Cacao
Encyclopedia
Theobroma cacao also cacao tree and cocoa tree, is a small (4–8 m or 15–26 ft tall) evergreen
Evergreen
In botany, an evergreen plant is a plant that has leaves in all seasons. This contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.There are many different kinds of evergreen plants, both trees and shrubs...

 tree
Tree
A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to...

 in the family Sterculiaceae
Sterculiaceae
Sterculiaceae is a botanical name for a group of flowering plants at the rank of family, which is now considered obsolete. As is true for any botanical name, the circumscription, status and placement of the taxon has varied with taxonomic point of view...

 (alternatively Malvaceae
Malvaceae
Malvaceae, or the mallow family, is a family of flowering plants containing over 200 genera with close to 2,300 species. Judd & al. Well known members of this family include okra, jute and cacao...

), native to the deep tropical region of the Americas. Its seeds are used to make cocoa powder and chocolate
Chocolate
Chocolate is a raw or processed food produced from the seed of the tropical Theobroma cacao tree. Cacao has been cultivated for at least three millennia in Mexico, Central and South America. Its earliest documented use is around 1100 BC...

. There are two prominent competing hypotheses about the origins of the domestication of the originally wild Theobroma cacao tree. One is that wild examples were originally distributed from southeastern Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 to the Amazon basin
Amazon Basin
The Amazon Basin is the part of South America drained by the Amazon River and its tributaries that drains an area of about , or roughly 40 percent of South America. The basin is located in the countries of Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela...

, with domestication taking place both in the Lacandon area of Mexico and in lowland South America
South America
South America is a continent situated in the Western Hemisphere, mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere. The continent is also considered a subcontinent of the Americas. It is bordered on the west by the Pacific Ocean and on the north and east...

. But recent studies of Theobroma cacao genetics seem to show that the plant originated in the Amazon and was distributed by humans throughout Central America
Central America
Central America is the central geographic region of the Americas. It is the southernmost, isthmian portion of the North American continent, which connects with South America on the southeast. When considered part of the unified continental model, it is considered a subcontinent...

 and Mesoamerica
Mesoamerica
Mesoamerica is a region and culture area in the Americas, extending approximately from central Mexico to Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica, within which a number of pre-Columbian societies flourished before the Spanish colonization of the Americas in the 15th and...

.

The tree is today found only growing wild in the low foothills of the Andes
Andes
The Andes is the world's longest continental mountain range. It is a continual range of highlands along the western coast of South America. This range is about long, about to wide , and of an average height of about .Along its length, the Andes is split into several ranges, which are separated...

 at elevations of around 200–400 m (650–1300 ft) in the Amazon
Amazon River
The Amazon of South America is the second longest river in the world and by far the largest by waterflow with an average discharge greater than the next seven largest rivers combined...

 and Orinoco
Orinoco
The Orinoco is one of the longest rivers in South America at . Its drainage basin, sometimes called the Orinoquia, covers , with 76.3% of it in Venezuela and the remainder in Colombia...

 river
River
A river is a natural watercourse, usually freshwater, flowing towards an ocean, a lake, a sea, or another river. In a few cases, a river simply flows into the ground or dries up completely before reaching another body of water. Small rivers may also be called by several other names, including...

 basins. It requires a humid climate with regular rainfall and good soil. It is an understory
Understory
Understory is the term for the area of a forest which grows at the lowest height level below the forest canopy. Plants in the understory consist of a mixture of seedlings and saplings of canopy trees together with understory shrubs and herbs...

 tree, growing best with some overhead shade. The leaves
Leaf
A leaf is an organ of a vascular plant, as defined in botanical terms, and in particular in plant morphology. Foliage is a mass noun that refers to leaves as a feature of plants....

 are alternate, entire, unlobed, 10–40 cm (4–16 in) long and 5–20 cm (2–8 in) broad.

