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Chalcid wasp

 
Chalcid Wasp

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Chalcid wasp



 
 
Chalcid wasps (superfamily Chalcidoidea) belong to the insect
Insect

Insects are the biggest class of arthropods and the only ones with wings. They are the most diverse group of animals on the planet. They are most diverse at the equator and their diversity declines toward the poles....
 order Hymenoptera
Hymenoptera

Hymenoptera is one of the larger order s of insects, comprising the sawfly, wasps, bees, and ants. The name refers to the membranous wings of the insects, and is derived from the Ancient Greek language wikt:???? : membrane and wikt:pte??? : wing....
, and are one of the largest groups within the order, with some 22,000 known species, and an estimated total diversity of anywhere from 60,000 to more than 500,000 species, meaning the vast majority have yet to be discovered and described.

Most of the species are parasitoid
Parasitoid

A parasitoid is an organism that spends a significant portion of its biological life cycle attached to or within a single host organism which it ultimately kills in the process....
s of other insects, attacking the egg or larval stage of their host, though many other life cycles are known.






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Ic Leucospis Gigas
Chalcid wasps (superfamily Chalcidoidea) belong to the insect
Insect

Insects are the biggest class of arthropods and the only ones with wings. They are the most diverse group of animals on the planet. They are most diverse at the equator and their diversity declines toward the poles....
 order Hymenoptera
Hymenoptera

Hymenoptera is one of the larger order s of insects, comprising the sawfly, wasps, bees, and ants. The name refers to the membranous wings of the insects, and is derived from the Ancient Greek language wikt:???? : membrane and wikt:pte??? : wing....
, and are one of the largest groups within the order, with some 22,000 known species, and an estimated total diversity of anywhere from 60,000 to more than 500,000 species, meaning the vast majority have yet to be discovered and described.

Most of the species are parasitoid
Parasitoid

A parasitoid is an organism that spends a significant portion of its biological life cycle attached to or within a single host organism which it ultimately kills in the process....
s of other insects, attacking the egg or larval stage of their host, though many other life cycles are known. These hosts are to be found in at least 12 different insect orders including Lepidoptera
Lepidoptera

Lepidoptera is an order of insect that includes moths and butterfly. It is one of the most speciose orders in the class Insecta, encompassing moths and the three superfamilies of butterfly, skipper , and Hedylidae....
 (butterflies and moths), Diptera (true flies), Coleoptera
Beetle

Beetles are the group of insects with the largest number of known species. They are placed in the order Coleoptera , which contains more described species than in any other order in the animal, constituting about 25% of all known life-forms....
 (beetles), Hemiptera
Hemiptera

Hemiptera is an order of insects, comprising around 80,000 species of cicadas, aphids, planthoppers, leafhoppers, shield bugs, and others. They range in size from 1 mm to around 15 cm, and share a common arrangement of sucking mouthparts ....
 (true bugs), and other Hymenoptera, as well as two orders of Arachnida, and even one family of nematode
Nematode

The "roundworms" or "nematodes" are the most diverse phylum of body cavity, and one of the most diverse of all animals. Nematode species are very difficult to distinguish; over 80,000 have been described, of which over 15,000 are parasite....
s.

A few species are phytophagous and the larvae feed inside seeds, stems, and gall
Gall

Galls or plant galls are abnormal outgrowths of plant tissues and can be caused by various parasites, from fungi and bacterium, to insects and mites....
s. Generally beneficial to humans as a group, chalcids keep various crop pests under control, and many species have been imported to control insect pests.

They can be found almost everywhere, particularly on flowers, foliage, and in leaf litter, but are often overlooked due to their very small size, which includes the smallest of all known insects (see Dicopomorpha echmepterygis
Dicopomorpha echmepterygis

Dicopomorpha echmepterygis is a parasitic wasp in the family Mymaridae. The males of this species are the smallest of all known insects. They are blind and wingless and may be no more than 0.139 mm in length ....
).

Chalcids are tiny, dark-coloured wasps, often metallic blue or green with complex sculpturing on the body. They are also recognized by the characteristic reduced wing venation, similar to that seen in other superfamilies of parasitic wasp
Parasitic wasp

The term Parasitoid wasp refers to a large evolutionary grade of hymenopteran Superfamily, mainly in the Apocrita. They are primarily parasitoids of other animals, mostly other arthropods....
s. They have a significant part to play in ecosystem
Ecosystem

An ecosystem is a natural unit consisting of all plants, animals and micro-organisms in an area functioning together with all of the non-living physical factors of the environment....
s and their true importance must not be underestimated.

Taxonomy


Chalcidoidea is a superfamily of Hymenoptera
Hymenoptera

Hymenoptera is one of the larger order s of insects, comprising the sawfly, wasps, bees, and ants. The name refers to the membranous wings of the insects, and is derived from the Ancient Greek language wikt:???? : membrane and wikt:pte??? : wing....
, whose family constituency is in constant flux, as new hypotheses of relationships are constantly being proposed and rejected; with the advent of molecular systematics, it seems that the future will see further revisions of the classification in use today.

