Hadrosaurinae
Encyclopedia
Saurolophinae is a subfamily of hadrosaurid
Hadrosaurid
Hadrosaurids or duck-billed dinosaurs are members of the family Hadrosauridae, and include ornithopods such as Edmontosaurus and Parasaurolophus. They were common herbivores in the Upper Cretaceous Period of what are now Asia, Europe and North America. They are descendants of the Upper...

 dinosaur
Dinosaur
Dinosaurs are a diverse group of animals of the clade and superorder Dinosauria. They were the dominant terrestrial vertebrates for over 160 million years, from the late Triassic period until the end of the Cretaceous , when the Cretaceous–Paleogene extinction event led to the extinction of...

s. Members of the subfamily derive from the Late Cretaceous
Late Cretaceous
The Late Cretaceous is the younger of two epochs into which the Cretaceous period is divided in the geologic timescale. Rock strata from this epoch form the Upper Cretaceous series...

 of Antarctica, Argentina
Argentina
Argentina , officially the Argentine Republic , is the second largest country in South America by land area, after Brazil. It is constituted as a federation of 23 provinces and an autonomous city, Buenos Aires...

, Canada
Canada
Canada is a North American country consisting of ten provinces and three territories. Located in the northern part of the continent, it extends from the Atlantic Ocean in the east to the Pacific Ocean in the west, and northward into the Arctic Ocean...

 (Alberta
Alberta
Alberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...

 and Saskatchewan
Saskatchewan
Saskatchewan is a prairie province in Canada, which has an area of . Saskatchewan is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, and on the south by the U.S. states of Montana and North Dakota....

), China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

, Kazakhstan
Kazakhstan
Kazakhstan , officially the Republic of Kazakhstan, is a transcontinental country in Central Asia and Eastern Europe. Ranked as the ninth largest country in the world, it is also the world's largest landlocked country; its territory of is greater than Western Europe...

, Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

, Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

, the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 (Alabama
Alabama
Alabama is a state located in the southeastern region of the United States. It is bordered by Tennessee to the north, Georgia to the east, Florida and the Gulf of Mexico to the south, and Mississippi to the west. Alabama ranks 30th in total land area and ranks second in the size of its inland...

, Colorado
Colorado
Colorado is a U.S. state that encompasses much of the Rocky Mountains as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains...

, Georgia, Mississippi
Mississippi
Mississippi is a U.S. state located in the Southern United States. Jackson is the state capital and largest city. The name of the state derives from the Mississippi River, which flows along its western boundary, whose name comes from the Ojibwe word misi-ziibi...

, Montana
Montana
Montana is a state in the Western United States. The western third of Montana contains numerous mountain ranges. Smaller, "island ranges" are found in the central third of the state, for a total of 77 named ranges of the Rocky Mountains. This geographical fact is reflected in the state's name,...

, New Jersey
New Jersey
New Jersey is a state in the Northeastern and Middle Atlantic regions of the United States. , its population was 8,791,894. It is bordered on the north and east by the state of New York, on the southeast and south by the Atlantic Ocean, on the west by Pennsylvania and on the southwest by Delaware...

, New Mexico
New Mexico
New Mexico is a state located in the southwest and western regions of the United States. New Mexico is also usually considered one of the Mountain States. With a population density of 16 per square mile, New Mexico is the sixth-most sparsely inhabited U.S...

, North Carolina
North Carolina
North Carolina is a state located in the southeastern United States. The state borders South Carolina and Georgia to the south, Tennessee to the west and Virginia to the north. North Carolina contains 100 counties. Its capital is Raleigh, and its largest city is Charlotte...

, North Dakota
North Dakota
North Dakota is a state located in the Midwestern region of the United States of America, along the Canadian border. The state is bordered by Canada to the north, Minnesota to the east, South Dakota to the south and Montana to the west. North Dakota is the 19th-largest state by area in the U.S....

, South Dakota
South Dakota
South Dakota is a state located in the Midwestern region of the United States. It is named after the Lakota and Dakota Sioux American Indian tribes. Once a part of Dakota Territory, South Dakota became a state on November 2, 1889. The state has an area of and an estimated population of just over...

, and Wyoming
Wyoming
Wyoming is a state in the mountain region of the Western United States. The western two thirds of the state is covered mostly with the mountain ranges and rangelands in the foothills of the Eastern Rocky Mountains, while the eastern third of the state is high elevation prairie known as the High...

) and Uzbekistan
Uzbekistan
Uzbekistan , officially the Republic of Uzbekistan is a doubly landlocked country in Central Asia and one of the six independent Turkic states. It shares borders with Kazakhstan to the west and to the north, Kyrgyzstan and Tajikistan to the east, and Afghanistan and Turkmenistan to the south....

.

Classification

Saurolophinae was first defined as a clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

 in a 2010 phylogenetic analysis by Prieto-Márquez. Traditionally, the "crestless" branch of the family Hadrosauridae had been named Hadrosaurinae
Hadrosaurinae
Saurolophinae is a subfamily of hadrosaurid dinosaurs. Members of the subfamily derive from the Late Cretaceous of Antarctica, Argentina, Canada , China, Kazakhstan, Mexico, Russia, the United States and...