The flower
Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to effect reproduction, usually by providing a mechanism for the union of sperm with eggs...

s are produced in clusters directly on the trunk
Trunk (botany)
In botany, trunk refers to the main wooden axis of a tree that supports the branches and is supported by and directly attached to the roots. The trunk is covered by the bark, which is an important diagnostic feature in tree identification, and which often differs markedly from the bottom of the...

 and older branches; they are small, 1–2 cm (1/2–1 in) diameter, with pink calyx. While many of the world's flowers are pollinated by bees (Hymenoptera
Hymenoptera
Hymenoptera is one of the largest orders of insects, comprising the sawflies, wasps, bees and ants. There are over 130,000 recognized species, with many more remaining to be described. The name refers to the heavy wings of the insects, and is derived from the Ancient Greek ὑμήν : membrane and...

) or butterflies/moths
Moths
Moths may refer to:* Gustav Moths , German rower* The Moths!, an English indie rock band* MOTHS, members of the Memorable Order of Tin Hats...

 (Lepidoptera
Lepidoptera
Lepidoptera is a large order of insects that includes moths and butterflies . It is one of the most widespread and widely recognizable insect orders in the world, encompassing moths and the three superfamilies of butterflies, skipper butterflies, and moth-butterflies...

), cacao flowers are pollinated by tiny flies, Forcipomyia
Forcipomyiinae
Forcipomyiinae is a subfamily of biting midges. In this subfamily, both anterior and posterior prolegs are present on larvae. Larvae are both terrestrial and aquatic, and feed primarily on algae and fungi. Some species in the genus Forcipomyia are important pollinators of cacao.- References :*...

 midges in the order Diptera
Diptera
Diptera , or true flies, is the order of insects possessing only a single pair of wings on the mesothorax; the metathorax bears a pair of drumstick like structures called the halteres, the remnants of the hind wings. It is a large order, containing an estimated 240,000 species, although under half...

. The fruit
Fruit
In broad terms, a fruit is a structure of a plant that contains its seeds.The term has different meanings dependent on context. In non-technical usage, such as food preparation, fruit normally means the fleshy seed-associated structures of certain plants that are sweet and edible in the raw state,...

, called a cacao pod, is ovoid, 15–30 cm (6–12 in) long and 8–10 cm (3–4 in) wide, ripening yellow to orange, and weighs about 500 g (1 lb) when ripe. The pod contains 20 to 60 seed
Seed
A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food. It is the product of the ripened ovule of gymnosperm and angiosperm plants which occurs after fertilization and some growth within the mother plant...

s, usually called "beans", embedded in a white pulp. The seeds are the main ingredient of chocolate
Chocolate
Chocolate is a raw or processed food produced from the seed of the tropical Theobroma cacao tree. Cacao has been cultivated for at least three millennia in Mexico, Central and South America. Its earliest documented use is around 1100 BC...

, while the pulp is used in some countries to prepare a refreshing juice
Juice
Juice is the liquid that is naturally contained in fruit or vegetable tissue.Juice is prepared by mechanically squeezing or macerating fruit or vegetable flesh without the application of heat or solvents. For example, orange juice is the liquid extract of the fruit of the orange tree...

. Each seed contains a significant amount of fat (40–50%) as cocoa butter
Cocoa butter
Cocoa butter, also called theobroma oil, is a pale-yellow, pure edible vegetable fat extracted from the cocoa bean. It is used to make chocolate, biscuits, and baked goods, as well as some pharmaceuticals, ointments, and toiletries...

. Their most noted active constituent is theobromine
Theobromine
Theobromine , also known as xantheose, is a bitter alkaloid of the cacao plant, with the chemical formula C7H8N4O2. It is found in chocolate, as well as in a number of other foods, including the leaves of the tea plant, and the kola nut...

, a compound similar to caffeine
Caffeine
Caffeine is a bitter, white crystalline xanthine alkaloid that acts as a stimulant drug. Caffeine is found in varying quantities in the seeds, leaves, and fruit of some plants, where it acts as a natural pesticide that paralyzes and kills certain insects feeding on the plants...

.

