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Housefly



 
 
The housefly (also house fly, house-fly or common housefly), Musca domestica, is the most common of all flies fluttering in homes, and indeed one of the most widely distributed insects; it is often considered a pest
Pest (animal)

A pest is an organism which has characteristics that are regarded by humans as injurious or unwanted. This is most often because it causes damage to agriculture through feeding on crops or parasitising livestock, such as codling moth on apples, or boll weevil on cotton....
 that can carry serious diseases.

adults are 6–9 mm long. Their thorax
Thorax

The thorax is a division of an animal's body that lies between the head and the abdomen.In mammals, the thorax is the region of the body formed by the sternum, the thoracic vertebrae and the ribs....
 is gray, with four longitudinal dark lines on the back.






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The housefly (also house fly, house-fly or common housefly), Musca domestica, is the most common of all flies fluttering in homes, and indeed one of the most widely distributed insects; it is often considered a pest
Pest (animal)

A pest is an organism which has characteristics that are regarded by humans as injurious or unwanted. This is most often because it causes damage to agriculture through feeding on crops or parasitising livestock, such as codling moth on apples, or boll weevil on cotton....
 that can carry serious diseases.

Physical description

The adults are 6–9 mm long. Their thorax
Thorax

The thorax is a division of an animal's body that lies between the head and the abdomen.In mammals, the thorax is the region of the body formed by the sternum, the thoracic vertebrae and the ribs....
 is gray, with four longitudinal dark lines on the back. The underside of their abdomen is yellow, and their whole body is covered with hair. The females are slightly larger than the males, and have a much larger space between their red compound eyes.

Like most Diptera
Fly

True flies are insects of the Order Diptera , possessing a single pair of insect wing on the mesothorax and a pair of halteres, derived from the hind wings, on the metathorax....
 (meaning "two-winged"), houseflies have only one pair of wings
Insect wing

Insect wings are outgrowths of the insect exoskeleton that enable insects to Insect flight. They are found on the second and third thorax segments , and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments....
; the hind pair is reduced to small halteres
Halteres

Halteres are small knobbed structures found as a pair in some two-winged insects. They are flapped rapidly and function as accelerometers to help the insect maintain stability in flight, analogous to an aircraft's attitude indicator....
 that aid in flight
Flight

Flight is the process by which an object moves either through the air, or movement beyond earth's atmosphere , by aerodynamically generating Lift , propulsion or Lighter than air using buoyancy, or by simple ballistic movement....
 stability. Characteristically, the media vein (M1+2 or fourth long vein
Insect wing

Insect wings are outgrowths of the insect exoskeleton that enable insects to Insect flight. They are found on the second and third thorax segments , and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments....
 of the wing) shows a sharp upward bend.

Species that appear similar to the housefly include:
  • The lesser house fly
    Lesser house fly

    The lesser house fly or little house fly, Fannia canicularis, is somewhat smaller than the common housefly. It is slender, and the median vein in the wing is straight....
    , Fannia canicularis, is somewhat smaller, more slender, and the media vein is straight.
  • The stable fly
    Stable fly

    Stomoxys calcitrans is commonly called the "stable fly", "horse fly", or "dog fly". Rather unusual for a member of the family Muscidae is that it Hematophagy from mammals....
    , Stomoxys calcitrans, has piercing mouthparts and the media vein is only slightly curved.


Life cycle

Each female fly can lay approximately 500 eggs in several batches of about 75 to 150, eggs
Egg (biology)

In most birds and reptiles, an egg is the zygote, resulting from fertilization of the ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing embryo....
 . The eggs are white and are about 1.2 mm in length. Within a day, larvae (maggot
Maggot

Maggot is the common name of the larval phase of development in insects of the order Diptera . Sometimes the word is used to denote the larval stage of any insects....
s) hatch from the eggs; they live and feed in (usually dead and decaying) organic material, such as garbage or faeces. They are pale-whitish, 3-9 mm long, thinner at the mouth end, and have no legs. They live at least one week. At the end of their third instar
Instar

An instar is a developmental stage of arthropods, such as insects, between each ecdysis , until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form....
, the maggots crawl to a dry cool place and transform into pupa
Pupa

A pupa is the life stage of some insects undergoing transformation. The pupal stage is found only in Holometabolism insects, those that undergo a complete metamorphosis, going through four life stages; embryo, larva, pupa and imago....
e, colored reddish or brown and about 8 mm long. The adult flies then emerge from the pupae. (This whole cycle is known as complete metamorphosis.) The adults live from two weeks to a month in the wild, or longer in benign laboratory conditions. After having emerged from the pupae, the flies cease to grow; small flies are not young flies, but are indeed the result of getting insufficient food during the larval stage.

Some 36 hours after having emerged from the pupa, the female is receptive for mating. The male mounts her from behind to inject sperm
Sperm

The term sperm is derived from the Greek word sperma and refers to the male reproductive Cell . In the types of sexual reproduction known as anisogamy and oogamy, there is a marked difference in the size of the gametes with the smaller one being termed the "male" or sperm cell....
. Normally the female mates only once, storing the sperm to use it repeatedly for laying several sets of eggs. Males are territorial: they will defend a certain territory against other males and will attempt to mount any females that enter that territory.

