Pandanus tectorius
Encyclopedia
Pandanus tectorius is a species of Pandanus
Pandanus
Pandanus is a genus of monocots with about 600 known species. They are numerous palmlike dioecious trees and shrubs native of the Old World tropics and subtropics. They are classified in the order Pandanales, family Pandanaceae.-Overview:...

(screwpine) that is native to Malesia
Malesia
Malesia is a biogeographical region straddling the boundaries of the Indomalaya ecozone and Australasia ecozone, and also a phytogeographical floristic region in the Paleotropical Kingdom.-Floristic province:...

, eastern Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

, and the Pacific Islands
Pacific Islands
The Pacific Islands comprise 20,000 to 30,000 islands in the Pacific Ocean. The islands are also sometimes collectively called Oceania, although Oceania is sometimes defined as also including Australasia and the Malay Archipelago....

. Common names include Thatch Screwpine, Hala (Hawaiian
Hawaiian language
The Hawaiian language is a Polynesian language that takes its name from Hawaii, the largest island in the tropical North Pacific archipelago where it developed. Hawaiian, along with English, is an official language of the state of Hawaii...

), Bacua (Spanish
Spanish language
Spanish , also known as Castilian , is a Romance language in the Ibero-Romance group that evolved from several languages and dialects in central-northern Iberia around the 9th century and gradually spread with the expansion of the Kingdom of Castile into central and southern Iberia during the...

), and Vacquois (French
French language
French is a Romance language spoken as a first language in France, the Romandy region in Switzerland, Wallonia and Brussels in Belgium, Monaco, the regions of Quebec and Acadia in Canada, and by various communities elsewhere. Second-language speakers of French are distributed throughout many parts...

).

Description

P. tectorius is a tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 that to grows to 4–14 m (13.1–45.9 ft) tall. The single trunk
Trunk (botany)
In botany, trunk refers to the main wooden axis of a tree that supports the branches and is supported by and directly attached to the roots. The trunk is covered by the bark, which is an important diagnostic feature in tree identification, and which often differs markedly from the bottom of the...

 is spiny and forks at a height of 4–8 m (13.1–26.2 ft). It is supported by prop root
Aerial root
Aerial roots are roots above the ground. They are almost always adventitious. They are found in diverse plant species, including epiphytes such as orchids, tropical coastal swamp trees such as mangroves, the resourceful banyan trees, the warm-temperate rainforest rātā and pōhutukawa Aerial roots...

s that firmly anchor the tree to the ground. Its leaves are usually 90–150 cm (3–4.9 ft) long and 5–7 cm (2–2.8 in) wide with saw-like margins.

Flowers

P. tectorius is dioecious
Plant sexuality
Plant sexuality covers the wide variety of sexual reproduction systems found across the plant kingdom. This article describes morphological aspects of sexual reproduction of plants....

, with very different male and female flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s. Male flowers are small, fragrant, form clusters or raceme
Raceme
A raceme is a type of inflorescence that is unbranched and indeterminate and bears pedicellate flowers — flowers having short floral stalks called pedicels — along the axis. In botany, axis means a shoot, in this case one bearing the flowers. In a raceme, the oldest flowers are borne...

s, and short lived, lasting only a single day. Female flowers resemble pineapple
Pineapple
Pineapple is the common name for a tropical plant and its edible fruit, which is actually a multiple fruit consisting of coalesced berries. It was given the name pineapple due to its resemblance to a pine cone. The pineapple is by far the most economically important plant in the Bromeliaceae...

s.

Fruit

The Thatch Screwpine's fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

 is either ovoid
Oval
An oval is any curve resembling an egg or an ellipse, such as a Cassini oval. The term does not have a precise mathematical definition except in one area oval , but it may also refer to:* A sporting arena of oval shape** a cricket field...

, ellipsoid, subglobose or globose
Sphere
A sphere is a perfectly round geometrical object in three-dimensional space, such as the shape of a round ball. Like a circle in two dimensions, a perfect sphere is completely symmetrical around its center, with all points on the surface lying the same distance r from the center point...

 with a diameter of 4–20 cm (1.6–7.9 in) and a length of 8–30 cm (3.1–11.8 in). The fruit is made up of 38–200 wedge-like phalanges, which have an outer fibrous husk
Husk
Husk in botany is the outer shell or coating of a seed. It often refers to the leafy outer covering of an ear of maize as it grows on the plant. Literally, a husk or hull includes the protective outer covering of a seed, fruit or vegetable...