The scientific name Theobroma
Theobroma
Theobroma is a genus of flowering plants in the mallow family, Malvaceae, that is sometimes classified as a member of Sterculiaceae. It contains roughly 20 species of small understory trees native to the tropical forests of Central and South America...

from Greek
Greek language
Greek is an independent branch of the Indo-European family of languages. Native to the southern Balkans, it has the longest documented history of any Indo-European language, spanning 34 centuries of written records. Its writing system has been the Greek alphabet for the majority of its history;...

 θεοβρῶμα means "food
Food
Food is any substance consumed to provide nutritional support for the body. It is usually of plant or animal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals...

 of the gods
Deity
A deity is a recognized preternatural or supernatural immortal being, who may be thought of as holy, divine, or sacred, held in high regard, and respected by believers....

". The word cacao itself derives from the Nahuatl
Nahuatl
Nahuatl is thought to mean "a good, clear sound" This language name has several spellings, among them náhuatl , Naoatl, Nauatl, Nahuatl, Nawatl. In a back formation from the name of the language, the ethnic group of Nahuatl speakers are called Nahua...

 (Aztec
Aztec
The Aztec people were certain ethnic groups of central Mexico, particularly those groups who spoke the Nahuatl language and who dominated large parts of Mesoamerica in the 14th, 15th and 16th centuries, a period referred to as the late post-classic period in Mesoamerican chronology.Aztec is the...

 language) word cacahuatl, learned at the time of the conquest when it was first encountered by the Spanish. Similar words for the plant and its by-products are attested in a number of other indigenous Mesoamerican languages
Mesoamerican languages
Mesoamerican languages are the languages indigenous to the Mesoamerican cultural area, which covers southern Mexico, all of Guatemala and Belize and parts of Honduras and El Salvador. The area is characterized by extensive linguistic diversity containing several hundred different languages and...

.

History of cultivation

Cultivation, use, and cultural elaboration of cacao were early and extensive in Mesoamerica
Mesoamerica
Mesoamerica is a region and culture area in the Americas, extending approximately from central Mexico to Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica, within which a number of pre-Columbian societies flourished before the Spanish colonization of the Americas in the 15th and...

. Ceramic vessels with residues from the preparation of cacao beverages have been found at archaeological sites dating back to the Early Formative
Mesoamerican chronology
Mesoamerican chronology divides the history of pre-Columbian Mesoamerica into several periods: the Paleo-Indian , the Archaic , the Preclassic , the Classic , and the Postclassic...

 (1900-900 BC) period. For example, one such vessel found at an Olmec
Olmec
The Olmec were the first major Pre-Columbian civilization in Mexico. They lived in the tropical lowlands of south-central Mexico, in the modern-day states of Veracruz and Tabasco....

 archaeological site on the Gulf Coast of Veracruz
Veracruz
Veracruz, formally Veracruz de Ignacio de la Llave officially Estado Libre y Soberano de Veracruz de Ignacio de la Llave , is one of the 31 states that, along with the Federal District, comprise the 32 federative entities of Mexico. It is divided in 212 municipalities and its capital city is...

, Mexico
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 dates cacao's preparation by pre-Olmec peoples as early as 1750 BC. On the Pacific coast of Chiapas
Chiapas
Chiapas officially Estado Libre y Soberano de Chiapas is one of the 31 states that, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 118 municipalities and its capital city is Tuxtla Gutierrez. Other important cites in Chiapas include San Cristóbal de las...

, Mexico, a Mokaya
Mokaya
Mokaya is the term used to describe pre-Olmec cultures of the Soconusco region in Mexico and parts of the Pacific coast of western Guatemala, an archaeological culture that developed a number of Mesoamerica’s earliest-known sedentary settlements...

 archaeological site provides evidence of cacao beverages dating even earlier, to 1900 BC.

