European Jackal
Encyclopedia
The European Jackal also known as the Caucasian Jackal or Reed Wolf is a subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 of golden jackal
Golden Jackal
The golden jackal , also known as the common jackal, Asiatic jackal, thos or gold-wolf is a Canid of the genus Canis indigenous to north and northeastern Africa, southeastern and central Europe , Asia Minor, the Middle East and southeast Asia...

 native to Southeast Europe
Southeast Europe
Southeast Europe or Southeastern Europe is a relatively recent political designation for the states of the Balkans. Writers such as Maria Todorova and Vesna Goldsworthy have suggested the use of the term Southeastern Europe to replace the word Balkans for the region, to minimize potential...

, Asia Minor
Asia Minor
Asia Minor is a geographical location at the westernmost protrusion of Asia, also called Anatolia, and corresponds to the western two thirds of the Asian part of Turkey...

 and the Caucasus
Caucasus
The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

. Its Latin name, moreoticus, means "of Morea
Morea
The Morea was the name of the Peloponnese peninsula in southern Greece during the Middle Ages and the early modern period. It also referred to a Byzantine province in the region, known as the Despotate of Morea.-Origins of the name:...

". It is one of the rarest and least known native canids in Europe, and is not recognized as a large carnivore by the Large Carnivore Initiative for Europe (LCIE), despite being the same size as the Iberian lynx
Iberian Lynx
The Iberian lynx, Lynx pardinus, is a critically endangered species native to the Iberian Peninsula in Southern Europe. It is one of the most endangered cat species in the world. According to the conservation group SOS Lynx, if this species died out, it would be one of the few feline extinctions...

, which is itself classed as such. Though mostly found in scattered populations within Eastern Europe, its range has grown to encompass parts of its former Eastern European range, as well as in Western Europe, which is thought to be attributable to a decline in wolf populations.

Physical description

It is one of the largest jackals in the world, with probably only the Egyptian jackal exceeding it in size; animals of both sexes average 120–125 cm (47–49 in) in total length and 10–13 kg (20-29 lb) in body weight. One adult male in North-Eastern Italy is recorded to have reached 14.9 kg (33 lb). The fur is coarse, and is generally brightly coloured with blackish tones on the back. The thighs, upper legs, ears and forehead are bright reddish chestnut. Jackals in Northern Dalmatia
Dalmatia
Dalmatia is a historical region on the eastern coast of the Adriatic Sea. It stretches from the island of Rab in the northwest to the Bay of Kotor in the southeast. The hinterland, the Dalmatian Zagora, ranges from fifty kilometers in width in the north to just a few kilometers in the south....

 have broader than average skulls, which is thought to result from human induced isolation from other populations, thus resulting in a new morphotype.

Diet

In the Caucasus
Caucasus
The Caucasus, also Caucas or Caucasia , is a geopolitical region at the border of Europe and Asia, and situated between the Black and the Caspian sea...

, jackals mainly hunt hare
Hare
Hares and jackrabbits are leporids belonging to the genus Lepus. Hares less than one year old are called leverets. Four species commonly known as types of hare are classified outside of Lepus: the hispid hare , and three species known as red rock hares .Hares are very fast-moving...

s, small rodents, pheasant
Pheasant
Pheasants refer to some members of the Phasianinae subfamily of Phasianidae in the order Galliformes.Pheasants are characterised by strong sexual dimorphism, males being highly ornate with bright colours and adornments such as wattles and long tails. Males are usually larger than females and have...

s, partridge
Partridge
Partridges are birds in the pheasant family, Phasianidae. They are a non-migratory Old World group.These are medium-sized birds, intermediate between the larger pheasants and the smaller quails. Partridges are native to Europe, Asia, Africa, and the Middle East...

s, duck
Duck
Duck is the common name for a large number of species in the Anatidae family of birds, which also includes swans and geese. The ducks are divided among several subfamilies in the Anatidae family; they do not represent a monophyletic group but a form taxon, since swans and geese are not considered...

s, coot
Coot
Coots are medium-sized water birds that are members of the rail family Rallidae. They constitute the genus Fulica. Coots have predominantly black plumage, and, unlike many of the rails, they are usually easy to see, often swimming in open water...

s, moorhen
Moorhen
Moorhens, sometimes called marsh hens, are medium-sized water birds that are members of the rail family Rallidae. They constitute the genus Gallinula....

