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Mule



 
 
For other uses, see Mule (disambiguation)
Mule (disambiguation)

A mule is a hybrid equid, a cross of a female horse and a male donkey, often used for transport in rugged terrainMule or MULE can also refer to:...


In its common modern meaning, a mule is the offspring of a male donkey
Donkey

The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
 and a female horse
Horse

The horse is a hoofed mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolution of the horse over the past 45 to 55 million years from a small multi-toed creature into the large, odd-toed ungulate animal of today....
. (The much rarer offspring of a male horse and a female donkey is called a hinny
Hinny

A hinny is a domestication equine hybrid which is the offspring of a male horse and a female donkey . It is similar to the more common mule, but mules are the product of a female horse and a male donkey....
.) Mules are classified as an F1 hybrid
F1 hybrid

F1 hybrid is a term used in genetics and selective breeding. F1 stands for Filial 1, the first filial generation seeds/plants or animal offspring resulting from a cross mating of distinctly different parental types....
. The term "mule" (Latin
Latin

Latin is an Italic language, historically spoken in Latium and Ancient Rome. Through the Military history of the Roman Empire, Latin spread throughout the Mediterranean and a large part of Europe....
 mulus) was formerly applied to the infertile offspring of any two creatures of different species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
.

The size of a mule and work to which it is put depends largely on the breeding of the mule's dam
Mare

Mare most commonly refers to mare.Mare and similar may also refer to:...
.






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For other uses, see Mule (disambiguation)
Mule (disambiguation)

A mule is a hybrid equid, a cross of a female horse and a male donkey, often used for transport in rugged terrainMule or MULE can also refer to:...


In its common modern meaning, a mule is the offspring of a male donkey
Donkey

The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
 and a female horse
Horse

The horse is a hoofed mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolution of the horse over the past 45 to 55 million years from a small multi-toed creature into the large, odd-toed ungulate animal of today....
. (The much rarer offspring of a male horse and a female donkey is called a hinny
Hinny

A hinny is a domestication equine hybrid which is the offspring of a male horse and a female donkey . It is similar to the more common mule, but mules are the product of a female horse and a male donkey....
.) Mules are classified as an F1 hybrid
F1 hybrid

F1 hybrid is a term used in genetics and selective breeding. F1 stands for Filial 1, the first filial generation seeds/plants or animal offspring resulting from a cross mating of distinctly different parental types....
. The term "mule" (Latin
Latin

Latin is an Italic language, historically spoken in Latium and Ancient Rome. Through the Military history of the Roman Empire, Latin spread throughout the Mediterranean and a large part of Europe....
 mulus) was formerly applied to the infertile offspring of any two creatures of different species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
.

The size of a mule and work to which it is put depends largely on the breeding of the mule's dam
Mare

Mare most commonly refers to mare.Mare and similar may also refer to:...
. Mules can be lightweight, medium weight, or even, when produced from draught horse mare
Mare

Mare most commonly refers to mare.Mare and similar may also refer to:...
s, of moderate heavy weight.

The mule, easier to breed and usually larger in size than a hinny, has monopolized the attention of breeders . Reproductive success occurs more often when a donkey is the sire
Father

The father is defined as the male parent of an offspring. The adjective "paternal" refers to father, parallel to "maternal" for mother.According to the anthropologist Maurice Godelier, the parental role assumed by human males is a critical difference between human society and that of humans' closest biological relatives - chimpanzees and b...
 and the horse
Horse

The horse is a hoofed mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolution of the horse over the past 45 to 55 million years from a small multi-toed creature into the large, odd-toed ungulate animal of today....
 is the dam
Mother

A mother is a biological and/or Maternal bond female parent of an offspring. Because of the complexity and differences of the social, cultural, and religious definitions and roles, it is challenging to define a mother in a universally accepted definition....
. Sometimes people let a stallion
Stallion (horse)

A stallion is a male horse that has not been castration, or gelding.Stallions will follow the horse conformation and phenotype of their list of horse breeds, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" neck as well as a somewhat more muscular physique as compared to female...
 (male horse) run with a jenny (female donkey) for as long as six years before she becomes pregnant. Mules and hinnies are almost always sterile
Infertility

Infertility primarily refers to the biological inability of a person to contribute to fertilization. Infertility may also refer to the state of a woman who is unable to carry a pregnancy to full term....
 (see fertile mules
Mule

In its common modern meaning, a mule is the offspring of a male donkey and a female horse.Mules are classified as an F1 hybrid.The term "mule" was formerly applied to the infertile offspring of any two creatures of different species....
 below for rare cases). The sterility is attributed to the differing number of chromosome
Chromosome

A chromosome is an organized structure of DNA and protein that is found in Cell . A chromosome is a single piece of DNA that contains many genes, regulatory sequence and other genetic sequence....
s of the two species: donkeys have 62 chromosomes, whereas horses have 64.