In a study of Theobroma cacao tree genetics by Motamayor et al., they suggest a domestication and spread from lowland Amazonia, contesting an earlier hypothesis that the tree was domesticated independently in both the Lacandon
Lacandon
The Lacandon are one of the Maya peoples who live in the jungles of the Mexican state of Chiapas, near the southern border with Guatemala. Their homeland, the Lacandon Jungle, lies along the Mexican side of the Usumacinta River and its tributaries. The Lacandon are one of the most isolated and...

 area of Mexico, and in Amazonia. The cacao tree belongs to the Theobroma genus, in the Sterculiaceae family, that contains 22 species. Today, the most common of the cultivated species is Theobroma cacao, with two subspecies and three forms. Wild cacaos falling into two groups. The South American subspecies spaerocarpum has a fairly smooth melon-like fruit. In contrast, the Mesoamerican cacao subspecies has ridged, elongated fruits. At some unknown early date, the subspecies T. cacao reached the southern lowlands of Mesoamerica and came into wide usage.
The Maya
Maya civilization
The Maya is a Mesoamerican civilization, noted for the only known fully developed written language of the pre-Columbian Americas, as well as for its art, architecture, and mathematical and astronomical systems. Initially established during the Pre-Classic period The Maya is a Mesoamerican...

 believed that the kakaw (cacao) was discovered by the gods in a mountain that also contained other delectable foods to be used by the Maya. According to Maya mythology
Maya mythology
Mayan mythology is part of Mesoamerican mythology and comprises all of the Mayan tales in which personified forces of nature, deities, and the heroes interacting with these play the main roles...

, the Plumed Serpent gave cacao to the Maya after humans were created from maize by divine grandmother goddess Xmucane (Bogin 1997, Coe 1996, Montejo 1999, Tedlock 1985). The Maya celebrated an annual festival in April to honor their cacao god, Ek Chuah, an event that included the sacrifice of a dog
Dogs in Mesoamerica
Various sorts of dogs are known to have existed in pre-Spanish Mesoamerica, as shown by archaeological and iconographical sources, and the testimonies of the 16th-century Spaniards. In the Central Mexican area, there were three races: the medium-sized furred dog , the medium-sized hairless dog ,...

 with cacao colored markings; additional animal sacrifices; offerings of cacao, feathers and incense; and an exchange of gifts. In a similar creation story, the Mexica
Mexica
The Mexica were a pre-Columbian people of central Mexico.Mexica may also refer to:*Mexica , a board game designed by Wolfgang Kramer and Michael Kiesling*Mexica , a 2005 novel by Norman Spinrad...

 (Aztec
Aztec
The Aztec people were certain ethnic groups of central Mexico, particularly those groups who spoke the Nahuatl language and who dominated large parts of Mesoamerica in the 14th, 15th and 16th centuries, a period referred to as the late post-classic period in Mesoamerican chronology.Aztec is the...

) god Quetzalcoatl
Quetzalcoatl
Quetzalcoatl is a Mesoamerican deity whose name comes from the Nahuatl language and has the meaning of "feathered serpent". The worship of a feathered serpent deity is first documented in Teotihuacan in the first century BCE or first century CE...

 discovered cacao (cacahuatl: "'bitter water"'), in a mountain filled with other plant foods (Coe 1996, Townsend 1992). Cacao was offered regularly to a pantheon of Mexica
Mexica
The Mexica were a pre-Columbian people of central Mexico.Mexica may also refer to:*Mexica , a board game designed by Wolfgang Kramer and Michael Kiesling*Mexica , a 2005 novel by Norman Spinrad...

 deities and the Madrid Codex depicts priests lancing their ear lobes (auto-sacrifice) and covering the cacao with blood as a suitable sacrifice to the gods. The cacao beverage as ritual were used only by men, as it was believed to be toxic for women and children.

There are several mixtures of cacao described in ancient texts, for ceremonial, medicinal uses as well as culinary purposes. Some mixtures included maize
Maize
Maize known in many English-speaking countries as corn or mielie/mealie, is a grain domesticated by indigenous peoples in Mesoamerica in prehistoric times. The leafy stalk produces ears which contain seeds called kernels. Though technically a grain, maize kernels are used in cooking as a vegetable...

, chili
Chile powder
Chili powder is the ground, dried fruit of one or more varieties of chili pepper, sometimes with the addition of other spices . It is used as a spice to add piquance and flavor to dishes. In American English the name is usually spelled "chili", or, less commonly, "chile"...

