European Beech
Encyclopedia
Fagus sylvatica, the European Beech or Common Beech, is a deciduous
Deciduous
Deciduous means "falling off at maturity" or "tending to fall off", and is typically used in reference to trees or shrubs that lose their leaves seasonally, and to the shedding of other plant structures such as petals after flowering or fruit when ripe...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 belonging to the beech
Beech
Beech is a genus of ten species of deciduous trees in the family Fagaceae, native to temperate Europe, Asia and North America.-Habit:...

 family Fagaceae
Fagaceae
The family Fagaceae, or beech family, comprises about 900 species of both evergreen and deciduous trees and shrubs, which are characterized by alternate simple leaves with pinnate venation, unisexual flowers in the form of catkins, and fruit in the form of cup-like nuts. Fagaceous leaves are often...

.

Natural range

The natural range extends from southern Sweden
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

 (with some isolated locations in southern Norway) to central Italy
Italy
Italy , officially the Italian Republic languages]] under the European Charter for Regional or Minority Languages. In each of these, Italy's official name is as follows:;;;;;;;;), is a unitary parliamentary republic in South-Central Europe. To the north it borders France, Switzerland, Austria and...

, west to France
France
The French Republic , The French Republic , The French Republic , (commonly known as France , is a unitary semi-presidential republic in Western Europe with several overseas territories and islands located on other continents and in the Indian, Pacific, and Atlantic oceans. Metropolitan France...

, southern England
England
England is a country that is part of the United Kingdom. It shares land borders with Scotland to the north and Wales to the west; the Irish Sea is to the north west, the Celtic Sea to the south west, with the North Sea to the east and the English Channel to the south separating it from continental...

, northern Portugal
Portugal
Portugal , officially the Portuguese Republic is a country situated in southwestern Europe on the Iberian Peninsula. Portugal is the westernmost country of Europe, and is bordered by the Atlantic Ocean to the West and South and by Spain to the North and East. The Atlantic archipelagos of the...

, and central Spain
Spain
Spain , officially the Kingdom of Spain languages]] under the European Charter for Regional or Minority Languages. In each of these, Spain's official name is as follows:;;;;;;), is a country and member state of the European Union located in southwestern Europe on the Iberian Peninsula...

, and east to northwest Turkey
Turkey
Turkey , known officially as the Republic of Turkey , is a Eurasian country located in Western Asia and in East Thrace in Southeastern Europe...

, where it intergrades with the Oriental Beech
Oriental Beech
Fagus orientalis, the Oriental Beech, is a deciduous tree in the beech family Fagaceae. The natural range extends from southeast Bulgaria's Strandja mountain through northwest Turkey east to the Caucasus and Alborz Mountains of Iran. It is restricted to mountain forests, at 500-2,100 m altitude...

 (F. orientalis), which replaces it further east. In the Balkans
Balkans
The Balkans is a geopolitical and cultural region of southeastern Europe...

, it shows some hybridisation with Oriental Beech; these hybrid trees are named Fagus x taurica. In the southern part of its range around the Mediterranean, it grows only in mountain
Mountain
Image:Himalaya_annotated.jpg|thumb|right|The Himalayan mountain range with Mount Everestrect 58 14 160 49 Chomo Lonzorect 200 28 335 52 Makalurect 378 24 566 45 Mount Everestrect 188 581 920 656 Tibetan Plateaurect 250 406 340 427 Rong River...

 forests, at 600-1,800 m altitude. Although often regarded as native in southern England, recent evidence suggests that it did not arrive in England until about 4000 BC, or 2,000 years after the English Channel
English Channel
The English Channel , often referred to simply as the Channel, is an arm of the Atlantic Ocean that separates southern England from northern France, and joins the North Sea to the Atlantic. It is about long and varies in width from at its widest to in the Strait of Dover...

 formed after the ice age
Ice age
An ice age or, more precisely, glacial age, is a generic geological period of long-term reduction in the temperature of the Earth's surface and atmosphere, resulting in the presence or expansion of continental ice sheets, polar ice sheets and alpine glaciers...

s; it could have been an early introduction by Stone age
Stone Age
The Stone Age is a broad prehistoric period, lasting about 2.5 million years , during which humans and their predecessor species in the genus Homo, as well as the earlier partly contemporary genera Australopithecus and Paranthropus, widely used exclusively stone as their hard material in the...

 man
Human
Humans are the only living species in the Homo genus...