A female mule that has estrus cycles and can carry a fetus is called a "molly" and can occasionally occur naturally as well as through embryo transfer
Embryo transfer

Embryo transfer refers to a step in the process of in vitro fertilisation whereby one or several embryos are placed into the uterus of the female with the intent to establish a pregnancy....
.

Biology


Size and Performance

The median weight range for a mule is between .

Although it depends on the individual animal, a Pakistani army mule can "carry up to 172 kg and walk 26 km without resting."

Characteristics

With its short thick head, long ears, thin limbs, small narrow hooves, short mane
Mane

Mane can have the following meanings:*The mane is used to describe the line of hair along the spine of the neck, starting behind the ears and ending just above the withers....
, absence of chestnuts (horny growths) inside the hocks, the mule looks like a donkey
Donkey

The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
; in height and body, shape of neck and croup, uniformity of coat, and teeth, it appears horse-like; the mule comes in all sizes, shapes and conformities. There are mules that resemble quarter horses
American Quarter Horse

The American Quarter Horse is an American Horse breeds of horse that excels at sprinting short distances. Its name came from its ability to outdistance other breeds of horses in races of a quarter mile or less, where some individuals have been clocked at speeds up to 55 mph....
, huge draft mules, fine-boned racing mules, shaggy pony
PONY

PONY may refer to:*PONY Baseball and Softball*PONY MagazineA PONY can refer to a small keg of beer....
 mules and many more types.

A mule does not sound exactly like a donkey or a horse. Instead, a mule makes a sound that is similar to a donkey's but also has the whinnying characteristics of a horse (often starts with a whinny, ends in a hee-haw). Sometimes, mules whimper. The coats of mules come in the same varieties as those of horses. Common colors are sorrel
Sorrel (horse)

Sorrel is one of the most common equine coat colors in horses. While it is usually used to refer to a copper-red shade of chestnut horse, in some places it is used generically in place of the term "chestnut" to describe any reddish horse with a same-color or lighter mane and tail, ranging from reddish-gold to a deep burgundy or chocolate s...
, bay, black
Black (horse)

Black is a equine coat color of horses in which the entire hair coat is black. Black is a relatively uncommon coat color, and novices frequently mistake dark Chestnut or Bay for black....
, and grey
Gray (horse)

Gray or grey is a Equine coat color of horses characterized by progressive silvering of the colored hairs of the coat. Most gray horses have black skin and dark eyes; unlike many depigmentation genes, gray does not affect skin or eye color Their adult hair coat is white, dappled, or white intermingled with hairs of other colors....
. Less common are white
White (horse)

True "white" horses, especially those that carry the proposed White or "W" gene, are rare. Most horses that are commonly referred to as "white" are actually Gray whose hair coats are completely white.ge:Cremello5.jpg|thumb|right|150px|This light palomino foal has dark skin and a creme-colored coat that can be mistaken for white....
, roans
Roan (color)

Roan is a coat color found in many animals, notably horses, cattle and dogs. It is defined generally as an even mixture of white and pigmented hairs that does not Gray as the animal ages....
 (both blue and red), palomino
Palomino

Palomino is a equine coat color in horses, consisting of a gold coat and white mane and tail. Genetically, the palomino color is created by a single allele of a dilution gene called the cream gene working on a red base coat....
, dun
Dun gene

File:Mesteno.jpgThe dun gene is a dilution gene that affects both red and black pigments in the equine coat color of a horse. The dun gene has the ability to affect the appearance of all black , bay , or chestnut -based horses to some degree by lightening the base body coat and suppressing the underlying base color to the mane, tail, legs...
, and buckskin. Least common are paint
Pinto horse

A Pinto horse has a equine coat color that consists of large patches of white and another color. In nations using British English, the term used to describe this pattern is #Colors....
 mules or tobiano
Tobiano

Tobiano is a spotted color pattern commonly seen in Pinto horses, produced by a dominant gene. The tobiano gene produces white-haired, pink-skinned patches on a base coat color....
s.