. Phalanges contain two seeds on average, with a maximum of eight reported. The phalanges are buoyant
Buoyancy
In physics, buoyancy is a force exerted by a fluid that opposes an object's weight. In a column of fluid, pressure increases with depth as a result of the weight of the overlying fluid. Thus a column of fluid, or an object submerged in the fluid, experiences greater pressure at the bottom of the...

, and the seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s within them can remain viable for many months while being transported
Drift seed
Drift seeds and drift fruits are seeds and fruits adapted for long distance dispersal by water. Most are produced by tropical trees, and they can be found on distant beaches after drifting thousands of miles through ocean currents...

 by ocean current
Ocean current
An ocean current is a continuous, directed movement of ocean water generated by the forces acting upon this mean flow, such as breaking waves, wind, Coriolis effect, cabbeling, temperature and salinity differences and tides caused by the gravitational pull of the Moon and the Sun...

s.

Habitat

P. tectorius naturally grows in coast
Coast
A coastline or seashore is the area where land meets the sea or ocean. A precise line that can be called a coastline cannot be determined due to the dynamic nature of tides. The term "coastal zone" can be used instead, which is a spatial zone where interaction of the sea and land processes occurs...

al regions, such as on mangrove
Mangrove
Mangroves are various kinds of trees up to medium height and shrubs that grow in saline coastal sediment habitats in the tropics and subtropics – mainly between latitudes N and S...

 margins and beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

es, at elevations from sea level to 610 m (2,001.3 ft). It requires 1500 – of annual rainfall. Thatch Screwpine is well adapted to grow in the many soil
Soil
Soil is a natural body consisting of layers of mineral constituents of variable thicknesses, which differ from the parent materials in their morphological, physical, chemical, and mineralogical characteristics...

 types present on coasts, including quartz
Quartz
Quartz is the second-most-abundant mineral in the Earth's continental crust, after feldspar. It is made up of a continuous framework of SiO4 silicon–oxygen tetrahedra, with each oxygen being shared between two tetrahedra, giving an overall formula SiO2. There are many different varieties of quartz,...

 sand
Sand
Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.The composition of sand is highly variable, depending on the local rock sources and conditions, but the most common constituent of sand in inland continental settings and non-tropical coastal...

, coral sand
Coral sand
Coral sand is sand of particles originating in tropical and sub-tropical marine environments from bioerosion of limestone skeletal material of marine organisms. One example of this process is that of parrot fishes which bite off pieces of coral, digest the living tissue, and excrete the inorganic...

, and peat
Peat
Peat is an accumulation of partially decayed vegetation matter or histosol. Peat forms in wetland bogs, moors, muskegs, pocosins, mires, and peat swamp forests. Peat is harvested as an important source of fuel in certain parts of the world...

, as well as in limestone
Limestone
Limestone is a sedimentary rock composed largely of the minerals calcite and aragonite, which are different crystal forms of calcium carbonate . Many limestones are composed from skeletal fragments of marine organisms such as coral or foraminifera....

 and basalt
Basalt
Basalt is a common extrusive volcanic rock. It is usually grey to black and fine-grained due to rapid cooling of lava at the surface of a planet. It may be porphyritic containing larger crystals in a fine matrix, or vesicular, or frothy scoria. Unweathered basalt is black or grey...