s and passerine
Passerine
A passerine is a bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds or, less accurately, as songbirds, the passerines form one of the most diverse terrestrial vertebrate orders: with over 5,000 identified species, it has roughly...

s. They readily eat lizard
Lizard
Lizards are a widespread group of squamate reptiles, with nearly 3800 species, ranging across all continents except Antarctica as well as most oceanic island chains...

s, snake
Snake
Snakes are elongate, legless, carnivorous reptiles of the suborder Serpentes that can be distinguished from legless lizards by their lack of eyelids and external ears. Like all squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales...

s, frog
Frog
Frogs are amphibians in the order Anura , formerly referred to as Salientia . Most frogs are characterized by a short body, webbed digits , protruding eyes and the absence of a tail...

s, fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

, molluscs and insects. During the winter period, they will kill many nutrias and waterfowl
Waterfowl
Waterfowl are certain wildfowl of the order Anseriformes, especially members of the family Anatidae, which includes ducks, geese, and swans....

. During such times, jackals will surplus kill
Surplus killing
Surplus killing is the behavior predators exhibit when they kill more prey than they can immediately use. They may partially consume, cache, or abandon intact prey...

 and cache what they do not eat. Jackals will feed on fruits such as pear
Pear
The pear is any of several tree species of genus Pyrus and also the name of the pomaceous fruit of these trees. Several species of pear are valued by humans for their edible fruit, but the fruit of other species is small, hard, and astringent....

s, hawthorn
Crataegus
Crataegus , commonly called hawthorn or thornapple, is a large genus of shrubs and trees in the rose family, Rosaceae, native to temperate regions of the Northern Hemisphere in Europe, Asia and North America. The name hawthorn was originally applied to the species native to northern Europe,...

, dogwood
Dogwood
The genus Cornus is a group of about 30-60 species of woody plants in the family Cornaceae, commonly known as dogwoods. Most dogwoods are deciduous trees or shrubs, but a few species are nearly herbaceous perennial subshrubs, and a few of the woody species are evergreen...

 and the cones of Common Medlar
Common Medlar
Mespilus germanica, known as the common medlar is a large shrub or small tree, and the name of the fruit of this tree. Despite its Latin name, which means German or Germanic medlar, it is indigenous to southwest Asia and also southeastern Europe, mostly the Black Sea coasts of modern Turkey...

s. European jackals tend not to be as damaging to livestock as wolves and red fox
Red Fox
The red fox is the largest of the true foxes, as well as being the most geographically spread member of the Carnivora, being distributed across the entire northern hemisphere from the Arctic Circle to North Africa, Central America, and the steppes of Asia...

es are, though they can become a serious nuisance to small sized stock when in high numbers. The highest number of livestock damages occurred in southern Bulgaria
Bulgaria
Bulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...

: 1,053 attacks on small stock, mainly sheep and lambs, were recorded between 1982–87, along with some damages to newborn deer in game farms. In Greece
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

, rodents, insects, carrion, and fruits comprise the jackal’s diet, though they rarely eat garbage, due to large numbers of stray dogs preventing them access to places with high human density. Jackals in Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

 have been known to eat the eggs of the endangered green turtle. In Hungary
Hungary
Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...

, their most frequent prey are common vole
Common Vole
The Common Vole, Microtus arvalis, is a European mammal.- Distribution and habitat :The common vole is hardly restricted in means of distribution and habitat and inhabits large areas of Eurasia. As Microtus arvalis followed human civilisation, primary and secondary habitats can be distinguished...

s and bank vole
Bank Vole
The bank vole is a small vole with red-brown fur and some grey patches, with a tail about half as long as its body. A rodent, it lives in woodland areas and is around in length. The bank vole is found in western Europe and northern Asia...

s. Information on the diet of jackals in North-Eastern Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

 is scant, but it is certain that they prey on small roe deer
Roe Deer
The European Roe Deer , also known as the Western Roe Deer, chevreuil or just Roe Deer, is a Eurasian species of deer. It is relatively small, reddish and grey-brown, and well-adapted to cold environments. Roe Deer are widespread in Western Europe, from the Mediterranean to Scandinavia, and from...

 and hares.