The mule possesses the sobriety, patience, endurance and sure-footedness of the donkey
Donkey

The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
, and the vigour, strength and courage of the horse. Operators of working animal
Working animal

A working animal is an animal that is kept by humans and trained to perform tasks. They may be close members of the family, such as guide dogs, or domestications such as logging elephants....
s generally find mules preferable to horses: mules show less impatience under the pressure of heavy weights, and their skin, harder and less sensitive than that of horses, renders them more capable of resisting sun and rain. Their hooves are harder than horses', and they show a natural resistance to disease and insects. Many North American farmer
Farmer

A farmer is a person who raises living organisms for food or raw materials....
s with clay
Clay

Clay is a naturally occurring material composed primarily of fine-grained minerals, which show plasticity through a variable range of water content, and which can be hardened when dried and/or fired....
 soil found mules superior as plow animals, especially in the U.S. state of Missouri
Missouri

Missouri is a U.S. state in the Midwestern United States of the United States bordered by Iowa, Illinois, Kentucky, Tennessee, Arkansas, Oklahoma, Kansas and Nebraska....
, hence the expression "stubborn as a Missouri mule".

Mules are generally less tolerant towards dogs than horses are. They are also capable of striking out with any of their hooves in any direction, even sideways if needed.

Mules exhibit a higher cognitive intelligence than their parent species - horses and donkeys. This is believed to be the result of hybrid vigour
Heterosis

Heterosis is a term used in genetics and selective breeding. The term heterosis, also known as hybrid vigour or outbreeding enhancement, describes the increased strength of different characteristics in Hybrid ; the possibility to obtain a genetically superior individual by combining the virtues of its parents....
, similar to how mules acquire greater height and endurance than either parents.

Mules are highly intelligent. They tend to be curious by nature. A mule generally will not let the rider put it in harm's way. However, the stereotype of the mule as being stubborn is somewhat unfair and inaccurate.

Distribution

FAO reports that China was the top market for mules in 2003 closely followed by Mexico and many Central and South American nations.

Fertility


Several female mules have produced offspring when mated with a purebred horse or donkey. Since 1527 there have been more than 60 documented cases of foals born to female mules around the world. There are no recorded cases of fertile mule stallions. Mules and Hinnies have 63 chromosomes that are a mixture of one from each parent. The different structure and number usually prevents the chromosomes from pairing up properly and creating successful embryos.

There are reports that a mule in China produced a foal in 1984.

In Morocco
Morocco

Morocco , officially the Kingdom of Morocco , is a country located in North Africa with a population of nearly 34 million and an area just under 447,000 km2....
, in early 2002, a mare mule produced a rare foal.

In 2007 a mule named Kate gave birth to a mule son in Colorado. Blood and hair samples were tested verifying that the mother was a mule and the colt was indeed her offspring.

The modern mule

After World War II
World War II

World War II, or the Second World War , was a global military conflict which involved a Participants in World War II, including all of the great powers, organised into two opposing military alliances: the Allies of World War II and the Axis powers....
, mules fell on hard times. The use of mules for farming and transportation of agricultural products gave way to modern tractors and trucks. A dedicated number of mule breeders, however, continued the tradition as hobby and continued breeding the great lines of mammoth jacks started in the United States
United States

The United States of America is a Federal government constitutional republic comprising U.S. state and a federal district. The country is situated mostly in central North America, where its Contiguous United States and Washington, D.C., the Capital districts and territories, lie between the Pacific Ocean and Atlantic Oceans, Borders of the U...
 by George Washington
George Washington

George Washington was the leader of the Continental Army in the American Revolutionary War and served as the List of Presidents of the United States President of the United States of the United States of Americas ....
 with the gift from the King of Spain of two Catalan
Catalonia

Catalonia , is an Autonomous Community in northeast Spain.Catalonia covers an area of 32,114 km? and has an official population of 7,210,508. It borders France and Andorra to the north, Aragon to the west, the Valencian Community to the south, and the Mediterranean Sea to the east ....
 jacks. These hobby breeders began to utilize better mares for mule production until today's modern saddle mule emerged. Exhibition shows where mules pulled heavy loads have now been joined with mules competing in Western
Western riding

Western riding is a style of Equestrianism which evolved from the ranching and warfare traditions brought to the Americas by the Spain Conquistadors, and both equipment and riding style evolved to meet the working needs of the cowboy in the American West....
 and English pleasure
English Pleasure

English pleasure is generic term for a number of different English riding classes seen at horse shows in the United States, where the horse is ridden in either hunt seat or saddle seat tack....
 riding, as well as dressage
Dressage

Dressage is a path and destination of competitive horse training, with competitions held at all levels from amateur to the Equestrian at the Summer Olympics....
 and show jumping
Show jumping

Show jumping, also known as "stadium jumping" or "jumpers," is a member of a family of English riding equestrianism events that also includes dressage, eventing, Show hunter and equitation....
 competition. There is now a cable TV show dedicated to the training of donkeys and mules. Mules, once snubbed at traditional horse shows, have been accepted for competition at the most exclusive horse shows in the world in all disciplines.

