Calamistrum
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In spider
Spider
Spiders are air-breathing arthropods that have eight legs, and chelicerae with fangs that inject venom. They are the largest order of arachnids and rank seventh in total species diversity among all other groups of organisms...

s, the calamistrum is a row of specialized leg bristles used to comb out fine bands of silk
Spider silk
Spider silk is a protein fiber spun by spiders. Spiders use their silk to make webs or other structures, which function as nets to catch other animals, or as nests or cocoons for protection for their offspring...

. It is only found on cribellate spiders, that is, spiders that possess the spinning organ known as the cribellum
Cribellum
The cribellum is a silk spinning organ found in certain spiders. Unlike normal spinnerets, the cribellum consists of one or more plates covered in thousands of tiny spigots. These spigots produce extremely fine fibers which are combed out by the spider's calamistrum, producing silk with a wooly...

. The calamistrum and cribellum are used to form the hackled bands of silk which are characteristic of the webs of these spiders. The calamistrum is found on the upper margin of the metatarsus
Arthropod leg
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments are of Latin origin, and may be confused with terms for bones: coxa , trochanter , femur, tibia, tarsus, ischium, metatarsus, carpus, dactylus ,...

 of the hind legs. Each bristle of the calamistrum is serrated on one side and smooth on the other.

The length of a spider's calamistrum is always equal to or greater than the width of the cribellum. The ratio between calamistrum length and cribellum width varies greatly, however, even among related species. This is likely due to differences in spinning behavior and as well as differences in the size and shape of the legs and abdomen.

When male cribellate spiders reach sexual maturity
Sexual maturity
Sexual maturity is the age or stage when an organism can reproduce. It is sometimes considered synonymous with adulthood, though the two are distinct...

, they either lose the cribellum and calamistrum or retain them in a vestigal form.
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