Housefly Pupae Killed By Wasp Larvae
The flies depend on warm temperatures; generally, the warmer the temperature the faster the flies will develop. In winter, most of them survive in the larval or the pupa stage in some protected warm location.

The average lifespan of an adult housefly is approximately 15 to 25 days. Since they can complete their lifecycle in as little as seven to ten days, flies typically live between 22 and 25 days from egg to death. 

Behavior


Houseflies can take in only liquid foods. They spit out saliva
Saliva

Saliva is the watery and usually frothy substance produced in the mouths of humans and most other animals. Saliva is produced in and secreted from the salivary glands....
 on solid foods to predigest it, and then suck it back inside. They also regurgitate partly digested matter and pass it again to the abdomen.

The flies can walk on vertical planes, and can even hang upside-down from ceilings. This is accomplished with the surface tension
Surface tension

Surface tension is an attractive property of the surface of a liquid. It is what causes the surface portion of liquid to be attracted to another surface, such as that of another portion of liquid ....
 of liquids secreted by gland
Gland

A gland is an Organ in an animal's body that synthesizes a substance for release such as hormones or breast milk, often into the bloodstream or into cavities inside the body or its outer surface ....
s near their feet. When they are not flying, flies continually preen themselves, cleaning their eyes with their forelegs and dusting off their legs by rubbing them together. They do this because most of their taste and smell receptors lie on the hair of their legs.

Flies have a very highly-evolved evasion reaction which helps to ensure their survival. It is possible to confuse a fly's evasion system by swatting it with two objects simultaneously from different directions. The holes in a fly swatter minimize the air current that warns the fly as being hit, whilst reducing air resistance and increasing speed of the swat.

Research in 2008 by Professor Michael Dickinson of the California Institute of Technology in Pasadena discovered that a fly anticipates the approach of a threat, calculating the angle of attack, and alters its stance ready to make the most appropriate getaway. Professor Dickinson showed that this evasive action, which allows the fly to jump away in the opposite direction, takes place within 200 ms.

Houseflies release a pheromone
Pheromone

A pheromone is a chemical that triggers a natural behavioral response in another member of the opposite gender of the same species. There are alarm signal pheromones, food trail pheromones, sex pheromones, and many others that affect behavior or physiology....
 called that serves both as aggregation
Swarm

The term swarm is applied to fish, insects, birds and microorganisms, such as bacteria, and describes a behavior of an aggregation of animals of similar size and body orientation, generally cruising in the same direction....
 and sexual attraction
Sexual attraction

Sexual attraction refers to a person's ability to Attractiveness in a sexual or erotic manner the interest of another person.Which aspects of sexual attraction have had greatest influence to humans at different points in time have differed between cultures and regions....
 purposes.

The housefly is an object of biological research, mainly because of one remarkable quality: the sex determination
Sex-determination system

A sex-determination system is a biology system that determines the development of sex in an organism. Most sexual organisms have two sexes. In many cases, sex determination is genetic: males and females have different alleles or even different genes that specify their sexual Comparative anatomy....
 mechanism. Although a wide variety of sex determination mechanisms exist in nature (e.g. male and female heterogamy
Heterogamy

Heterogamy has a number of biological definitions:*In reproductive biology, heterogamy is the alternation of differently organized generations, applied to the alternation between parthenogenesis and a sexual reproduction generation....
, haplodiploidy, environmental factors) the way sex is determined is usually fixed within one species
Species

In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring....
. However, the housefly exhibits many different mechanisms for sex determination, such as male heterogamy (like most insect
Insect

Insects are the biggest class of arthropods and the only ones with wings. They are the most diverse group of animals on the planet. They are most diverse at the equator and their diversity declines toward the poles....
s and mammal
Mammal

Mammals are a class of vertebrate animals whose name is derived from their distinctive feature, mammary glands, with which they feed their young....
s), female heterogamy (like birds) and maternal control over offspring sex. This makes the housefly one of the most suitable species to study the evolution
Evolution

In biology, evolution is change in the heritability trait of a population of organisms from one generation to the next. These changes are caused by a combination of three main processes: variation, reproduction, and selection....
 of sex determination.

Evolution

Even though the order of flies (Diptera) is much older, true houseflies evolved in the beginning of the Cenozoic
Cenozoic

The Cenozoic Era...
 era, some 65 million years ago. They are thought to have originated in the southern Palearctic
Palearctic

The Palearctic or Palaearctic is one of the eight ecozones dividing the Earth surface.Physically, the Palearctic is the largest ecozone....
 region, particularly the Middle East. Because of their close, commensal relationship with man, they probably owe their worldwide dispersal to co-migration with humans.

