Capelin
Encyclopedia
The capelin or caplin, Mallotus villosus, is a small forage fish
Forage fish
Forage fish, also called prey fish or bait fish, are small fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the food chain on plankton, often by filter feeding...

 of the smelt family found in the Atlantic
Atlantic Ocean
The Atlantic Ocean is the second-largest of the world's oceanic divisions. With a total area of about , it covers approximately 20% of the Earth's surface and about 26% of its water surface area...

 and Arctic
Arctic Ocean
The Arctic Ocean, located in the Northern Hemisphere and mostly in the Arctic north polar region, is the smallest and shallowest of the world's five major oceanic divisions...

 oceans. In summer, it grazes on dense swarms of plankton
Plankton
Plankton are any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. That is, plankton are defined by their ecological niche rather than phylogenetic or taxonomic classification...

 at the edge of the ice shelf. Larger capelin also eat a great deal of krill
Krill
Krill is the common name given to the order Euphausiacea of shrimp-like marine crustaceans. Also known as euphausiids, these small invertebrates are found in all oceans of the world...

 and other crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s. Whale
Whale
Whale is the common name for various marine mammals of the order Cetacea. The term whale sometimes refers to all cetaceans, but more often it excludes dolphins and porpoises, which belong to suborder Odontoceti . This suborder also includes the sperm whale, killer whale, pilot whale, and beluga...

s, seal
Pinniped
Pinnipeds or fin-footed mammals are a widely distributed and diverse group of semiaquatic marine mammals comprising the families Odobenidae , Otariidae , and Phocidae .-Overview: Pinnipeds are typically sleek-bodied and barrel-shaped...

s, cod, squid, mackerel, beluga whales and seabird
Seabird
Seabirds are birds that have adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding niches have resulted in similar adaptations...

s all prey on capelin in particular during the spawning season of the capelin while it migrates southwards. Capelin spawn on sandy beaches and sandy bottom at the age of 2–6 years, and have an extremely high mortality rate on the beaches after spawning, for males close to 100% mortality.
Males reach 20 cm in length, while females are up to 25 cm long. They are olive-colored dorsally, shading to silver on sides. Males have a translucent ridge on both sides of their bodies. The ventral aspects of the males iridesce reddish at the time of spawn.

In years with large quantities of herring
Herring
Herring is an oily fish of the genus Clupea, found in the shallow, temperate waters of the North Pacific and the North Atlantic oceans, including the Baltic Sea. Three species of Clupea are recognized. The main taxa, the Atlantic herring and the Pacific herring may each be divided into subspecies...

 in the Barents Sea
Barents Sea
The Barents Sea is a marginal sea of the Arctic Ocean, located north of Norway and Russia. Known in the Middle Ages as the Murman Sea, the sea takes its current name from the Dutch navigator Willem Barents...

, capelin seem to be heavily affected. Probably both food competition and herring feeding on capelin larvae lead to collapses in the capelin stock.

Commercially, capelin is used for fish meal
Fish meal
Fish meal, or fishmeal, is a commercial product made from both whole fish and the bones and offal from processed fish. It is a brown powder or cake obtained by rendering pressing the cooked whole fish or fish trimmings to remove most of the fish oil and water, and then ground...

 and oil
Fish oil
Fish oil is oil derived from the tissues of oily fish. Fish oils contain the omega-3 fatty acids eicosapentaenoic acid , and docosahexaenoic acid , precursors of certain eicosanoids that are known to reduce inflammation throughout the body, and are thought to have many health benefits.Fish do not...

 industry products, but is also appreciated as food. The flesh is agreeable in flavor, resembling herring. Capelin roe
Roe
Roe or hard roe is the fully ripe internal egg masses in the ovaries, or the released external egg masses of fish and certain marine animals, such as shrimp, scallop and sea urchins...

 ("masago") is considered as a high value product. It is also commonly mixed with wasabi
Wasabi
, also known as Japanese horseradish, is a member of the Brassicaceae family, which includes cabbages, horseradish, and mustard. Its root is used as a condiment and has an extremely strong flavor. Its hotness is more akin to that of a hot mustard rather than the capsaicin in a chili pepper,...

 and sold as "wasabi caviar". Sometimes masago is used as a substitute for tobiko
Tobiko
Tobiko is the Japanese word for the flying fish roe used to create certain types of sushi. Tobiko is sometimes used as an ingredient in California rolls....

