Paraptenodytes
Encyclopedia
Paraptenodytes is an extinct genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 of penguins
Penguin
Penguins are a group of aquatic, flightless birds living almost exclusively in the southern hemisphere, especially in Antarctica. Highly adapted for life in the water, penguins have countershaded dark and white plumage, and their wings have become flippers...

 which contains two or three species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 sized between a Magellanic Penguin
Magellanic Penguin
The Magellanic Penguin, Spheniscus magellanicus, is a South American penguin, breeding in coastal Argentina, Chile and the Falkland Islands, with some migrating to Brazil where they are occasionally seen as far north as Rio de Janeiro. It is the most numerous of the Spheniscus penguins. Its nearest...

 and an Emperor Penguin
Emperor Penguin
The Emperor Penguin is the tallest and heaviest of all living penguin species and is endemic to Antarctica. The male and female are similar in plumage and size, reaching in height and weighing anywhere from . The dorsal side and head are black and sharply delineated from the white belly,...

 (Aptenodytes forsteri). They are known from fossil
Fossil
Fossils are the preserved remains or traces of animals , plants, and other organisms from the remote past...

 bones ranging from a partial skeleton and some additional material in the case of P. antarcticus, and a single humerus
Humerus
The humerus is a long bone in the arm or forelimb that runs from the shoulder to the elbow....

 in the case of P. brodkorbi. The latter species is therefore often considered invalid; a recent study considers it indeed valid, but distinct enough not to belong into Paraptenodytes. The fossils were found in the Santa Cruz
Santa Cruz Province (Argentina)
Santa Cruz is a province of Argentina, located in the southern part of the country, in Patagonia. It borders Chubut province to the north, and Chile to the west and south. To the east is the Atlantic Ocean...

 and Chubut Province
Chubut Province
Chubut a province in the southern part of Argentina situated between the 42nd parallel south and the 46th parallel south , the Andes range separating Argentina from Chile, and the Atlantic ocean...

s of Patagonia
Patagonia
Patagonia is a region located in Argentina and Chile, integrating the southernmost section of the Andes mountains to the southwest towards the Pacific ocean and from the east of the cordillera to the valleys it follows south through Colorado River towards Carmen de Patagones in the Atlantic Ocean...

, Argentina
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

, in Patagonian Molasse Formation rocks of Early Miocene
Miocene
The Miocene is a geological epoch of the Neogene Period and extends from about . The Miocene was named by Sir Charles Lyell. Its name comes from the Greek words and and means "less recent" because it has 18% fewer modern sea invertebrates than the Pliocene. The Miocene follows the Oligocene...

 age; later occurrences are apparently from Late Miocene or possibly even Early Pliocene
Pliocene
The Pliocene Epoch is the period in the geologic timescale that extends from 5.332 million to 2.588 million years before present. It is the second and youngest epoch of the Neogene Period in the Cenozoic Era. The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch...

 deposits.

Together with the related genus Arthrodytes, they form the subfamily Paraptenodytinae, which is not an ancestor of modern penguins.
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