There are nineteen extant families recognized at present:

  • Agaonidae Walker
    Francis Walker (entomologist)

    Francis Walker was an England entomologist. He was one of the most prolific authors in entomology, and stirred controversy during his later life as his publications resulted in a huge number of junior synonyms....
    , 1846
  • Aphelinidae
    Aphelinidae

    Aphelinidae is a moderate-sized family of tiny parasitic wasps, with some 1160 described species in some 35 genera. These minute insects are challenging to study as they deteriorate rapidly after death unless extreme care is taken , making identification of most museum specimens difficult....
     Thomson
    Carl Gustaf Thomson

    Carl Gustaf Thomson was a Sweden entomologist.Thomson studied at Lund. He was the author of Coleoptera Scandinaviae , Skandinaviens inseckta and Skandinaviens Hymenoptera ....
    , 1876
  • Chalcididae
    Chalcididae

    The Chalcididae are a moderate-sized family within the Chalcidoidea, composed mostly of parasitoids and a few hyperparasites. The family is apparently polyphyletic, though the different subfamilies may each be monophyletic, and some may be elevated to family status in the near future....
     Latreille
    Pierre André Latreille

    Pierre Andr? Latreille was a France entomology. His works describing insects assigned many of the insect taxa still in use today.Latreille was born into a humble family of Brive-la-Gaillarde, Corr?ze, and in 1778 entered the Jean Lemoine in Paris....
    , 1817
  • Encyrtidae
    Encyrtidae

    Encyrtidae is a large family of parasitic wasps, with some 3710 described species in some 455 genera . The larvae of the majority are primary parasitoids on Hemiptera, though other hosts are attacked, and details of the life history can be variable ....
     Walker
    Francis Walker (entomologist)

    Francis Walker was an England entomologist. He was one of the most prolific authors in entomology, and stirred controversy during his later life as his publications resulted in a huge number of junior synonyms....
    , 1837
  • Eucharitidae
    Eucharitidae

    The Eucharitidae are a specialized group of ant parasitoids within the Chalcidoidea. There are 53 genera and >470 species worldwide, mostly in the tropics; very few genera are shared between the Old World and New World....
     Latreille
    Pierre André Latreille

    Pierre Andr? Latreille was a France entomology. His works describing insects assigned many of the insect taxa still in use today.Latreille was born into a humble family of Brive-la-Gaillarde, Corr?ze, and in 1778 entered the Jean Lemoine in Paris....
    , 1809
  • Eulophidae
    Eulophidae

    Eulophidae is a large family of hymenopteran insects, with over 4,300 described species in some 300 genera . The family as presently defined also includes the genus Elasmus, which was previously treated as a separate family, "Elasmidae", and is now treated as a subfamily of Eulophidae....
     Westwood
    John Obadiah Westwood

    John Obadiah Westwood was an England entomologist and archaeologist also noted for his artistic talents.Born in Sheffield, he studied to be a lawyer but abandoned that for his scientific interests....
    , 1829
    (including Elasmidae)
  • Eupelmidae
    Eupelmidae

    Eupelmidae is a family of parasitic wasps in the superfamily Chalcidoidea. The group is apparently polyphyletic, though the different subfamilies may each be monophyletic, and may be elevated to family status in the near future....
     Walker
    Francis Walker (entomologist)

    Francis Walker was an England entomologist. He was one of the most prolific authors in entomology, and stirred controversy during his later life as his publications resulted in a huge number of junior synonyms....
    , 1833
  • Eurytomidae
    Eurytomidae

    Eurytomidae is a family within the superfamily Chalcidoidea. The group is apparently polyphyletic, though the different subfamilies may each be monophyletic, and may be elevated to family status in the near future....
     Walker
    Francis Walker (entomologist)

    Francis Walker was an England entomologist. He was one of the most prolific authors in entomology, and stirred controversy during his later life as his publications resulted in a huge number of junior synonyms....
    , 1832
  • Leucospidae
    Leucospidae

    The Leucospidae are a small, specialized group within the Chalcidoidea, composed exclusively of ectoparasitoids of aculeata or bees. They are typically mimicry of bees or stinging wasps, often black with yellow, red, or white markings, sometimes metallic, with a robust mesosoma and very strong sculpturing....
     Fabricius
    Johan Christian Fabricius

    Johan Christian Fabricius was a Denmark entomologist and economist.Fabricius was born at T?nder in the duchy of Schleswig. He studied at the gymnasium at Altona and entered the University of Copenhagen in 1762....
    , 1775
  • Mymaridae Haliday
    Alexander Henry Haliday

