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Osmoregulation



 
 
Osmoregulation is the active regulation of the osmotic pressure
Osmotic pressure

Osmotic pressure is the Fluid_statics#Hydrostatic_pressure produced by a difference in concentration between solutions on the two sides of a surface such as a differentially permeable membrane....
 of bodily fluids to maintain the homeostasis
Homeostasis

Homeostasis is the property of a system, either open system or closed system, that regulates its internal environment and tends to maintain a stable, constant condition....
 of the body's water
Water

Water is a common chemical substance that is essential for the survival of all known forms of life. In typical usage, water refers only to its liquid form or States of matter, but the substance also has a solid state, ice, and a gaseous state, water vapor or steam....
 content; that is it keeps the body's fluids from becoming too dilute or too concentrated. Osmotic pressure is a measure of the tendency of water to move into one solution
Solution

In chemistry, a solution is a homogeneous mixture composed of two or more substances. In such a mixture, a solute is dissolved in another substance, known as a solvent....
 from another by osmosis
Osmosis

Osmosis is the diffusion of a solvent through a Semipermeable membrane, from a solution of low solute concentration to a solution with high solute concentration , up a solute concentration gradient....
. The higher the osmotic pressure of a solution the more water wants to go into the solution. Pressure must be exerted on the hypertonic
Tonicity

Tonicity measures the ability of a solution to exert an osmotic pressure upon the membrane. Osmolality and osmolarity measure concentration of the solutes independently on their ability to cross the membrane....
 side of a selectively-permeable membrane
Biological membrane

A biological membrane or biomembrane is an enclosing or separating amphipathic layer that acts as a barrier within or around a cell . It is, almost invariably, a lipid bilayer, composed of a double layer of lipid-class molecules, specifically phospholipids and cholesterol, with occasional integral membrane protein intertwined, some o...
 to prevent diffusion
Diffusion

Molecular diffusion, often called simply diffusion, is a net transport of molecules from a region of higher concentration to one of lower concentration by random molecular motion....
 of water by osmosis from the side containing pure water.

Organisms in all environments (aquatic and terrestrial) must maintain the right concentration of solutes and amount of water in their body fluids; this involves excretion
Excretion

Excretion is the process of eliminating waste products of metabolism and other non-useful materials. It is an essential process in all forms of life....
: getting rid of metabolic wastes and other substances such as hormone
Hormone

Hormones are chemicals released by cells that affect cells in other parts of the body. Only a small amount of hormone is required to alter cell metabolism....
s that would be toxic if allowed to accumulate in the blood
Blood

Blood is a specialized bodily fluid that delivers necessary substances to the body's Cell s ? such as nutrients and oxygen ? and transports waste products away from those same cells....
 via organs such as the skin
Skin

The skin is the outer covering of the body, also known as the epidermis. It is the largest organ of the integumentary system made up of multiple layers of epithelial biological tissue, and guards the underlying muscles, bones, ligaments and organ s....
 and the kidneys; keeping the water and dissolved solutes in balance is referred to as osmoregulation.

a class="link1" onMouseover='showByLink("m3713281",this)' onMouseout='hide("m3713281")'href="http://www.absoluteastronomy.com/topics/Osmoconformer">Osmoconformer
Osmoconformer

An osmoconformer is a marine invertebrate that maintains its internal salinity such that it is always equal to the surrounding seawater. Osmoconformers do not actively exchange solutes with the environment, but keep their body fluids tonicity to the external environment by actively regulating their internal concentration of amino acids, ions,...
s match their body osmolarity to their environment .