Range

Bulgaria
Bulgaria
Bulgaria , officially the Republic of Bulgaria , is a parliamentary democracy within a unitary constitutional republic in Southeast Europe. The country borders Romania to the north, Serbia and Macedonia to the west, Greece and Turkey to the south, as well as the Black Sea to the east...

 has the largest European jackal population, which went through a 33–fold increase in range from the early sixties to mid-eighties. Factors aiding this increase include the replacement of natural forests with dense scrub, an increase in animal carcasses from state game farms, reductions in wolf populations and the abandonement of poisoning campaigns. In the early ‘90s, it was estimated that up to 5000 jackals populated Bulgaria. The population increased in 1994, and appears to have stabilised.
In Greece
Greece
Greece , officially the Hellenic Republic , and historically Hellas or the Republic of Greece in English, is a country in southeastern Europe....

, golden jackals are the rarest of the three wild extant canids there, having disappeared from Central and Western Greece and are now limited to discontinuous, isolated population clusters in Peloponnese
Peloponnese
The Peloponnese, Peloponnesos or Peloponnesus , is a large peninsula , located in a region of southern Greece, forming the part of the country south of the Gulf of Corinth...

, Fokida, Samos Island
Samos Island
Samos is a Greek island in the eastern Aegean Sea, south of Chios, north of Patmos and the Dodecanese, and off the coast of Asia Minor, from which it is separated by the -wide Mycale Strait. It is also a separate regional unit of the North Aegean region, and the only municipality of the regional...

, Halkidiki and North-eastern Greece. Currently, the largest population cluster is located in Nestos, north-eastern Greece. Although listed as “vulnerable” in the Red Data Book for Greek Vertebrates, the species has neither been officially declared as a game species nor as a protected one.

The subspecies is common in the Balkan Peninsula. Jackal populations have been increasing in Serbia
Serbia
Serbia , officially the Republic of Serbia , is a landlocked country located at the crossroads of Central and Southeast Europe, covering the southern part of the Carpathian basin and the central part of the Balkans...

 since the late 70's, and occur mainly in north-eastern Serbia and lower Srem
Srem
Śrem is a town on the Warta river in central Poland. It has been situated in the Greater Poland Voivodeship since 1999; from 1975 to 1998 it was part of the Poznań Voivodeship...

. Jackals are especially common near Negotin and Bela Palanka, where during the 1990s, about 500 specimens were shot. In Croatia
Croatia
Croatia , officially the Republic of Croatia , is a unitary democratic parliamentary republic in Europe at the crossroads of the Mitteleuropa, the Balkans, and the Mediterranean. Its capital and largest city is Zagreb. The country is divided into 20 counties and the city of Zagreb. Croatia covers ...

, 19 jackal packs have been confirmed in the north-western part of Ravni kotari
Ravni Kotari
Ravni Kotari is a geographical region in Croatia. It is situated in northern Dalmatia, around Zadar. It is bordered by Bukovica to the northwest, lower Krka on the southeast and the Adriatic Sea. The biggest settlement of the region is the town of Benkovac...

 and two on Vir Island. Golden jackals are listed as a protected species in Slovenia
Slovenia
Slovenia , officially the Republic of Slovenia , is a country in Central and Southeastern Europe touching the Alps and bordering the Mediterranean. Slovenia borders Italy to the west, Croatia to the south and east, Hungary to the northeast, and Austria to the north, and also has a small portion of...

, where they were first spotted in 1952 and have also established permanent territories there. In 2005, a probably vagrant female was accidentally shot near Gornji Grad
Gornji Grad
Gornji Grad is a town and a municipality in Slovenia. It lies on the Dreta River at the foothills of the Savinja Alps. Traditionally it belonged to the region of Styria and is now included in the Savinja statistical region....

 in the Upper Savinja Valley, Northern Slovenia. In 2009, two territorial groups of golden jackal were recorded in the Ljubljana Marshes area, Central Slovenia. It seems that the species continues to expand towards Central Europe. Jackal populations in Albania
Albania
Albania , officially known as the Republic of Albania , is a country in Southeastern Europe, in the Balkans region. It is bordered by Montenegro to the northwest, Kosovo to the northeast, the Republic of Macedonia to the east and Greece to the south and southeast. It has a coast on the Adriatic Sea...

 however are on the verge of extinction with possible occurrence in only three lowland wetland locations along the Adriatic Sea.