. P. tectorius is salt
Halophyte
A halophyte is a plant that grows where it is affected by salinity in the root area or by salt spray, such as in saline semi-deserts, mangrove swamps, marshes and sloughs, and seashores. An example of a halophyte is the salt marsh grass Spartina alterniflora . Relatively few plant species are...

 and wind tolerant and favors slightly acid
Acid
An acid is a substance which reacts with a base. Commonly, acids can be identified as tasting sour, reacting with metals such as calcium, and bases like sodium carbonate. Aqueous acids have a pH of less than 7, where an acid of lower pH is typically stronger, and turn blue litmus paper red...

ic to basic
Base (chemistry)
For the term in genetics, see base A base in chemistry is a substance that can accept hydrogen ions or more generally, donate electron pairs. A soluble base is referred to as an alkali if it contains and releases hydroxide ions quantitatively...

 soil (pH
Soil pH
The soil pH is a measure of the acidity or basicity in soils. pH is defined as the negative logarithm of the activity of hydrogen ions in solution. It ranges from 0 to 14, with 7 being neutral. A pH below 7 is acidic and above 7 is basic. Soil pH is considered a master variable in soils as it...

 of 6-10). It prefers to grow in full sunlight, but grows well with 30-50% shade.

Range

P. tectorius occurs from near Port Macquarie in New South Wales
New South Wales
New South Wales is a state of :Australia, located in the east of the country. It is bordered by Queensland, Victoria and South Australia to the north, south and west respectively. To the east, the state is bordered by the Tasman Sea, which forms part of the Pacific Ocean. New South Wales...

 to northern Queensland
Queensland
Queensland is a state of Australia, occupying the north-eastern section of the mainland continent. It is bordered by the Northern Territory, South Australia and New South Wales to the west, south-west and south respectively. To the east, Queensland is bordered by the Coral Sea and Pacific Ocean...

, Australia
Australia
Australia , officially the Commonwealth of Australia, is a country in the Southern Hemisphere comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands in the Indian and Pacific Oceans. It is the world's sixth-largest country by total area...

 and Indonesia
Indonesia
Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

 east through the islands
Pacific Islands
The Pacific Islands comprise 20,000 to 30,000 islands in the Pacific Ocean. The islands are also sometimes collectively called Oceania, although Oceania is sometimes defined as also including Australasia and the Malay Archipelago....

 of the tropical Pacific Ocean
Pacific Ocean
The Pacific Ocean is the largest of the Earth's oceanic divisions. It extends from the Arctic in the north to the Southern Ocean in the south, bounded by Asia and Australia in the west, and the Americas in the east.At 165.2 million square kilometres in area, this largest division of the World...

 to Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

. Its exact native range is unknown due to extensive cultivation; it may be an early Polynesian
Polynesian culture
Polynesian culture refers to the indigenous peoples' culture of Polynesia who share common traits in language, customs and society. Chronologically, the development of Polynesian culture can be divided into four different historical eras:...

 introduction
Canoe plants
Canoe plants, or Polynesian introductions, are plants taken from ancient Polynesia and transplanted to other Pacific IslandsThe term is particularly used to refer to plants brought to Hawaii 1,700 years ago by Polynesian explorers....

 to many of the more isolated Pacific islands on which it occurs. P. tectorius is known to have predated human settlement on Hawaii due to seed and pollen samples taken from Kauai
Kauai
Kauai or Kauai, known as Tauai in the ancient Kaua'i dialect, is geologically the oldest of the main Hawaiian Islands. With an area of , it is the fourth largest of the main islands in the Hawaiian archipelago, and the 21st largest island in the United States. Known also as the "Garden Isle",...

's Makauwahi Cave
Makauwahi Cave
The Makauwahi Cave is the largest limestone cave found in Hawaii. It lies on the south coast of the island of Kauai, in the Māhāulepū Valley close to Māhāulepū Beach, and is important for its paleoecological and archaeological values...

.

Uses

The fruit can be eaten raw or cooked and is a major source of food in Micronesia
Micronesia
Micronesia is a subregion of Oceania, comprising thousands of small islands in the western Pacific Ocean. It is distinct from Melanesia to the south, and Polynesia to the east. The Philippines lie to the west, and Indonesia to the southwest....

, especially in the atoll
Atoll
An atoll is a coral island that encircles a lagoon partially or completely.- Usage :The word atoll comes from the Dhivehi word atholhu OED...

s. The fibrous nature of the fruit also serves as a natural dental floss
Dental floss
Dental floss is made of either a bundle of thin nylon filaments or a plastic ribbon used to remove food and dental plaque from teeth. The floss is gently inserted between the teeth and scraped along the teeth sides, especially close to the gums. Dental floss may be flavored or unflavored, and...