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Osmoregulation is the active regulation of the osmotic pressure
Osmotic pressure

Osmotic pressure is the Fluid_statics#Hydrostatic_pressure produced by a difference in concentration between solutions on the two sides of a surface such as a differentially permeable membrane....
 of bodily fluids to maintain the homeostasis
Homeostasis

Homeostasis is the property of a system, either open system or closed system, that regulates its internal environment and tends to maintain a stable, constant condition....
 of the body's water
Water

Water is a common chemical substance that is essential for the survival of all known forms of life. In typical usage, water refers only to its liquid form or States of matter, but the substance also has a solid state, ice, and a gaseous state, water vapor or steam....
 content; that is it keeps the body's fluids from becoming too dilute or too concentrated. Osmotic pressure is a measure of the tendency of water to move into one solution
Solution

In chemistry, a solution is a homogeneous mixture composed of two or more substances. In such a mixture, a solute is dissolved in another substance, known as a solvent....
 from another by osmosis
Osmosis

Osmosis is the diffusion of a solvent through a Semipermeable membrane, from a solution of low solute concentration to a solution with high solute concentration , up a solute concentration gradient....
. The higher the osmotic pressure of a solution the more water wants to go into the solution. Pressure must be exerted on the hypertonic
Tonicity

Tonicity measures the ability of a solution to exert an osmotic pressure upon the membrane. Osmolality and osmolarity measure concentration of the solutes independently on their ability to cross the membrane....
 side of a selectively-permeable membrane
Biological membrane

A biological membrane or biomembrane is an enclosing or separating amphipathic layer that acts as a barrier within or around a cell . It is, almost invariably, a lipid bilayer, composed of a double layer of lipid-class molecules, specifically phospholipids and cholesterol, with occasional integral membrane protein intertwined, some o...
 to prevent diffusion
Diffusion

Molecular diffusion, often called simply diffusion, is a net transport of molecules from a region of higher concentration to one of lower concentration by random molecular motion....
 of water by osmosis from the side containing pure water.

Organisms in all environments (aquatic and terrestrial) must maintain the right concentration of solutes and amount of water in their body fluids; this involves excretion
Excretion

Excretion is the process of eliminating waste products of metabolism and other non-useful materials. It is an essential process in all forms of life....
: getting rid of metabolic wastes and other substances such as hormone
Hormone

Hormones are chemicals released by cells that affect cells in other parts of the body. Only a small amount of hormone is required to alter cell metabolism....
s that would be toxic if allowed to accumulate in the blood
Blood

Blood is a specialized bodily fluid that delivers necessary substances to the body's Cell s ? such as nutrients and oxygen ? and transports waste products away from those same cells....
 via organs such as the skin
Skin

The skin is the outer covering of the body, also known as the epidermis. It is the largest organ of the integumentary system made up of multiple layers of epithelial biological tissue, and guards the underlying muscles, bones, ligaments and organ s....
 and the kidneys; keeping the water and dissolved solutes in balance is referred to as osmoregulation.

Regulators and conformers


Two major types of osmoregulation are osmoconformers and osmoregulators. Osmoconformer
Osmoconformer

An osmoconformer is a marine invertebrate that maintains its internal salinity such that it is always equal to the surrounding seawater. Osmoconformers do not actively exchange solutes with the environment, but keep their body fluids tonicity to the external environment by actively regulating their internal concentration of amino acids, ions,...
s
match their body osmolarity to their environment . It can be either active or passive. Most marine invertebrates are osmoconformers, although their ionic composition may be different from that of seawater.

Osmoregulators tightly regulate their body osmolarity, which always stays constant and are more common in the animal kingdom. Osmoregulators actively control salt concentrations despite the salt concentrations in the environment. An example is freshwater fish. The gills actively uptake salt from the environment by the use of mitochondria-rich (MR) cells. Water will diffuse into the fish, so it excretes a very hypotonic
Tonicity

Tonicity measures the ability of a solution to exert an osmotic pressure upon the membrane. Osmolality and osmolarity measure concentration of the solutes independently on their ability to cross the membrane....
 urine to expel all the excess water. A marine fish
Fish