, who used the nuts
Nut (fruit)
A nut is a hard-shelled fruit of some plants having an indehiscent seed. While a wide variety of dried seeds and fruits are called nuts in English, only a certain number of them are considered by biologists to be true nuts...

 for food. The beech is classified as a native in the south of England and as a non-native in the north where it is often removed from 'native' woods. Localised pollen records have been recorded in the North of England from the Iron Age by Sir Harry Godwin. Changing climatic conditions may put beech populations in southern England under increased stress and while it may not be possible to maintain the current levels of beech in some sites it is thought that conditions for beech in north-west England will remain favourable or even improve. It is often planted in Britain.

Description

It is a large tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

, capable of reaching heights of up to 49 m (160 ft) tall and 3 m (10 ft) trunk diameter, though more typically 25–35 m (80–115 ft) tall and up to 1.5 m (5 ft) trunk diameter. A 10-year-old sapling will stand about 4 m (13 ft) tall. It has a typical lifespan of 150 to 200 years, though sometimes up to 300 years. The appearance varies according to its habitat
Habitat
* Habitat , a place where a species lives and grows*Human habitat, a place where humans live, work or play** Space habitat, a space station intended as a permanent settlement...

; in forest conditions, it tends to have a long, slender light-gray trunk with a narrow crown and erect branches, in isolation with good side light the trunk is short with a large and widely spreading crown with very long branches.

The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are alternate, simple, and entire or with a slightly crenate margin, 5–10 cm long and 3–7 cm broad, with 6-7 veins on each side of the leaf (7-10 veins in Fagus orientalis). When crenate, there is one point at each vein tip, never any points between the veins. The bud
Bud
In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of the stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be specialized to develop flowers or short shoots, or may have...

s are long and slender, 15–30 mm long and 2–3 mm thick, but thicker (to 4–5 mm) where the buds include flower buds.

The leaves of beech are often not abscissed
Abscission
Abscission is a term used in several areas of biology. In plant sciences it most commonly refers to the process by which a plant drops one or more of its parts, such as a leaf, fruit, flower or seed...

 in the autumn and instead remain on the tree until the spring. This process is called marcescence
Marcescence
Marcescence is the retention of dead plant organs that normally are shed. It is most obvious in deciduous trees that retain leaves through the winter. Several trees normally have marcescent leaves such as oak , beech and hornbeam . Marcescent leaves of pin oak complete development of their...

. This particularly occurs when trees are saplings or when plants are clipped as a hedge (making beech hedges attractive screens, even in winter), but it also often continues to occur on the lower branches when the tree is mature.

The European Beech starts to flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

 when it is between 30–80 years old. The flowers are small catkin
Catkin
A catkin or ament is a slim, cylindrical flower cluster, with inconspicuous or no petals, usually wind-pollinated but sometimes insect pollinated . They contain many, usually unisexual flowers, arranged closely along a central stem which is often drooping...

s which appear shortly after the leaves in spring. The seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s, called beechnuts, are small triangular nuts
Nut (fruit)
A nut is a hard-shelled fruit of some plants having an indehiscent seed. While a wide variety of dried seeds and fruits are called nuts in English, only a certain number of them are considered by biologists to be true nuts...