Flies and humans

In colder climates, houseflies survive only with humans. They have a tendency to aggregate and are difficult to dispel. They are capable of carrying over 100 pathogen
Pathogen

A pathogen , infectious agent, or germ, is a biological agent that causes disease or illness to its Host .There are several substrates and pathways whereby pathogens can invade a host; the principal pathways have different episodic time frames, but soil contamination has the longest or most persistent potential for harboring...
s, such as typhoid, cholera
Cholera

Cholera, sometimes known as Asiatic or epidemic cholera, is an infectious gastroenteritis caused by enterotoxin-producing strains of the bacterium Vibrio cholerae....
, Salmonella
Salmonella

Salmonella is a genus of rod-shaped Gram-negative enterobacteriaceae that causes typhoid fever, paratyphoid fever, and the foodborne illness salmonellosis....
, bacillary dysentery
Bacillary dysentery

Bacillary dysentery is a type of dysentery caused by shigellosis.Bacillary dysentery is associated with species of bacteria from the Enterobacteriaceae family....
, tuberculosis
Tuberculosis

Tuberculosis is a common and often deadly infectious disease caused by mycobacterium, mainly Mycobacterium tuberculosis . Tuberculosis usually attacks the lungs but can also affect the central nervous system, the lymphatic system, the circulatory system, the genitourinary system, the gastrointestinal system, bones, joints, and even the...
, anthrax
Anthrax

Anthrax is an Acute disease in humans and animals caused by the bacterium Bacillus anthracis, which is highly lethal in some forms. There are effective vaccines against anthrax, and some forms of the disease respond well to antibiotic treatment....
, ophthalmia
Ophthalmia

Ophthalmia is inflammation of the eye. It is a medical sign which may be indicative of various conditions, including sympathetic ophthalmia , ophthalmia neonatorum , and actinic conjunctivitis ....
, and parasitic worm
Parasitic worm

See also Parasitic worm Parasitic worms or helminths are a division of eukaroytic parasites that, unlike external parasites such as lice and fleas, live inside their host....
s. The flies in poorer and lower-hygienic areas usually carry more pathogens. Some strains have become immune to most common insecticide
Insecticide

An insecticide is a pesticide used against insects in all developmental forms. They include ovicides and larvicides used against the Egg and larvae of insects respectively....
s.

House flies feed on liquid or semi-liquid substances beside solid material which has been softened by saliva or vomit. Because of their high intake of food, they deposit faeces constantly, one of the factors that makes the insect a very dangerous and heinous carrier of pathogens. Although they are domestic flies, usually confined to the human habitations, they can fly for several miles from the breeding place. They are active only in daytime and rest at night e.g. at the corners of rooms, ceiling hangings, etc.

Housefly as a vector of disease

Mechanical transmission of organisms on its hairs, mouthparts, vomitus and feces:
  • parasitic diseases: Cysts of protozoa
    Protozoa

    Protozoan are microorganisms classified as unicellular eukaryotes. While there is no exact definition of the term "protozoan", most scientists use the word to refer to a unicellular heterotrophic protist, such as an amoeba or a ciliate....
     e.g. Entamoeba histolytica
    Entamoeba histolytica

    For the infection and disease caused by this parasite, refer to Amoebiasis.Entamoeba histolytica is an anaerobic organism parasitic protozoan, part of the genus Entamoeba....
    , Giardia lamblia
    Giardia lamblia

    File:Giardia life cycle en.svgGiardia lamblia is a flagellated protozoan parasite that colonises and reproduces in the small intestine, causing giardiasis....
     and eggs of helminths e.g.:Ascaris lumbricoides
    Ascaris lumbricoides

    Ascaris lumbricoides is the member of the Ascaris family responsible for the disease Ascariasis.It can reach a length of up to 35 cm....
    , Trichuros trichura, Haemenolypes nana, Enterobius vermicularis.
  • bacterial diseases: Typhoid, cholera, dysentery, pyogenic cocci...etc.
  • Viruses: Enterovirus
    Enterovirus

    The enteroviruses are a genus of Virus classification associated with several human and mammalian diseases. Historically the most significant has been the Poliovirus....
    es: Poliomyelitis
    Poliomyelitis

    Poliomyelitis, often called polio or infantile paralysis, is an acute virus infectious disease spread from person to person, primarily via the fecal-oral route....
    , infective hepatitis (A & E)..etc


As a causative agent of diseases

Accidental myiasis
Myiasis

Myiasis is an animal or human disease caused by Parasite Diptera fly larvae feeding on the host 's necrotic or living Biological tissue. Colloquialisms for Myiasis include fly-strike and fly-blown....


External links

  • on the UF
    University of Florida

    The University of Florida is a Public university land-grant university, sea grant colleges, Space grant colleges major research university located on a campus in Gainesville, Florida, in the United States....
     / IFAS
    Institute of Food and Agricultural Sciences

    The University of Florida?s Institute of Food and Agricultural Sciences is a federal-state-county partnership dedicated to developing knowledge in agriculture, human and natural resources, and the life sciences, and enhancing and sustaining the quality of human life by making that information accessible....
     Featured Creatures Web site
  • , by Anthony DeBartolo, Chicago Tribune
  • From The Straight Dope website:
  • , from Project Gutenberg
    Project Gutenberg

    Project Gutenberg, abbreviated as PG, is a volunteer effort to digitize, archive and distribute cultural works, as founder Michael Hart said "To encourage the creation and distribution of eBooks."....