, flying fish roe, due to its similarity and taste.

Capelin is an important forage fish
Forage fish
Forage fish, also called prey fish or bait fish, are small fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the food chain on plankton, often by filter feeding...

, and is essential as the key food of the Atlantic cod
Atlantic cod
The Atlantic cod, Gadus morhua, is a well-known demersal food fish belonging to the family Gadidae. It is also commercially known as cod, codling or haberdine....

. The North-East Atlantic Cod and Capelin fisheries therefore are managed by a multi-species approach developed by the main resource owners Norway and Russia.

In the provinces of Quebec
Quebec
Quebec or is a province in east-central Canada. It is the only Canadian province with a predominantly French-speaking population and the only one whose sole official language is French at the provincial level....

 (particularly in the Gaspé peninsula
Gaspé Peninsula
The Gaspésie , or Gaspé Peninsula or the Gaspé, is a peninsula along the south shore of the Saint Lawrence River in Quebec, Canada, extending into the Gulf of Saint Lawrence...

) and Newfoundland and Labrador
Newfoundland and Labrador
Newfoundland and Labrador is the easternmost province of Canada. Situated in the country's Atlantic region, it incorporates the island of Newfoundland and mainland Labrador with a combined area of . As of April 2011, the province's estimated population is 508,400...

 in Canada, it is a regular summertime practice to go to the beach
Beach
A beach is a geological landform along the shoreline of an ocean, sea, lake or river. It usually consists of loose particles which are often composed of rock, such as sand, gravel, shingle, pebbles or cobblestones...

 and scoop the capelin up in nets or whatever is available, as the capelin "roll in" in the millions each year at the end of June or in early July.

Capelin migration

Capelin move inshore in large schools
Shoaling and schooling
In biology, any group of fish that stay together for social reasons are said to be shoaling , and if, in addition, the group is swimming in the same direction in a coordinated manner, they are said to be schooling . In common usage, the terms are sometimes used rather loosely...

 to spawn
Spawn (biology)
Spawn refers to the eggs and sperm released or deposited, usually into water, by aquatic animals. As a verb, spawn refers to the process of releasing the eggs and sperm, also called spawning...

 and migrate
Fish migration
Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousands of kilometres...

 in spring and summer to feed in the plankton
Plankton
Plankton are any drifting organisms that inhabit the pelagic zone of oceans, seas, or bodies of fresh water. That is, plankton are defined by their ecological niche rather than phylogenetic or taxonomic classification...

-rich oceanic area between Iceland
Iceland
Iceland , described as the Republic of Iceland, is a Nordic and European island country in the North Atlantic Ocean, on the Mid-Atlantic Ridge. Iceland also refers to the main island of the country, which contains almost all the population and almost all the land area. The country has a population...

, Greenland
Greenland
Greenland is an autonomous country within the Kingdom of Denmark, located between the Arctic and Atlantic Oceans, east of the Canadian Arctic Archipelago. Though physiographically a part of the continent of North America, Greenland has been politically and culturally associated with Europe for...

, and Jan Mayen
Jan Mayen
Jan Mayen Island is a volcanic island in the Arctic Ocean and part of the Kingdom of Norway. It is long and 373 km2 in area, partly covered by glaciers . It has two parts: larger northeast Nord-Jan and smaller Sør-Jan, linked by an isthmus wide...

. Capelin distribution and migration is linked with ocean current
Ocean current
An ocean current is a continuous, directed movement of ocean water generated by the forces acting upon this mean flow, such as breaking waves, wind, Coriolis effect, cabbeling, temperature and salinity differences and tides caused by the gravitational pull of the Moon and the Sun...

s and water masses. Around Iceland, maturing capelin usually undertake extensive northward feeding migration
Fish migration
Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousands of kilometres...

s in spring and summer and the return migration takes place in September to November. The spawning migration starts from north of Iceland in December to January. In a paper published in 2009, researchers from Iceland recount their application of an interacting particle model to the capelin stock around Iceland, successfully predicting the spawning migration route for 2008.
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