 15–20 mm long and 7–10 mm wide at the base; there are two nuts in each cupule, maturing in the autumn 5–6 months after pollination. Flower and seed production is particularly abundant in years following a hot, sunny and dry summer, though rarely for two years in a row. The nuts are an important food for bird
Bird
Birds are feathered, winged, bipedal, endothermic , egg-laying, vertebrate animals. Around 10,000 living species and 188 families makes them the most speciose class of tetrapod vertebrates. They inhabit ecosystems across the globe, from the Arctic to the Antarctic. Extant birds range in size from...

s, rodent
Rodent
Rodentia is an order of mammals also known as rodents, characterised by two continuously growing incisors in the upper and lower jaws which must be kept short by gnawing....

s and in the past also human
Human
Humans are the only living species in the Homo genus...

s. Slightly toxic to humans if eaten in large quantities due to the tannin
Tannin
A tannin is an astringent, bitter plant polyphenolic compound that binds to and precipitates proteins and various other organic compounds including amino acids and alkaloids.The term tannin refers to the use of...

s they contain, the nuts were nonetheless pressed to obtain an oil in 19th century England that was used for cooking and in lamps. They were also ground to make flour, which could be eaten after the tannin
Tannin
A tannin is an astringent, bitter plant polyphenolic compound that binds to and precipitates proteins and various other organic compounds including amino acids and alkaloids.The term tannin refers to the use of...

s were leached out by soaking.

Habitat

Climate and temperatures vary, though humidity needs to be constant. Little is required of the soil so long as it is well drained. Though not demanding of its soil type, the European Beech has several significant requirements: a humid atmosphere (precipitation well distributed throughout the year and frequent fogs) and well drained soil (it can not handle excessive stagnant water). It prefers moderately fertile ground, calcified or lightly acidic, therefore it is found more often on the side of a hill than at the bottom of a clayey basin. It tolerates rigorous winter cold, but is sensitive to spring frost. In Norway
Norway
Norway , officially the Kingdom of Norway, is a Nordic unitary constitutional monarchy whose territory comprises the western portion of the Scandinavian Peninsula, Jan Mayen, and the Arctic archipelago of Svalbard and Bouvet Island. Norway has a total area of and a population of about 4.9 million...

's oceanic climate
Oceanic climate
An oceanic climate, also called marine west coast climate, maritime climate, Cascadian climate and British climate for Köppen climate classification Cfb and subtropical highland for Köppen Cfb or Cwb, is a type of climate typically found along the west coasts at the middle latitudes of some of the...

 planted trees grow well as far north as in Trondheim
Trondheim
Trondheim , historically, Nidaros and Trondhjem, is a city and municipality in Sør-Trøndelag county, Norway. With a population of 173,486, it is the third most populous municipality and city in the country, although the fourth largest metropolitan area. It is the administrative centre of...

.

A beech forest is very dark and few species of plant are able to survive there, where the sun barely reaches the ground. Young beeches prefer some shade and may grow poorly in full sunlight. In a clear-cut forest a European Beech will germinate and then die of excessive dryness. Under oak
Oak
An oak is a tree or shrub in the genus Quercus , of which about 600 species exist. "Oak" may also appear in the names of species in related genera, notably Lithocarpus...

s with sparse leaf cover it will quickly surpass them in height and, due to the beech's dense foliage, the oaks will die from lack of sunlight. Foresters may assure the oaks' survival by cutting young beeches with a billhook 10 cm off the ground, which can produce magnificent bonsai
Bonsai
is a Japanese art form using miniature trees grown in containers. Similar practices exist in other cultures, including the Chinese tradition of penjing from which the art originated, and the miniature living landscapes of Vietnamese hòn non bộ...

.

The root system is shallow, even superficial, with large roots spreading out in all directions. European Beech forms ectomycorrhizas with a range of fungi including members of the genera Amanita
Amanita
The genus Amanita contains about 600 species of agarics including some of the most toxic known mushrooms found worldwide. This genus is responsible for approximately 95% of the fatalities resulting from mushroom poisoning, with the death cap accounting for about 50% on its own...

, Boletus
Boletus
Boletus is a genus of mushroom, comprising over 100 species. The genus Boletus was originally broadly defined and described by Elias Magnus Fries in 1821, essentially containing all fungi with pores...