. The tree's leaves are often used as flavoring for sweet dishes such as kaya jam, and are also said to have medicinal properties. It is also used in Sri Lankan cookery, where the leaves are used to flavour a variety of curries. Leaves were used by the Polynesians
Polynesians
The Polynesian peoples is a grouping of various ethnic groups that speak Polynesian languages, a branch of the Oceanic languages within the Austronesian languages, and inhabit Polynesia. They number approximately 1,500,000 people...

 to make baskets, mats, outrigger canoe
Outrigger canoe
The outrigger canoe is a type of canoe featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull...

 sails, thatch
Thatching
Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge , rushes, or heather, layering the vegetation so as to shed water away from the inner roof. It is a very old roofing method and has been used in both tropical and temperate climates...

 roofs, and grass skirt
Grass skirt
A grass skirt is a traditional part of dress wear on tropical islands all over the world . This is referred to many kinds of dress wear. These are popular with Hula girls on Hawaii. Many of these are seen at Hawaiian parties....

s.

A large shrub or small tree of immense cultural, health, and economic importance in the Pacific, second only to coconut on atolls. It grows wild mainly in semi­natural vegetation in littoral habitats throughout the tropical and subtropical Pacific, where it can withstand drought, strong winds, and salt spray. It propagates readily from seed, but it is also widely propagated from branch cuttings by local people for farms and homegardens. It grows fairly quickly, and all parts are used, from the nutritious fruits of edible varieties, to the poles and branches in construction, to the leaves for weaving and garlands. The plant is prominent in Pacific culture and tradition, including local medicine.
Hundreds of cultivated varieties, are known by their local names and character­istics of fruits, branches, and leaves. At present, there is evidence that this diversity is declining, with certain va­rieties becoming difficult to find. The reasons include less replanting, deforestation, fire, flagging interest by the new generation, and rapid population growth leading to urban­ization. Planting pandanus should be promoted in both rural and urban areas. Special attempts should be made to collect and replant endangered varieties that are valued for their edible fruits and other uses.

Culture

The seal of Punahou School
Punahou School
Punahou School, once known as Oahu College, is a private, co-educational, college preparatory school located in Honolulu CDP, City and County of Honolulu in the U.S. State of Hawaii...

 in Honolulu, Hawaii
Hawaii
Hawaii is the newest of the 50 U.S. states , and is the only U.S. state made up entirely of islands. It is the northernmost island group in Polynesia, occupying most of an archipelago in the central Pacific Ocean, southwest of the continental United States, southeast of Japan, and northeast of...

 features the hala tree, in part because lauhala
Lauhala
Lauhala, lau meaning "leaf" in the Hawaiian language, refers to the leaves of the hala tree .-Uses:The hala tree is of great cultural, health and economic importance in many Pacific Islands. The fruit of the tree is used as a food source in many Pacific Islands. The fruits are oftentimes consumed...

, the art of weaving with the leaves of that tree, is pivotal to the history of the island
History of Hawaii
The human history of Hawaii includes phases of early Polynesian settlement, British arrival, unification, Euro-American and Asian immigrators, the overthrow of the Hawaiian monarchy, a brief period as the Republic of Hawaii, and admission to the United States as Hawaii Territory and then as the...

, with everything from houses to pillows being made in this fashion. Local legend tells of an aged Hawaiian couple who lived long ago above the present Punahou campus, and had to travel far for water. They prayed each night for a spring, but to no avail. Finally one night, in a dream answering their prayers, they were told to uproot the stump of an old hala tree. They did as they were told and found a spring of clear, sweet water, which they named Ka Punahou, the New Spring. According to legend, Punahou School's lily pond is fed by this same spring.

Ecology

The stick insect
Phasmatidae
Phasmatidae is a family of the stick insects . They belong to the superfamily Anareolatae of suborder Verophasmatodea.Like many of their relatives, Phasmatidae are capable of regenerating limbs and commonly reproduce by parthenogenesis...

 Megacrania batesii
Megacrania batesii
The Peppermint Stick Insect is a bluish-green coloured stick insect that only lives on the midribs of the leaves of the "screwpine" Pandanus tectorius. If disturbed during the day, it emits a milky substance with a smell resembling peppermint...

lives and feeds only on this species.
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