In Hungary
Hungary
Hungary , officially the Republic of Hungary , is a landlocked country in Central Europe. It is situated in the Carpathian Basin and is bordered by Slovakia to the north, Ukraine and Romania to the east, Serbia and Croatia to the south, Slovenia to the southwest and Austria to the west. The...

, where they are sometimes called "reed wolves", golden jackals had disappeared in the 1950s through hunting and habitat destruction, only to return in the late 70's, with the first breeding pairs being detected near the southern border in Transdanubia
Transdanubia
Transdanubia is a traditional region of Hungary.-Traditional interpretation:The borders of Transdanubia are the Danube river , the Drava and Mura rivers and the foothills of the Alps roughly along the border between Hungary and Austria .Transdanubia comprises the counties of Győr-Moson-Sopron,...

, then between the River Danube and Tisza
Tisza
The Tisza or Tisa is one of the main rivers of Central Europe. It rises in Ukraine, and is formed near Rakhiv by the junction of headwaters White Tisa, whose source is in the Chornohora mountains and Black Tisa, which springs in the Gorgany range...

. Golden jackals have increased greatly in number year by year, with some estimates indicating that they now outnumber red foxes. The sighting of a jackal near the Austrian border in the summer of 2007 indicated that they have spread throughout the country.

In Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

 the species at present is distributed in Friuli Venezia Giulia, Veneto
Veneto
Veneto is one of the 20 regions of Italy. Its population is about 5 million, ranking 5th in Italy.Veneto had been for more than a millennium an independent state, the Republic of Venice, until it was eventually annexed by Italy in 1866 after brief Austrian and French rule...

 and Trentino-Alto Adige/Südtirol. In the High Adriatic Hinterland its distribution has been recently udated by Lapini et al. (2009). It was hypothesized that the earliest evidence of golden jackals in Italy came from fossils found in Puglia dating back to the late Pleistocene
Pleistocene
The Pleistocene is the epoch from 2,588,000 to 11,700 years BP that spans the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....

, on the basis of fossil remains found in Romanelli Cave. Such samples were discovered in layers located under the isotopic stadium MIS 5, and consequently they can be more precisely attributed to the end of the middle Pleistocene. Moreover, these materials have been recently re-determined and correctly attributed to Canis moschbachensis, a small, ancient relative to the wolf. For these reasons Canis aureus should be considered a post-glacial invader of north-eastern Italy. In 1984 Canis aureus had reached the Province of Belluno
Province of Belluno
TheThe Province of Belluno is a province in the Veneto region of Italy. Its capital is the city of Belluno.It has an area of 3,678 km², and a total population of 214,026 .-Geography:...

, in 1985 a pack reproduced near Udine
Udine
Udine is a city and comune in northeastern Italy, in the middle of Friuli-Venezia Giulia region, between the Adriatic sea and the Alps , less than 40 km from the Slovenian border. Its population was 99,439 in 2009, and that of its urban area was 175,000.- History :Udine is the historical...

 (this group was eliminated in 1987), a road-killed jackal was collected in Veneto in 1992, and their presence was then confirmed in the Province of Gorizia
Province of Gorizia
The Province of Gorizia is a province in the autonomous Friuli–Venezia Giulia region of Italy.-Overview:Its capital is the city of Gorizia. It belonged to the Province of Udine between 1924 and 1927 and the communes of Sonzia, Plezzo, Bergogna, Caporetto, Tolmino, Circhina, Santa Lucia d'Isonzo,...

 and in the hinterland of the Gulf of Trieste
Gulf of Trieste
The Gulf of Trieste is a shallow bay of the Adriatic Sea, in the extreme northern part of the Mediterranean Sea. It is part of the Gulf of Venice and is shared by Italy, Slovenia and Croatia...

. In these cases, the specimens were usually roaming male subadults, though a family-group was discovered in Agordino in 1994. A young dead female was discovered on 10 December 2009 in Carnia
Carnia
Carnia is a historical-geographic region of Friuli, whose municipalities all belong to the province of Udine, which is part of the Friuli-Venezia Giulia Region.It covers the western and central part of the mountainous region of the Province of Udine...

, indicating that the species' range has continued to expand. Moreover, in the late summer of 2009, the species was also signalled in Trentino-Alto Adige/Südtirol, where it has surely reached the Puster Valley. The Italian branch of WWF estimates that jackal numbers in Italy may be underestimated. The golden jackal is a protected animal in Italy.