, Cantharellus
Cantharellus
Cantharellus is a genus of popular edible mushrooms, commonly known as chanterelles . They are mycorrhizal fungi, meaning they form symbiotic associations with plants, making them very difficult to cultivate...

, Hebeloma
Hebeloma
Hebeloma is a genus of fungi. Found worldwide, it contains the poison pie or fairy cakes and the ghoul fungus Hebeloma is a genus of fungi. Found worldwide, it contains the poison pie or fairy cakes (Hebeloma crustuliniforme) and the ghoul fungus Hebeloma is a genus of fungi. Found worldwide, it...

and Lactarius
Lactarius
Lactarius is a genus of mushroom-producing fungi. The genus, collectively known commonly as milk-caps, are characterized by the fact that they exude a milky fluid if cut or damaged...

; these fungi are important in enhancing uptake of water and nutrients from the soil.

In the woodlands of southern Britain, beech is dominant over oak and elm south of a line from about north Suffolk across to Cardigan. Oak are the dominant forest trees north of this line. One of the most beautiful European Beech forests called Sonian Forest
Sonian Forest
The Sonian Forest is a forest that lies across the south-eastern part of Brussels, Belgium.The forest lies in the Flemish municipalities of Sint-Genesius-Rode, Hoeilaart, Overijse and Tervuren, in Uccle, Watermael-Boitsfort, Auderghem and Woluwe-Saint-Pierre in the Brussels-Capital Region and in...

 (Forêt de Soignes/Zoniënwoud) is found in the southeast of Brussels
Brussels
Brussels , officially the Brussels Region or Brussels-Capital Region , is the capital of Belgium and the de facto capital of the European Union...

, Belgium
Belgium
Belgium , officially the Kingdom of Belgium, is a federal state in Western Europe. It is a founding member of the European Union and hosts the EU's headquarters, and those of several other major international organisations such as NATO.Belgium is also a member of, or affiliated to, many...

. Beech is a dominant tree species in France and constitutes about 10% of French forests.

Phenology

Spring leaf budding by the European Beech is triggered by a combination of day length and temperature. Bud break each year is from the middle of April to the beginning of May, often with remarkable precision (within a few days). It is more precise in the north of its range than the south, and at 600 m than at sea level
Sea level
Mean sea level is a measure of the average height of the ocean's surface ; used as a standard in reckoning land elevation...

.

The European Beech invests significantly in summer and autumn for the following spring. Conditions in summer, particularly good rainfall, determine the number of leaves included in the buds. In autumn, the tree builds the reserves that will sustain it into spring. Given good conditions, a bud can produce a shoot with up to ten or more leaves. The terminal bud emits a hormonal substance in the spring that halts the development of additional buds. This tendency, though very strong at the beginning of their existence, becomes weaker in older trees.

It is only after the budding that root growth of the year begins. The first roots to appear are very thin (with a diameter of less than 0.5 mm). Later, after a wave of above ground growth, thicker roots grow in a steady fashion.

Uses

European Beech is a very popular ornamental tree in park
Park
A park is a protected area, in its natural or semi-natural state, or planted, and set aside for human recreation and enjoyment, or for the protection of wildlife or natural habitats. It may consist of rocks, soil, water, flora and fauna and grass areas. Many parks are legally protected by...

s and large garden
Garden
A garden is a planned space, usually outdoors, set aside for the display, cultivation, and enjoyment of plants and other forms of nature. The garden can incorporate both natural and man-made materials. The most common form today is known as a residential garden, but the term garden has...

s, not only in Europe, but also in North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

 and New Zealand
New Zealand
New Zealand is an island country in the south-western Pacific Ocean comprising two main landmasses and numerous smaller islands. The country is situated some east of Australia across the Tasman Sea, and roughly south of the Pacific island nations of New Caledonia, Fiji, and Tonga...