    Alexander Henry Haliday, also known as Enrico Alessandro Haliday and Alexis Heinrich Haliday , was an Ireland entomologist. He is primarily known for his work on Hymenoptera, Diptera and Thysanoptera, but Haliday worked on all insect orders and on many aspects of entomology....
    , 1833
  • Ormyridae
    Ormyridae

    Ormyridae is a small family of parasitic wasps in the superfamily Chalcidoidea. They are either parasitoids or hyperparasite on gall-forming insects, primarily Cynipidae and Tephritidae....
     Förster
    Arnold Förster

    Arnold F?rster was a Germany entomologist, who worked mainly on Coleoptera and Hymenoptera....
    , 1856
  • Perilampidae
    Perilampidae

    The Perilampidae are a small family within the Chalcidoidea, composed mostly of hyperparasites. The family is closely related to the Eucharitidae, and the eucharitids appear to have evolved from within the Perilampidae, thus rendering the family paraphyletic ....
     Latreille
    Pierre André Latreille

    Pierre Andr? Latreille was a France entomology. His works describing insects assigned many of the insect taxa still in use today.Latreille was born into a humble family of Brive-la-Gaillarde, Corr?ze, and in 1778 entered the Jean Lemoine in Paris....
    , 1809
  • Pteromalidae
    Pteromalidae

    Pteromalidae is a very large family of parasitic wasps, with some 3,450 described species in some 640 genera . The subfamily-level divisions of the family are highly contentious and unstable, and there is no question that the family is completely artificial, composed of numerous distantly-related groups ....
     Dalman
    Johan Wilhelm Dalman

    Johan Wilhelm Dalman was a Sweden physician and a natural history. He first studied at Christianfeld in Schleswig-Holstein then at the University of Lund and the University of Uppsala....
    , 1820
  • Rotoitidae
    Rotoitidae

    Rotoitidae is a very small family of rare, relictual parasitic wasps in the superfamily Chalcidoidea. There are only two species known, each in their own genus, one from New Zealand and one from Chile....
     Boucek & Noyes
    Noyes

    Noyes, a surname and place name, may refer to:...
    , 1987
  • Signiphoridae
    Signiphoridae

    Signiphoridae is a small family of parasitic wasps in the superfamily Chalcidoidea. They are almost exclusively associated with scale insects, either as parasitoids or parasitoids of other parasitic wasps or Diptera....
     Ashmead, 1880
  • Tanaostigmatidae
    Tanaostigmatidae

    Tanaostigmatidae is a small family of parasitic wasps in the superfamily Chalcidoidea. They are almost exclusively phytophagy insects, forming galls in plant stems, leaves, or seeds....
     Howard, 1890
  • Tetracampidae
    Tetracampidae

    Tetracampidae is a small family of parasitic wasps in the superfamily Chalcidoidea. They are parasitoids of phytophagy insects, primarily Diptera....
     Förster
    Arnold Förster

    Arnold F?rster was a Germany entomologist, who worked mainly on Coleoptera and Hymenoptera....
    , 1856
  • Torymidae
    Torymidae

    Torymidae is a family of wasps that consists of attractive metallic species with enlarged hind legs, and generally with a long ovipositor. Many are parasitoids on gall-forming insects, and some are phytophagy species, sometimes usurping the galls formed by other insects....
     Walker
    Francis Walker (entomologist)

    Francis Walker was an England entomologist. He was one of the most prolific authors in entomology, and stirred controversy during his later life as his publications resulted in a huge number of junior synonyms....
    , 1833
  • Trichogrammatidae
    Trichogrammatidae

    The family Trichogrammatidae are tiny wasps in the Chalcidoidea that include some of the smallest of all insects, with most species having adults less than 1 mm in length....
     Haliday
    Alexander Henry Haliday

    Alexander Henry Haliday, also known as Enrico Alessandro Haliday and Alexis Heinrich Haliday , was an Ireland entomologist. He is primarily known for his work on Hymenoptera, Diptera and Thysanoptera, but Haliday worked on all insect orders and on many aspects of entomology....
    , 1851


There is also one extinct family, Khutelchalcididae Rasnitsyn
Alex Rasnitsyn

Alexandr Pavlovich Rasnitsyn is one of the world's leading experts in palaeoentomology. His scientific interests are centered on the palaeontology, phylogeny, and taxonomy of the hymenopteran insects and of the insects in general....
, Basibuyuk & Quicke, 2004
.

Of these families, at least five are known to be artificial groups (paraphyletic), and are being - or will be - divided into several families, or perhaps fused with existing families. The most problematic, the Pteromalidae, is, in some classifications, supposed to contain as many as 8 independent lineages, grouped together because of superficial similarities.

Identification

  • Key to families Grissell, E. E., and M. E. Schauff. 1990. A handbook of the families of Nearctic Chalcidoidea (Hymenoptera).Entomological Society of Washington (Washington, D.C.) Handbook 1:1-85.Online at


External links

  • Wrongly titled (as Chalcididae) gallery of images..
  • Pictorial overview.