A fish is any marine biology vertebrate animal that is typically ectothermic , covered with scale , and equipped with two sets of paired fins and several unpaired fins....
 has an internal osmotic concentration lower than that of the surrounding seawater, so it tends to lose water and gain salt. It actively excretes salt
Salt

A salt, in chemistry, is defined as the product formed from the neutralisation reaction of acids and base . Salts are ionic compounds composed of cations and anions so that the product is electrically electric charge ....
 out from the gill
Gill

A gill is an anatomical structure found in many aquatic ecosystem organisms. It is a respiration organ whose function is the extraction of oxygen from water and the excretion of carbon dioxide....
s. Most fish are stenohaline
Stenohaline

Stenohaline describes an organism, usually fish, that cannot handle a wide fluctuation in the salt content of water. Stenohaline is derived from the words: "steno" meaning narrow, and "haline" meaning salt....
, which means they are restricted to either salt or fresh water and cannot survive in water with a different salt concentration than they are adapted to. However, some fish show a tremendous ability to effectively osmoregulate across a broad range of salinities; fish with this ability are known as euryhaline
Euryhaline

Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the Poecilia_sphenops which can live in Fresh_water, brackish, or Sea_water water....
 species.

Osmoregulation in plants


There are no specific osmoregulation organs in higher plants. Control of water intake and loss is by means of those internal and external factors that affect the rate of transpiration.

Plants share with animals the problems of obtaining water and in disposing of the surplus. Certain plants develop methods of water conservation. Xerophyte
Xerophyte

A xerophyte or xerophytic organism is a plant which is able to survive in an environment with little available water or moisture, usually in environments where potential evapotranspiration exceeds precipitation for all or part of the growing season....
s are plants in dry habitats, such as deserts, that are able to withstand prolonged periods of water shortage. Succulent plants such as the cactus
Cactus

A cactus is any member of the spine plant family Cactaceae, native to the Americas. They are often used as ornamental plants, but some are also Crop plants....
 have water stored in large parenchyma
Parenchyma

Parenchyma is a term used to describe a bulk of a substance. It is used in different ways in animals and in plants.The term is New Latin, from Greek language parenkhuma, visceral flesh, from parenkhein, to pour in beside : para-, beside + en-, in + khein, to pour....
 tissues. Other plants have leaf
Leaf

In botany, a leaf is an above-ground plant Organ specialized for photosynthesis. For this purpose, a leaf is typically flat and thin, to expose the cells containing chloroplast to light over a broad area, and to allow light to penetrate fully into the tissues....
 modifications to reduce water loss, such as needle-shaped leaves, sunken stomata, and thick, waxy cuticles as in the pine
Pine

Pines are Pinophyta trees in the genus Pinus, in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authorities accept between 105 and 125 species....
. The sand-dune marram grass has rolled leaves with stomata on the inner surface. Oncophyorans are also osmoregulators. Hdrophytesare plants in water habitates like water lily.

Osmoregulation in protists and animals


Amoeba
Amoeba

Amoeba is a term used either to describe protists that move by crawling via pseudopods, or to refer to a genus that includes species that move by this mechanism....
 make use of contractile vacuoles to collect excretory waste, such as ammonia
Ammonia

Ammonia is a chemical compound with the chemical formula nitrogenhydrogen. It is normally encountered as a gas with a characteristic pungent odor....
, from the intracellular fluid by both diffusion and active transport. As osmotic action pushes water from the environment into the cytoplasm, the vacuole moves to the surface and disposes the contents into the environment.

Kidneys play a very large role in human osmoregulation, regulating the amount of water in urine waste. With the help of hormones such as antidiuretic hormone, aldosterone
Aldosterone

Aldosterone is a hormone that increases the reabsorption of sodium and water and the release of potassium in the kidneys. This increases the volume of fluid in the body, and drives blood pressure up....
, and angiotensin II, the human body can increase the permeability of the collecting ducts in the kidney to reabsorb water and prevent it from being excreted.

