Neotropical fishes
Encyclopedia
The freshwater
Freshwater
Fresh water is naturally occurring water on the Earth's surface in ice sheets, ice caps, glaciers, bogs, ponds, lakes, rivers and streams, and underground as groundwater in aquifers and underground streams. Fresh water is generally characterized by having low concentrations of dissolved salts and...

 fishes of tropical South and Central America represent one of the most diverse aquatic ecosystem
Ecosystem
An ecosystem is a biological environment consisting of all the organisms living in a particular area, as well as all the nonliving , physical components of the environment with which the organisms interact, such as air, soil, water and sunlight....

s on Earth, with more than 5,600 species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

, representing about 10% all living vertebrate
Vertebrate
Vertebrates are animals that are members of the subphylum Vertebrata . Vertebrates are the largest group of chordates, with currently about 58,000 species described. Vertebrates include the jawless fishes, bony fishes, sharks and rays, amphibians, reptiles, mammals, and birds...

 species. The exceptional diversity of species, adaptations, and life histories observed in the Neotropic
Neotropic
In biogeography, the Neotropic or Neotropical zone is one of the eight terrestrial ecozones. This ecozone includes South and Central America, the Mexican lowlands, the Caribbean islands, and southern Florida, because these regions share a large number of plant and animal groups.It is sometimes used...

al ichthyofauna has been the focus of numerous books and scientific papers, especially the wonderfully complex aquatic ecosystems of the Amazon Basin
Amazon Basin
The Amazon Basin is the part of South America drained by the Amazon River and its tributaries that drains an area of about , or roughly 40 percent of South America. The basin is located in the countries of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela...

 and adjacent river basins (e.g., Goulding and Smith, 1996; Araujo-Lima and Goulding, 1997; Barthem and Goulding, 1997; Barthem, 2003; Goulding et al., 2003). Many of the advances in Neotropical ichthyology
Ichthyology
Ichthyology is the branch of zoology devoted to the study of fish. This includes skeletal fish , cartilaginous fish , and jawless fish...

 have been summarized in three edited volumes; Malabarba et al. (1998);Reis et al. (2003); Albert and Reis (2011).
The Neotropical ichthyofauna extends throughout the continental waters of Central and South America, from south of the Mesa Central in southern Mexico (~ 16° N) to the La Plata estuary in northern Argentina (~ 34° S). The fishes of this region are largely restricted to the humid tropical portions of the Neotropical realm as circumscribed by Sclater (1858) and Wallace (1876), being excluded from the arid Pacific slopes of Peru and northern Chile, and the boreal regions of the Southern Cone in Chile and Argentina (Arratia, 1997; Dyer, 2000). The vast Neotropical ichthyofaunal region extends over more than 17 million square km of moist tropical lowland forests, seasonally flooded wetlands and savannahs, and also several arid peripheral regions (e.g., Northwest Venezuela; Northeast Brazil; Chaco of Paraguay, Argentina and Bolivia). At the core of this system lies Amazonia, the greatest interconnected freshwater fluvial system on the planet. This system includes the drainages of the Amazon Basin itself, and of two large adjacent regions, the Orinoco Basin and the Guiana Shield. The Amazon river
Amazon River
The Amazon of South America is the second longest river in the world and by far the largest by waterflow with an average discharge greater than the next seven largest rivers combined...

is by any measure the largest in the world, discharging about 16% of the world's flowing freshwater into the Atlantic (Goulding et al., 2003b).
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