Forest fragmentation
Encyclopedia
Forest fragmentation is a form of habitat fragmentation
Habitat fragmentation
Habitat fragmentation as the name implies, describes the emergence of discontinuities in an organism's preferred environment , causing population fragmentation...

, occurring when forests are cut down in a manner that leaves relatively small, isolated patches of forest known as forest fragments or forest remnants. The intervening matrix that separates the remaining woodland patches can be natural open areas, farmland, or developed areas. Following the principles of island biogeography
Island biogeography
Island biogeography is a field within biogeography that attempts to establish and explain the factors that affect the species richness of natural communities. The theory was developed to explain species richness of actual islands...

, remnant woodlands act like islands of forest in a sea of pastures, fields, subdivisions, shopping malls, etc.

Natural Causes of Forest Fragmentation

Forests may also be fragmented by natural processes such as fire and changes in climate.

For example, 300 million years ago in the Carboniferous
Carboniferous
The Carboniferous is a geologic period and system that extends from the end of the Devonian Period, about 359.2 ± 2.5 Mya , to the beginning of the Permian Period, about 299.0 ± 0.8 Mya . The name is derived from the Latin word for coal, carbo. Carboniferous means "coal-bearing"...

 the tropical rainforests in Euramerica were fragmented due to a change in climate. There was a great loss of amphibian diversity and simultaneously the drier climate spurred the diversification of reptiles. These changes however, occurred gradually over million of years, not like the human driven destruction of tropical rainforests today.

Conservation implications

Forest fragmentation is one of the greatest threats to biodiversity
Biodiversity
Biodiversity is the degree of variation of life forms within a given ecosystem, biome, or an entire planet. Biodiversity is a measure of the health of ecosystems. Biodiversity is in part a function of climate. In terrestrial habitats, tropical regions are typically rich whereas polar regions...

 in forests, especially in the tropics. The problem of habitat destruction
Habitat destruction
Habitat destruction is the process in which natural habitat is rendered functionally unable to support the species present. In this process, the organisms that previously used the site are displaced or destroyed, reducing biodiversity. Habitat destruction by human activity mainly for the purpose of...

 that caused the fragmentation in the first place is compounded by :
  • the inability of individual forest fragments to support viable populations, especially of large vertebrates
  • the local extinction of species that do not have at least one fragment capable of supporting a viable population
  • edge effect
    Edge effect
    The edge effect in ecology is the effect of the juxtaposition or placing side by side of contrasting environments on an ecosystem.This term is commonly used in conjunction with the boundary between natural habitats, especially forests, and disturbed or developed land. Edge effects are especially...

    s that alter the conditions of the outer areas of the fragment, greatly reducing the amount of true forest interior habitat.


The effect of fragmentation on the flora
Flora
Flora is the plant life occurring in a particular region or time, generally the naturally occurring or indigenous—native plant life. The corresponding term for animals is fauna.-Etymology:...

 and fauna
Fauna
Fauna or faunæ is all of the animal life of any particular region or time. The corresponding term for plants is flora.Zoologists and paleontologists use fauna to refer to a typical collection of animals found in a specific time or place, e.g. the "Sonoran Desert fauna" or the "Burgess shale fauna"...

 of a forest patch depends on a) the size of the patch, and b) its degree of isolation. Isolation depends on the distance to the nearest similar patch, and the contrast with the surrounding areas. For example, if a cleared area is reforested
Reforestation
Reforestation is the natural or intentional restocking of existing forests and woodlands that have been depleted, usually through deforestation....

 or allowed to regenerate, the increasing structural diversity of the vegetation
Vegetation
Vegetation is a general term for the plant life of a region; it refers to the ground cover provided by plants. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic characteristics. It is broader...

will lessen the isolation of the forest fragments. However, when formerly forested lands are converted permanently to pastures, agricultural fields, or human-inhabited developed areas, the remaining forest fragments, and the biota within them, are often highly isolated.

Forest patches that are smaller or more isolated will lose species faster than those that are larger or less isolated. A large number of small forest "islands" typically cannot support the same biodiversity that a single contiguous forest would hold, even if their combined area is much greater than the single forest.
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