. It is frequently kept clipped to make attractive hedges. Since the early nineteenth century there have been a large number of ornamental cultivar
Cultivar
A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

s of European Beech made by horticultural selection, often repeatedly; they include:
  • Copper Beech or Purple Beech (Fagus sylvatica Purpurea Group) - leaves purple, in many selections turning deep spinach green by mid-summer. In the United States Charles Sprague Sargent noted the earliest appearance in a nurseryman's catalogue in 1820, but in 1859 "the finest Copper Beech in America... more than fifty feet high" was noted in the grounds of Thomas Ash, Esq., Throgs Neck, New York; it must have been more than forty years old at the time.
  • Fern-leaf Beech (Fagus sylvatica Heterophylla Group) - leaves deeply serrated to thread-like
  • Dwarf Beech
    Dwarf Beech
    The Dwarf Beech, Fagus sylvatica Tortuosa Group, is a rare Cultivar Group of the European Beech with less than 1500 older specimens in Europe. It is also known as Twisted Beech or Parasol Beech....

     (Fagus sylvatica Tortuosa Group) - distinctive twisted trunk and branches
  • Weeping Beech
    Weeping Beech
    The Weeping Beech, Fagus sylvatica "pendula", is a cultured variety of the deciduous European Beech. It is considered the most picturesque of all the weeping trees....

     (Fagus sylvatica Pendula Group) - branches pendulous
  • Dawyck Beech (Fagus sylvatica 'Dawyck') - fastigiate growth
  • Golden Beech (Fagus sylvatica 'Zlatia') - leaves golden in spring


The wood
Wood
Wood is a hard, fibrous tissue found in many trees. It has been used for hundreds of thousands of years for both fuel and as a construction material. It is an organic material, a natural composite of cellulose fibers embedded in a matrix of lignin which resists compression...

 of the European Beech is used in the manufacture of numerous objects and implements. Its fine and short grain makes it an easy wood to work with, easy to soak, dye (except its heartwood), varnish and glue. Steaming makes the wood even easier to machine. It has an excellent finish and is resistant to compression and splitting. Milling is sometimes difficult due to cracking and it is stiff when flexed. The density of the wood is 720 kg per cubic meter. It is particularly well suited for minor carpentry, particularly furniture. From chairs to parquetry (flooring) and staircases, the European Beech can do almost anything other than heavy structural support, so long as it is not left outdoors. Its hardness make it ideal for making wooden mallet
Mallet
A mallet is a kind of hammer, usually of rubber,or sometimes wood smaller than a maul or beetle and usually with a relatively large head.-Tools:Tool mallets come in different types, the most common of which are:...

s and workbench
Workbench
A workbench is sturdy table at which manual work is done. They range from simple flat surfaces to very complex designs that may be considered tools in themselves. Workbenches vary in size from tiny jewelers benches to the huge benches used by staircase makers...

 tops. The wood of the European Beech rots easily if it is not protected by a tar based on a distillate of its own bark (as used in railway sleepers). It is better for paper pulp than many other broadleaved trees though is only sometimes used for this. The code for its use in Europe is FASY (like FAgus SYlvatica). Common beech is also considered one of the best fire woods for fireplaces.

Primary Product AM 01, a smoke flavouring, is produced from Fagus sylvatica L.

The fruit of beeches are edible and have a nutty flavour. However it is not recommended to eat too many because they contain low concentrations of Trimethylamine
Trimethylamine
Trimethylamine is an organic compound with the formula N3. This colorless, hygroscopic, and flammable tertiary amine has a strong "fishy" odor in low concentrations and an ammonia-like odor at higher concentrations...

, which is slightly toxic. Roasting both improves flavour and reduces the amount of Trimethylamine.

Pathogens

Biscogniauxia nummularia
Biscogniauxia nummularia
Biscogniauxia nummularia is a plant pathogen in the Xylariaceae family, known as the Beech Tarcrust. The Latin 'nummus' meaning a coin, applied as the encrustations are often rounded and coin-like.-Views of the Beech Tarcrust:...

is an Ascomycete primary pathogen of beech trees, causing strip-canker and wood rot. It can be found at all times of year and is not edible.

External links

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