, vanilla
Vanilla
Vanilla is a flavoring derived from orchids of the genus Vanilla, primarily from the Mexican species, Flat-leaved Vanilla . The word vanilla derives from the Spanish word "", little pod...

 (Vanilla planifolia), and honey. Archaeological evidence for use of cacao, while relatively sparse, has come from the recovery of whole cacao beans at Uaxactun
Uaxactun
Uaxactun is an ancient ruin of the Maya civilization, located in the Petén Basin region of the Maya lowlands, in the present-day department of Petén, Guatemala. The site lies some north of the major center of Tikal...

, Guatemala
Guatemala
Guatemala is a country in Central America bordered by Mexico to the north and west, the Pacific Ocean to the southwest, Belize to the northeast, the Caribbean to the east, and Honduras and El Salvador to the southeast...

 (Kidder 1947) and from the preservation of wood fragments of the cacao tree at Belize
Belize
Belize is a constitutional monarchy and the northernmost country in Central America. Belize has a diverse society, comprising many cultures and languages. Even though Kriol and Spanish are spoken among the population, Belize is the only country in Central America where English is the official...

 sites including Cuello
Cuello
Cuello is a Maya archaeological site in northern Belize. The site is that of a farming village with a long occupational history stretching back to approximately 1200 BC, during the Middle Preclassic period. Its inhabitants lived in pole-and-thatch houses that were built on top of low plaster-coated...

 and Pulltrouser Swamp (Hammond and Miksicek 1981; Turner and Miksicek 1984). In addition, analysis of residues from ceramic vessels has found traces of theobromine
Theobromine
Theobromine , also known as xantheose, is a bitter alkaloid of the cacao plant, with the chemical formula C7H8N4O2. It is found in chocolate, as well as in a number of other foods, including the leaves of the tea plant, and the kola nut...

 and caffeine
Caffeine
Caffeine is a bitter, white crystalline xanthine alkaloid that acts as a stimulant drug. Caffeine is found in varying quantities in the seeds, leaves, and fruit of some plants, where it acts as a natural pesticide that paralyzes and kills certain insects feeding on the plants...

 in early formative vessels from Puerto Escondido, Honduras (1100-900 BC) and in middle formative vessels from Colha, Belize
Colha, Belize
Colha, Belize is a Maya archaeological site located in northern portion of the country, about 52 km. north of Belize City, near the city of Orange Walk. The site is one of the earliest in the Maya region and remains important to the archaeological record of the Maya culture well into the...

 (600-400 BC) using similar techniques to those used to extract chocolate residues from four classic period (ca. 400 AD) vessels from a tomb at the archaeological site of Rio Azul
Río Azul
Rio Azul is a Pre-Columbian archaeological site of the Maya civilization. in Río Azul National Park in present day Petén Department, in northern Guatemala...

. As cacao is the only known commodity from Mesoamerica containing both of these alkaloid
Alkaloid
Alkaloids are a group of naturally occurring chemical compounds that contain mostly basic nitrogen atoms. This group also includes some related compounds with neutral and even weakly acidic properties. Also some synthetic compounds of similar structure are attributed to alkaloids...

 compounds, it seems likely that these vessels were used as containers for cacao drinks. In addition, cacao is named in a hieroglyphic text on one of the Rio Azul
Río Azul
Rio Azul is a Pre-Columbian archaeological site of the Maya civilization. in Río Azul National Park in present day Petén Department, in northern Guatemala...

 vessels. Cacao was also believed to be ground by the Aztecs and mixed with tobacco for smoking purposes.