A dead adult was found close to the road near Podolí (Uherské Hradiště District)
Podolí (Uherské Hradiště District)
Podolí is a village and municipality in Uherské Hradiště District in the Zlín Region of the Czech Republic.The municipality covers an area of , and has a population of 806 ....

 in the Czech Republic
Czech Republic
The Czech Republic is a landlocked country in Central Europe. The country is bordered by Poland to the northeast, Slovakia to the east, Austria to the south, and Germany to the west and northwest....

, on 19 March 2006.

In Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

, Romania
Romania
Romania is a country located at the crossroads of Central and Southeastern Europe, on the Lower Danube, within and outside the Carpathian arch, bordering on the Black Sea...

, the North Black Sea coast and Caucasus region, the statuses of jackals there are largely unknown. There are indications of expansion in Romania and north-western Black Sea coast, while it is reportedly in decline in Turkey.

Origins and presence in European tradition

Although present in Europe, jackals are rarely featured in European folklore or literature. Being a rare and elusive animal, the European jackal was historically often assumed to be an invasive species
Invasive species
"Invasive species", or invasive exotics, is a nomenclature term and categorization phrase used for flora and fauna, and for specific restoration-preservation processes in native habitats, with several definitions....

 wherever it made its presence known: In Dalmatia
Dalmatia
Dalmatia is a historical region on the eastern coast of the Adriatic Sea. It stretches from the island of Rab in the northwest to the Bay of Kotor in the southeast. The hinterland, the Dalmatian Zagora, ranges from fifty kilometers in width in the north to just a few kilometers in the south....

, it was widely believed by the inhabitants of Korčula
Korcula
Korčula is an island in the Adriatic Sea, in the Dubrovnik-Neretva County of Croatia. The island has an area of ; long and on average wide — and lies just off the Dalmatian coast. Its 16,182 inhabitants make it the second most populous Adriatic island after Krk...

 Island that Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

n jackals were introduced to the island by the Republic of Venice
Republic of Venice
The Republic of Venice or Venetian Republic was a state originating from the city of Venice in Northeastern Italy. It existed for over a millennium, from the late 7th century until 1797. It was formally known as the Most Serene Republic of Venice and is often referred to as La Serenissima, in...

 to inflict damage on the Republic of Ragusa
Republic of Ragusa
The Republic of Ragusa or Republic of Dubrovnik was a maritime republic centered on the city of Dubrovnik in Dalmatia , that existed from 1358 to 1808...

. When in 1929 a male jackal appeared on Premuda
Premuda
Premuda is a little island in Croatia, off the northern Adriatic coast. It belongs to the north Dalmatian islands which are situated north-west from the county center Zadar. Premuda is approximately 10 km long, up to 1 km wide, and has an area of 9.2 square kilometers...

, the
islanders believed that it was brought to the island "out of a sheer malice." An African origin for Dalmatian jackals has been proven unlikely, as their skulls bare little similarities to those of African jackals, but are similar to those of jackals in Asia Minor. Sir William Jardine thought that jackals were first transported to Europe through the Muslim conquests
Muslim conquests
Muslim conquests also referred to as the Islamic conquests or Arab conquests, began with the Islamic prophet Muhammad. He established a new unified polity in the Arabian Peninsula which under the subsequent Rashidun and Umayyad Caliphates saw a century of rapid expansion of Muslim power.They...

. However, the fossil record indicates that the golden jackal likely colonised the European continent from Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

 during the Upper Holocene
Holocene
The Holocene is a geological epoch which began at the end of the Pleistocene and continues to the present. The Holocene is part of the Quaternary period. Its name comes from the Greek words and , meaning "entirely recent"...

 or late Pleistocene
Pleistocene
The Pleistocene is the epoch from 2,588,000 to 11,700 years BP that spans the world's recent period of repeated glaciations. The name pleistocene is derived from the Greek and ....

.

Surveys taken in the High Adriatic Hinterland indicate that the totality of people with first hand experience of jackals (hunters, game keepers and local people) regularly mistook red foxes affected by sarcoptic mange (or in a problematic state of moult) for golden jackals. The sighting of a true golden jackal however, was always referred to as a wolf, or a little wolf. This was verified both with photo-trapping sessions and with track studies, confirming previous observations on this matter. This erroneous and controversial perception of the golden jackal may be due to the fact that its presence is still not traditional, neither in Italian and Slovenian human culture, nor in hunting and game keeping traditions.
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