Grey Alder
Encyclopedia
Alnus incana is a species of alder
Alder
Alder is the common name of a genus of flowering plants belonging to the birch family . The genus comprises about 30 species of monoecious trees and shrubs, few reaching large size, distributed throughout the North Temperate Zone and in the Americas along the Andes southwards to...

 with a wide range across the cooler parts of the Northern Hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of its equator—the word hemisphere literally means “half sphere”. It is also that half of the celestial sphere north of the celestial equator...

.

It is a small to medium size tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 15-20 m tall with smooth grey bark even in old age, its life span being a maximum of 60-100 years. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are matt green, ovoid, 5-11 cm long and 4-8 cm broad. The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are catkin
Catkin
A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect pollinated . They contain many, usually unisexual flowers, arranged closely along a central stem which is often drooping...

s, appearing early in spring before the leaves emerge, the male catkins pendulous and 5-10 cm long, the female catkins 1.5 cm long and one cm broad when mature in late autumn. The seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s are small, 1-2 mm long, and light brown with a narrow encircling wing. The Grey Alder has a shallow root system, and is marked not only by vigorous production of stump suckers, but also by root suckers, especially in the northern parts of its range. The wood resembles that of the black alder, but is somewhat paler and of little value.

Subspecies

There are four to six subspecies, some treated as separate species by some authors:
  • Alnus incana subsp. incana (Grey Alder). Northern Europe
    Europe
    Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

     and northwestern Asia
    Asia
    Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

    , and central and southern Europe in mountains, mainly in the regions of the Alps
    Alps
    The Alps is one of the great mountain range systems of Europe, stretching from Austria and Slovenia in the east through Italy, Switzerland, Liechtenstein and Germany to France in the west....

    , Carpathians
    Carpathian Mountains
    The Carpathian Mountains or Carpathians are a range of mountains forming an arc roughly long across Central and Eastern Europe, making them the second-longest mountain range in Europe...

     and the Caucasus
    Caucasus
    The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

    .
  • Alnus incana subsp. hirsuta (Spach) Á. & D.Löve (=A. hirsuta Spach; Manchurian Alder). Northeastern Asia
    Asia
    Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

    , and central Asia in mountains.
  • Alnus incana subsp. kolaensis (N.I.Orlova) Á. & D.Löve. Subarctic northeast Europe.
  • Alnus incana subsp. oblongifolia (=A. oblongifolia; Arizona Alder). Southwestern North America
    North America
    North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

    .
  • Alnus incana subsp. rugosa (Du Roi) R.T.Clausen (=A. rugosa Du Roi; Speckled Alder). Northeastern North America
    North America
    North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

    .
  • Alnus incana subsp. tenuifolia (Nutt.) Breitung (=A. tenuifolia Nutt.; Thinleaf Alder or Mountain Alder). Northwestern North America
    North America
    North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

     and Alaska
    Alaska
    Alaska is the largest state in the United States by area. It is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait...

    .

Ecology

Alnus incana is a light-demanding, fast-growing tree that grows well on poorer soils. In central Europe, it is a colonist of alluvial land alongside mountain brooks and streams, occurring at elevations up to 1500 metres. However, it does not require moist soil, and will also colonize screes and shallow stony slopes. In the northern part of its range, it is a common tree species at sea level in forests, abandoned fields and on lakeshores. It is sometimes used for afforestation
Afforestation
Afforestation is the establishment of a forest or stand of trees in an area where there was no forest. Reforestation is the reestablishment of forest cover, either naturally or artificially...

 on non-fertile soils which it enriches by means of nitrogen fixing
Nitrogen fixation
Nitrogen fixation is the natural process, either biological or abiotic, by which nitrogen in the atmosphere is converted into ammonia . This process is essential for life because fixed nitrogen is required to biosynthesize the basic building blocks of life, e.g., nucleotides for DNA and RNA and...

 bacteria in its root nodules. Several species of Lepidoptera
Lepidoptera
Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

 use Grey Alder as a food plant for their caterpillar
Caterpillar
Caterpillars are the larval form of members of the order Lepidoptera . They are mostly herbivorous in food habit, although some species are insectivorous. Caterpillars are voracious feeders and many of them are considered to be pests in agriculture...

s. See List of Lepidoptera that feed on alders. In the Boreal forest area of Canada, A. incana is often associated with Black Spruce
Black Spruce
Picea mariana is a species of spruce native to northern North America, from Newfoundland west to Alaska, and south to northern New York, Minnesota and central British Columbia...

 in the forest type termed Black Spruce/Speckled Alder.
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