The first Europeans to encounter cacao were Christopher Columbus
Christopher Columbus
Christopher Columbus was an explorer, colonizer, and navigator, born in the Republic of Genoa, in northwestern Italy. Under the auspices of the Catholic Monarchs of Spain, he completed four voyages across the Atlantic Ocean that led to general European awareness of the American continents in the...

 and his crew in 1502, when they captured a canoe at Guanaja
Guanaja
Guanaja is one of the Bay Islands of Honduras, and is in the Caribbean. It is about 70 km off the north coast of Honduras, and 12 km from the island of Roatan. One of the cays off Guanaja, also called Guanaja or Bonnaca or Low Cay , is near the main island, and contains most of the...

 that contained a quantity of mysterious-looking "almonds." The first real European knowledge about chocolate came in the form of a beverage which was first introduced to the Spanish at their meeting with Moctezuma
Moctezuma II
Moctezuma , also known by a number of variant spellings including Montezuma, Moteuczoma, Motecuhzoma and referred to in full by early Nahuatl texts as Motecuhzoma Xocoyotzin, was the ninth tlatoani or ruler of Tenochtitlan, reigning from 1502 to 1520...

 in the Aztec
Aztec
The Aztec people were certain ethnic groups of central Mexico, particularly those groups who spoke the Nahuatl language and who dominated large parts of Mesoamerica in the 14th, 15th and 16th centuries, a period referred to as the late post-classic period in Mesoamerican chronology.Aztec is the...

 capital of Tenochtitlan in 1519. Cortez and others noted the vast quantities of this beverage that the Aztec
Aztec
The Aztec people were certain ethnic groups of central Mexico, particularly those groups who spoke the Nahuatl language and who dominated large parts of Mesoamerica in the 14th, 15th and 16th centuries, a period referred to as the late post-classic period in Mesoamerican chronology.Aztec is the...

 emperor consumed, and how it was carefully whipped by his attendants beforehand. Examples of cacao beans along with other agricultural products were brought back to Spain at that time, but it seems that the beverage made from cacao was introduced to the Spanish court in 1544 by Kekchi Maya nobles brought from the New World to Spain by Dominican
Dominican Order
The Order of Preachers , after the 15th century more commonly known as the Dominican Order or Dominicans, is a Catholic religious order founded by Saint Dominic and approved by Pope Honorius III on 22 December 1216 in France...

 friar
Friar
A friar is a member of one of the mendicant orders.-Friars and monks:...

s to meet Prince Philip
Philip II of Spain
Philip II was King of Spain, Portugal, Naples, Sicily, and, while married to Mary I, King of England and Ireland. He was lord of the Seventeen Provinces from 1556 until 1581, holding various titles for the individual territories such as duke or count....

 (Coe and Coe 1996). Within a century, the culinary and medical uses of chocolate had spread to France, England and elsewhere in Western Europe
Western Europe
Western Europe is a loose term for the collection of countries in the western most region of the European continents, though this definition is context-dependent and carries cultural and political connotations. One definition describes Western Europe as a geographic entity—the region lying in the...

. Demand for this beverage led the French to establish cacao plantation
Plantation
A plantation is a long artificially established forest, farm or estate, where crops are grown for sale, often in distant markets rather than for local on-site consumption...

s in the Caribbean
Caribbean
The Caribbean is a crescent-shaped group of islands more than 2,000 miles long separating the Gulf of Mexico and the Caribbean Sea, to the west and south, from the Atlantic Ocean, to the east and north...

, while Spain subsequently developed their cacao plantations in their Philippine colony (Bloom 1998, Coe 1996). The Nahuatl-derived Spanish word cacao entered scientific nomenclature in 1753 after the Swedish
Sweden
Sweden , officially the Kingdom of Sweden , is a Nordic country on the Scandinavian Peninsula in Northern Europe. Sweden borders with Norway and Finland and is connected to Denmark by a bridge-tunnel across the Öresund....

 naturalist Linnaeus published his taxonomic binomial system and coined the genus and species Theobroma ("food of the gods") cacao.

Traditional pre-Hispanic beverages made with cacao are still consumed in Mesoamerica
Mesoamerica
Mesoamerica is a region and culture area in the Americas, extending approximately from central Mexico to Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica, within which a number of pre-Columbian societies flourished before the Spanish colonization of the Americas in the 15th and...

