Chiselmouth
Encyclopedia
The chiselmouth is an unusual cyprinid
Cyprinid
The family Cyprinidae, from the Ancient Greek kyprînos , consists of the carps, the true minnows, and their relatives . Commonly called the carp family or the minnow family, its members are also known as cyprinids...

 fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

 of western North America
North America
North America is a continent wholly within the Northern Hemisphere and almost wholly within the Western Hemisphere. It is also considered a northern subcontinent of the Americas...

. It is named for the sharp hard plate on its lower jaw, which is used to scrape rocks for algae
Algae
Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms, such as the giant kelps that grow to 65 meters in length. They are photosynthetic like plants, and "simple" because their tissues are not organized into the many...

. It belongs into the monotypic
Monotypic
In biology, a monotypic taxon is a taxonomic group with only one biological type. The term's usage differs slightly between botany and zoology. The term monotypic has a separate use in conservation biology, monotypic habitat, regarding species habitat conversion eliminating biodiversity and...

 genus
Genus
In biology, a genus is a low-level taxonomic rank used in the biological classification of living and fossil organisms, which is an example of definition by genus and differentia...

 Acrocheilus and is a close relative of the Gila
Gila (genus)
Gila is a genus of fish belonging to the family Cyprinidae, native to North America and Central America. Species of Gila are collectively referred to as western chubs. The chiselmouth is a close relative...

western chubs (Simons & Mayden 1997).

The chiselmouth's body plan generally follows the standard cyprinid form, generally elongated and slightly compressed. The snout is very blunt, with the lower jaw's plate (which consists of cornified epithelium
Epithelium
Epithelium is one of the four basic types of animal tissue, along with connective tissue, muscle tissue and nervous tissue. Epithelial tissues line the cavities and surfaces of structures throughout the body, and also form many glands. Functions of epithelial cells include secretion, selective...

) jutting out slightly. Coloration is rather drab, dark brown above and lighter lower down. Many individuals also have a pattern of black dots, and younger fish may have a dark area at the base of the tail. The single dorsal fin
Dorsal fin
A dorsal fin is a fin located on the backs of various unrelated marine and freshwater vertebrates, including most fishes, marine mammals , and the ichthyosaurs...

 has 10 soft rays, while the anal fin and well-developed pelvic fins each have 9-10 rays. Chiselmouths can reach a length of 30 cm (12 in).

Young fish feed on surface insect
Insect
Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

s. When the chisel develops (at around 0.6 inches length), they shift to scraping, making short darting movements at the substrate to dislodge whatever is on it, and sucking it in. Although they consume filamentous algae, it seems not be digested much despite a long coiled intestine, and their primary food actually consists of diatom
Diatom
Diatoms are a major group of algae, and are one of the most common types of phytoplankton. Most diatoms are unicellular, although they can exist as colonies in the shape of filaments or ribbons , fans , zigzags , or stellate colonies . Diatoms are producers within the food chain...

s.

They are typically found in warmer parts of streams and rivers in the drainages of the Columbia River
Columbia River
The Columbia River is the largest river in the Pacific Northwest region of North America. The river rises in the Rocky Mountains of British Columbia, Canada, flows northwest and then south into the U.S. state of Washington, then turns west to form most of the border between Washington and the state...

, Fraser River
Fraser River
The Fraser River is the longest river within British Columbia, Canada, rising at Fraser Pass near Mount Robson in the Rocky Mountains and flowing for , into the Strait of Georgia at the city of Vancouver. It is the tenth longest river in Canada...

, and the Harney-Malheur system of the Great Basin
Great Basin
The Great Basin is the largest area of contiguous endorheic watersheds in North America and is noted for its arid conditions and Basin and Range topography that varies from the North American low point at Badwater Basin to the highest point of the contiguous United States, less than away at the...

. Some are found in lakes, migrating into streams to spawn. Although abundant in many parts of their range, behavior remains little-known.
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