A major way animals have evolved to osmoregulate is by controlling the amount of water excreted through the excretory system
Excretory system

This system involves the kidneys and the liver which is not part of the system but creates most of the excreted waistLeifangThe excretory system excretes wastes....
.

Vertebrate excretory systems


Waste products of nitrogen metabolism


Ammonia
Ammonia

Ammonia is a chemical compound with the chemical formula nitrogenhydrogen. It is normally encountered as a gas with a characteristic pungent odor....
 is a toxic by-product of protein
Protein

Proteins are organic compounds made of amino acids arranged in a linear chain and joined together by peptide bonds between the carboxyl and amino groups of adjacent amino acid Residue ....
 metabolism and is generally converted to less toxic substances after it is produced then excreted; mammal
Mammal

Mammals are a class of vertebrate animals whose name is derived from their distinctive feature, mammary glands, with which they feed their young....
s convert ammonia to urea, whereas bird
Bird

Birds are wing, Bipedalismal, endothermic , vertebrate animals that lay egg . There are around 10,000 living species, making them the most numerous tetrapod vertebrates....
s and reptile
Reptile

Reptiles, or members of the class Reptilia, are air-breathing, cold-blooded vertebrates that have skin covered in scale as opposed to hair or feathers....
s form uric acid to be excreted with other wastes via their cloaca
Cloaca

In zoological anatomy, a cloaca is the posterior opening that serves as the only such opening for the alimentary tract and urinary tract of certain animal species....
s.

Achieving osmoregulation in vertebrates


Four processes occur:

  • filtration - fluid portion of blood (plasma) is filtered from a nephron
    Nephron

    Nephron is the basic structural and functional unit of the kidney. Its chief function is to regulate the concentration of water and soluble substances like sodium salts by filtering the blood, reabsorbing what is needed and excreting the rest as urine....
     (functional unit of vertebrate kidney) structure known as the glomerulus
    Glomerulus

    A glomerulus is a capillary tuft surrounded by Bowman's capsule in nephrons of the vertebrate kidney. It receives its blood supply from an afferent arteriole of the renal circulation....
     into Bowman's capsule
    Bowman's capsule

    The Bowman's capsule is a cup-like sac at the beginning of the tubular component of a nephron in the mammalian kidney. A glomerulus is enclosed in the sac....
     or glomerular capsule (in the kidney's cortex) and flows down the proximal convoluted tubule to a "u-turn" called the Loop of Henle
    Loop of Henle

    In the kidney, the loop of Henle is the portion of the nephron that leads from the proximal convoluted tubule to the distal convoluted tubule. The loop has a Hairpin turn in the renal medulla....
     (loop of the nephron) in the medulla portion of the kidney.
  • reabsorption - most of the viscous glomerular filtrate is returned to blood vessels that surround the convoluted tubules.
  • secretion - the remaining fluid becomes urine
    Urine

    Urine is a liquid waste product of the body secreted by the kidneys by a process of filtration from blood called urination and excreted through the urethra....
    , which travels down collecting ducts to the medullary region of the kidney.
  • excretion - the urine (in mammals) is stored in the urinary bladder and exits via the urethra; in other vertebrates, the urine mixes with other wastes in the cloaca before leaving the body; ( frogs also have a urinary bladder).


See also

  • halotolerance
    Halotolerance

    Halotolerance is the adaptation of living organisms to conditions of high salinity. Halotolerant species tend to live in areas such as hypersaline lakes, dune, saline deserts, salt marshes, and inland salt seas and spring s....
  • salt gland
    Salt gland

    The salt gland is an organ for excreting excess salts. It is found in elasmobranchs, seabirds, and some reptiles. In sharks, salt glands are found in the rectum, but in birds and reptiles, they are found in the skull, in the area of the eyes, nostrils or mouth....


External links

  • , Prof. Chuck Holliday, Dept. of Biology, Lafayette College. Contains links to articles on osmoregulation in crustaceans.