. These include the Oaxaca
Oaxaca
Oaxaca , , officially Estado Libre y Soberano de Oaxaca is one of the 31 states which, along with the Federal District, comprise the 32 federative entities of Mexico. It is divided into 571 municipalities; of which 418 are governed by the system of customs and traditions...

n beverage known as tejate
Tejate
Tejate is a maize and cacao beverage traditionally made in Oaxaca, Mexico, originating from pre-Hispanic times. It remains very popular among the indigenous Mixtec and Zapotec peoples, especially in rural areas. It's also very popular for anyone who lives in Oaxaca and the surrounding regions...

.

Currency system

Cacao beans constituted both a ritual beverage and a major currency
Currency
In economics, currency refers to a generally accepted medium of exchange. These are usually the coins and banknotes of a particular government, which comprise the physical aspects of a nation's money supply...

 system in pre-Columbian
Pre-Columbian
The pre-Columbian era incorporates all period subdivisions in the history and prehistory of the Americas before the appearance of significant European influences on the American continents, spanning the time of the original settlement in the Upper Paleolithic period to European colonization during...

 Mesoamerica
Mesoamerica
Mesoamerica is a region and culture area in the Americas, extending approximately from central Mexico to Belize, Guatemala, El Salvador, Honduras, Nicaragua, and Costa Rica, within which a number of pre-Columbian societies flourished before the Spanish colonization of the Americas in the 15th and...

n civilizations. At one point the Aztec
Aztec
The Aztec people were certain ethnic groups of central Mexico, particularly those groups who spoke the Nahuatl language and who dominated large parts of Mesoamerica in the 14th, 15th and 16th centuries, a period referred to as the late post-classic period in Mesoamerican chronology.Aztec is the...

 empire received a yearly tribute of 980 loads (xiquipil in nahuatl) of cacao, in addition to other goods. Each load represented exactly 8,000 beans. The buying power of quality beans was such that 80-100 beans could buy a new cloth mantle. The use of cacao beans as currency is also known to have spawned counterfeiters during the Aztec empire.

In some areas, such as Yucatán
Yucatán
Yucatán officially Estado Libre y Soberano de Yucatán is one of the 31 states which, with the Federal District, comprise the 32 Federal Entities of Mexico. It is divided in 106 municipalities and its capital city is Mérida....

, cacao beans were still used in place of small coins as late as the 1840s.

Cultivation

Cacao is cultivated on roughly 17000000 acre worldwide. According to the Food and Agriculture Organization
Food and Agriculture Organization
The Food and Agriculture Organization of the United Nations is a specialised agency of the United Nations that leads international efforts to defeat hunger. Serving both developed and developing countries, FAO acts as a neutral forum where all nations meet as equals to negotiate agreements and...

 (FAO), the top 20 cacao-producing countries in 2005 were as follows:
Rank, Country Value
(Int'l $1,000*)
Production
(Metric Tons)
1  Côte d'Ivoire 1,024,339 1,330,000
2  Ghana 566,852 736,000
3  Indonesia 469,810 610,000
4  Nigeria 281,886 366,000
5  Brazil 164,644 213,774
6  Cameroon 138,632 180,000
7  Ecuador 105,652 137,178
8  Colombia 42,589 55,298
9  Mexico 37,281 48,405
10  Papua New Guinea 32,733 42,500
11  Malaysia 25,742 33,423
12  Dominican Republic 24,646 32,000
13  Peru 21,950 28,500
14  Venezuela 13,093 17,000
15  Sierra Leone 8,472 11,000
16  Togo 6,547 8,500
17  India 6,161 8,000
18  Philippines 4,352 5,650
19   Congo, Rep.
Republic of the Congo
The Republic of the Congo , sometimes known locally as Congo-Brazzaville, is a state in Central Africa. It is bordered by Gabon, Cameroon, the Central African Republic, the Democratic Republic of the Congo , the Angolan exclave province of Cabinda, and the Gulf of Guinea.The region was dominated by...