. However, the use of the term Hadrosaurinae was questioned in a comprehensive study of hadrosaurid relationships by Albert Prieto-Márquez in 2010. Prieto-Márquez noted that, though the name Hadrosaurinae had been used for the clade of mostly crestless hadrosaurids by nearly all previous studies, its type species, Hadrosaurus foulkii, has almost always been excluded from the clade that bears its name, in violation of the rules for naming animals set out by the ICZN
International Code of Zoological Nomenclature
The International Code of Zoological Nomenclature is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals...

. Prieto-Márquez defined Hadrosaurinae as only the lineage containing H. foulkii, and used the name Saurolophinae instead for the traditional grouping.

The following cladogram was recovered in the 2010 phylogenetic analysis by Prieto-Márquez after the removal of Barsboldia
Barsboldia
Barsboldia was a genus of large hadrosaurid dinosaur from the early Maastrichtian Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, partial pelvis, and some ribs...

.

Genera

Saurolophinae (formerly Hadrosaurinae) is usually considered to include the following genera:
  • Anasazisaurus
    Anasazisaurus
    Anasazisaurus is a genus of hadrosaurid ornithopod dinosaur that lived about 74 million years ago, in the Late Cretaceous Period. It was found in the Farmington Member of the Kirtland Formation, in the San Juan Basin of New Mexico, United States. Only a partial skull has been found to date...

  • Barsboldia
    Barsboldia
    Barsboldia was a genus of large hadrosaurid dinosaur from the early Maastrichtian Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, partial pelvis, and some ribs...

  • Brachylophosaurus
    Brachylophosaurus
    Brachylophosaurus was a mid-sized member of the hadrosaurid family of dinosaurs. It is known from several skeletons and bonebed material from the Judith River Formation of Montana and the Oldman Formation of Alberta, living about 76.5 million years ago....

  • Edmontosaurus
    Edmontosaurus
    Edmontosaurus is a genus of crestless hadrosaurid dinosaur. It contains two species: Edmontosaurus regalis and Edmontosaurus annectens. Fossils of E. regalis have been found in rocks of western North America that date from the late Campanian stage of the Cretaceous Period 73 million years ago,...

  • Gryposaurus
    Gryposaurus
    Gryposaurus was a genus of duckbilled dinosaur that lived about 83 to 75.5 million years ago, in the Late Cretaceous of North America...

  • Kerberosaurus
    Kerberosaurus
    Kerberosaurus was a genus of hadrosaurine duckbill dinosaur from the late Maastrichtian-age Upper Cretaceous Tsagayan Formation of Blagoveschensk, Amur Region, Russia...

  • Kritosaurus
    Kritosaurus
    Kritosaurus is an incompletely known but historically important genus of hadrosaurid dinosaur. It lived about 73 million years ago, in the Late Cretaceous of North America...

  • Lophorhothon
    Lophorhothon
    Lophorhothon is a genus of hadrosauroid dinosaur from the Late Cretaceous, the first genus of dinosaur discovered in Alabama.-Discovery and naming:...

  • Maiasaura
    Maiasaura
    Maiasaura is a large duck-billed dinosaur genus that lived in the area currently covered by the state of Montana in the Upper Cretaceous Period , about 74 million years ago....

  • Naashoibitosaurus
    Naashoibitosaurus
    Naashoibitosaurus is a genus of hadrosaurid dinosaur that lived about 73 million years ago, in the Late Cretaceous, and was found in the Kirtland Formation of the San Juan Basin in New Mexico, United States. Only a partial skeleton has been found to date...

  • Prosaurolophus
    Prosaurolophus
    Prosaurolophus is a genus of hadrosaurid dinosaur from the Late Cretaceous of North America. It is known from the remains of at least 25 individuals belonging to two species, including skulls and skeletons, but it remains obscure...

  • Saurolophus
    Saurolophus
    Saurolophus is a genus of large hadrosaurine duckbill that lived about 69.5-68.5 million years ago, in the Late Cretaceous of North America and Asia; it is one of the few genera of dinosaurs known from multiple continents. It is distinguished by a spike-like crest which projects up and back...

    (type
    Type genus
    In biological classification, a type genus is a representative genus, as with regard to a biological family. The term and concept is used much more often and much more formally in zoology than it is in botany, and the definition is dependent on the nomenclatural Code that applies:* In zoological...

    )
  • Shantungosaurus
    Shantungosaurus
    Shantungosaurus, meaning "Shandong Lizard", is a genus of saurolophine hadrosaurid dinosaurs found in the Late Cretaceous Wangshi Formation of the Shandong Peninsula in China.-Description:...

  • Willinakaqe
    Willinakaqe
    Willinakaqe is an extinct genus of saurolophine hadrosaurid dinosaur which lived during the late Cretaceous of the Río Negro Province of southern Argentina....

  • Wulagasaurus
    Wulagasaurus
    Wulagasaurus is a genus of hadrosaurine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China. Its remains were found in a bonebed in the latest Maastrichtian-age Yuliangze Formation, dated to 65.5 million years ago...

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