Amish
Amish

The various Amish or Amish Mennonite church fellowships are Christian religious denominations, and form a very traditional subgrouping of Mennonite churches....
 farmers, who reject tractors and most other modern technology, commonly use teams of six or eight mules to pull plows, diskers, and other farm equipment, though they use horses for pulling buggies on the road.

During the Soviet war in Afghanistan
Soviet war in Afghanistan

The Soviet war in Afghanistan was a nine-year war involving Soviet Union Military of the Soviet Union supporting the Marxism People's Democratic Party of Afghanistan government against the Mujahideen#Afghanistan resistance movement....
, the United States used large numbers of mules to carry weapons and supplies over Afghanistan's rugged terrain to the mujahideen.

The mule clone


In 2003, researchers at University of Idaho
University of Idaho

The University of Idaho is Idaho's flagship and oldest public university, located in the rural city of Moscow, Idaho in Latah County, Idaho. It is the state's land-grant university and primary research university....
 and Utah State University
Utah State University

Utah State University is a Public university land-grant university whose main campus is located in Logan, Utah.It was established in 1888, after Anthon H....
 found a way to reproduce mules -- by cloning the first mule as part of . The research team includes Gordon Woods, UI professor of animal
Animal science

Animal science is described as "studying the biology of animals that are under the control of mankind" . Historically the animals studied were farm animals but courses available now look at a far broader area to include companion animals for example dogs, cats, horses and captive animals....
 and veterinary science
Veterinary medicine

Veterinary medicine is that branch of medical science,which deals with the study of diagnosis,treatment and prevention of diseases in companion,domestic, exotic, wildlife and production animals....
, Kenneth L. White, USU professor of animal science, and Dirk Vanderwall, UI assistant professor of animal and veterinary science. The baby mule, Idaho Gem, was born May 4. It is the first clone of a hybrid animal. Veterinary examinations of the foal and its surrogate mother showed them to be in good health soon after birth. The foal's DNA comes from a fetal cell culture first established in 1998 at the University of Idaho.

Color and size variety


Mules today come in a variety of shapes, sizes and colors, from minis under 50 pounds to maxis over 1000 pounds, and in many different colors. Mules from Appaloosa
Appaloosa

The Appaloosa is a list of horse breeds known for its preferred leopard complex-spotted coat pattern and other distinctive physical characteristics....
 mares produce wildly colored mules, much like their Appaloosa horse relatives, but with even wilder skewed colors. The Appaloosa color is produced by a complex of genes known as the Leopard complex
Leopard complex

The leopard complex is a group of genetically-related Equine coat color in horses. These patterns range from progressive increases in interspersed white hair similar to Gray or Roan to distinctive, Dalmatian -like leopard spots on a white coat....
 (Lp). Mares homozygous for the Lp gene bred to any color donkey
Donkey

The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
 will produce an Appaloosa colored mule.

See also

  • African Wild Ass
    African Wild Ass

    The African Wild Ass is a wild member of the horse family, Equidae. This species is believed to be the ancestor of the domestic donkey which is usually placed within the same species....
  • Donkey
    Donkey

    The 'donkey' or 'ass', Equus africanus asinus, is a Domestication member of the Equidae or horse family, and an Odd-toed ungulates. The wild ancestor of the donkey is the Wild Ass, E....
  • Horse
    Horse

    The horse is a hoofed mammal, a subspecies of one of seven extant species of the family Equidae. The horse has evolution of the horse over the past 45 to 55 million years from a small multi-toed creature into the large, odd-toed ungulate animal of today....
  • Jennet
    Jennet

    A Jennet or Spanish Jennet was a small Spain horse. It was noted for a smooth naturally ambling horse gait, compact and well-muscled build, and a good disposition....
  • Kiang
    Kiang

    The kiang is the largest of the wild asses. This equine is native to the Tibetan Plateau, where it inhabits montane grasslands and shrublands from 4000 to 7000 meters elevation....
  • Hinny
    Hinny

    A hinny is a domestication equine hybrid which is the offspring of a male horse and a female donkey . It is similar to the more common mule, but mules are the product of a female horse and a male donkey....
  • Onager
    Onager

    The Onager is a large mammal belonging to the genus Equus of the family Equidae and native to the deserts of Syria, Iran, Pakistan, India, Israel, and Tibet....
  • 40 acres and a mule
    40 acres and a mule

    40 acres and a mule is a term for compensation that was promised to be awarded to freed African American slaves after the American Civil War? 40 acres of land to farm, and a mule with which to drag a plow so the land could be cultivated....


External links

  • , registers mules of all types/educational information. Also some fertile mule information in FAQ.
  • hosted by the