Spanish