Hexacorallia
Encyclopedia
Hexacorallia is a subclass of Anthozoa
Anthozoa
Anthozoa is a class within the phylum Cnidaria that contains the sea anemones and corals. Unlike other cnidarians, anthozoans do not have a medusa stage in their development. Instead, they release sperm and eggs that form a planula, which attaches to some substrate on which the cnidarian grows...

 comprising approximately 4,300 species of water-based organisms formed of colonial polyp
Polyp
A polyp in zoology is one of two forms found in the phylum Cnidaria, the other being the medusa. Polyps are approximately cylindrical in shape and elongated at the axis of the body...

s generally with 6-fold symmetry. This includes all of the stony coral
Coral
Corals are marine animals in class Anthozoa of phylum Cnidaria typically living in compact colonies of many identical individual "polyps". The group includes the important reef builders that inhabit tropical oceans and secrete calcium carbonate to form a hard skeleton.A coral "head" is a colony of...

s, which are vital for coral reef
Coral reef
Coral reefs are underwater structures made from calcium carbonate secreted by corals. Coral reefs are colonies of tiny living animals found in marine waters that contain few nutrients. Most coral reefs are built from stony corals, which in turn consist of polyps that cluster in groups. The polyps...

 formation, as well as all sea anemone
Sea anemone
Sea anemones are a group of water-dwelling, predatory animals of the order Actiniaria; they are named after the anemone, a terrestrial flower. Sea anemones are classified in the phylum Cnidaria, class Anthozoa, subclass Zoantharia. Anthozoa often have large polyps that allow for digestion of larger...

s, tube anemones, and zoanthids within six extant order
Order (biology)
In scientific classification used in biology, the order is# a taxonomic rank used in the classification of organisms. Other well-known ranks are life, domain, kingdom, phylum, class, family, genus, and species, with order fitting in between class and family...

s. They are distinguished from the other subclass of Anthozoa, Octocorallia, in having six or fewer lines of symmetry in their body structure and only single rows of tentacles. These organisms are formed of individual soft polyps which in some live in colonies and can secrete a calcite
Calcite
Calcite is a carbonate mineral and the most stable polymorph of calcium carbonate . The other polymorphs are the minerals aragonite and vaterite. Aragonite will change to calcite at 380-470°C, and vaterite is even less stable.-Properties:...

 skeleton. As with all Cnidaria
Cnidaria
Cnidaria is a phylum containing over 9,000 species of animals found exclusively in aquatic and mostly marine environments. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey. Their bodies consist of mesoglea, a non-living jelly-like substance,...

ns these organisms have a complex life cycle including a motile phase when they are considered plankton and later characteristic sessile
Sessility (zoology)
In zoology, sessility is a characteristic of animals which are not able to move about. They are usually permanently attached to a solid substrate of some kind, such as a part of a plant or dead tree trunk, a rock, or the hull of a ship in the case of barnacles. Corals lay down their own...

 phase. It also includes the significant extinct orders of the rugose corals and tabulate coral
Tabulate coral
The tabulate corals, forming the order Tabulata, are an extinct form of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to a honeycomb. Adjacent cells are joined by small pores...

s

Phylogeny

Hexacorallia is considered to be monophyletic, that is all contained species are descended from a common ancestor, however it has been suggested that many of the current orders are not. Historically Ceriantharia and Antipatharia were considered to be a separate subclass called Ceriantipatharia, though more recent genetic studies place them in Hexacorallia with Ceriantharia as the oldest, or basal, group.

The subclass includes important coral reef builders the stony corals (Scleractinia
Scleractinia
Scleractinia, also called stony corals, are exclusively marine animals; they are very similar to sea anemones but generate a hard skeleton. They first appeared in the Middle Triassic and replaced tabulate and rugose corals that went extinct at the end of the Permian...

), sea anemones (Actiniaria) and related tube-dwelling anemones (Ceriantharia), and zoanthids (Zoantharia
Zoantharia
Zoanthids are an order of cnidarians commonly found in coral reefs, the deep sea and many other marine environments around the world. These animals come in a variety of different colonizing formations and in numerous colors...

). Antipatharia contains the black corals and Corallimorpharia
Corallimorpharia
Corallimorpharia is an order of marine cnidarians closely related to the true sea anemones . They are mostly tropical, with a narrow column topped with a wide oral disc. The tentacles are usually short or very short, arranged in rows radiating from the mouth. Many species occur together in large...

 are similar to anemones. The extant orders are shown below:
A number of extinct orders of corals have been classified as their calcium skeleton forms a prolific fossil record. These are generally thought to be close to the ancestors of modern Scleractinians and existed during the Paleozoic
Paleozoic
The Paleozoic era is the earliest of three geologic eras of the Phanerozoic eon, spanning from roughly...

 period 570-245 million years ago:
  • Numidiaphyllida †
  • Kilbuchophyllida †
  • Heterocorallia †
  • Rugosa
    Rugosa
    Disambiguation:The Rugosa Rose is also sometimes just called "Rugosa". For the moon in Star Wars: The Clone Wars, see .The Rugosa, also called the Tetracoralla, are an extinct order of coral that were abundant in Middle Ordovician to Late Permian seas.Solitary rugosans are often referred to...

     †
  • Heliolitida †
  • Tabulata †
  • Cothoniida †
  • Tabuloconida †

Hexacorallia or Zoantharia

The terms Hexacorallia and Zoantharia have been used synonymously to avoid confusion with some of the extinct species not showing sixfold symmetry. For example, Rugosa
Rugosa
Disambiguation:The Rugosa Rose is also sometimes just called "Rugosa". For the moon in Star Wars: The Clone Wars, see .The Rugosa, also called the Tetracoralla, are an extinct order of coral that were abundant in Middle Ordovician to Late Permian seas.Solitary rugosans are often referred to...

 † is also called Tetracorallia, for its fourfold symmetry. Therefore Hexacorallia might appear misleading, and Zoantharia is sometime preferred in paleontological contexts. However, Zoantharia being the name used since 1899 until recent days for the hexacorallian order Zoantharia
Zoantharia
Zoanthids are an order of cnidarians commonly found in coral reefs, the deep sea and many other marine environments around the world. These animals come in a variety of different colonizing formations and in numerous colors...

the use of the term Hexacorallia is strongly recommended in order to avoid confusions.
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