4,336 5,630
20  Solomon Islands 3,851 5,000

*Based on 1999–2001 international prices
Cacao production has increased from 1.5 million tons in 1983-1984 to 3.5 million tons in 2003-2004, an increase that has almost entirely been due to the expansion of the production area rather than to yield increases. Cacao is grown both by large agro-industrial plantations and also by small producers.

A tree begins to bear when it is four or five years old. A mature tree may have 6,000 flowers in a year, yet only about 20 pods. About 300-600 seeds (10 pods) are required to produce 1 kg (2.2 lb) of cocoa paste.

There are three main cultivar groups
Cultivar
A cultivar'Cultivar has two meanings as explained under Formal definition. When used in reference to a taxon, the word does not apply to an individual plant but to all those plants sharing the unique characteristics that define the cultivar. is a plant or group of plants selected for desirable...

 of cacao beans used to make cocoa and chocolate. The most prized, rare, and expensive is the Criollo Group, the cocoa bean used by the Maya
Maya civilization
The Maya is a Mesoamerican civilization, noted for the only known fully developed written language of the pre-Columbian Americas, as well as for its art, architecture, and mathematical and astronomical systems. Initially established during the Pre-Classic period The Maya is a Mesoamerican...

. Only 10% of chocolate is made from Criollo, which is less bitter and more aromatic than any other bean. The cacao bean in 80% of chocolate is made using beans of the Forastero Group. Forastero trees are significantly hardier than Criollo trees, resulting in cheaper cacao beans. Trinitario, a hybrid of Criollo and Forastero, is used in about 10% of chocolate.
Major cocoa bean processors include: Hershey's
Hershey's
Hershey's may refer to:* Hershey's, a nickname for The Hershey Company* Hershey's Ice Cream produced by Hershey Creamery Company* Hershey's Chocolate World, a theme-park/visitor-center facility...

, Nestlé
Nestlé
Nestlé S.A. is the world's largest food and nutrition company. Founded and headquartered in Vevey, Switzerland, Nestlé originated in a 1905 merger of the Anglo-Swiss Milk Company, established in 1867 by brothers George Page and Charles Page, and Farine Lactée Henri Nestlé, founded in 1866 by Henri...

 and Mars
Mars, Incorporated
Mars, Incorporated is a worldwide manufacturer of confectionery, pet food, and other food products with US$30 billion in annual sales in 2010, and is ranked as the 5th largest privately held company in the United States by Forbes. Headquartered in McLean, unincorporated Fairfax County, Virginia,...

, all of which purchase cocoa beans via various sources.

In June 2009, Mars Botanicals, a division of Mars, launched Cirku, a cocoa extract product that provides cocoa flavanols made with a patented process that contains a high level of phytonutrients.

Pests

Various plant pests and diseases can cause serious problems for cacao production; see: Illustrated guide to pests and their management.
  • Insect
    Insect
    Insects are a class of living creatures within the arthropods that have a chitinous exoskeleton, a three-part body , three pairs of jointed legs, compound eyes, and two antennae...

    s
    • Cocoa mirids or capsids
      Miridae
      The large and diverse insect family Miridae contains the plant bugs, leaf bugs, and grass bugs, and may also be known as capsid bugs. It is the largest family of true bugs belonging to the suborder Heteroptera, with over 10,000 known species and new ones constantly being described...

       (Worldwide, but especially in West Africa
      West Africa
      West Africa or Western Africa is the westernmost region of the African continent. Geopolitically, the UN definition of Western Africa includes the following 16 countries and an area of approximately 5 million square km:-Flags of West Africa:...

      )
    • Conopomorpha cramerella
      Conopomorpha cramerella
      The Cocoa Pod Borer is a moth of the Gracillariidae family. It is known from Saudi Arabia, China, India , Thailand, Brunei, Indonesia , Malaysia , Vietnam, Australia, New Britain, the Philippines, Samoa, the Solomon Islands, Sri Lanka, Taiwan...

      ("Cocoa pod borer" - in S.E. Asia)
  • Fungi
    Fungus
    A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom, Fungi, which is separate from plants, animals, and bacteria...

    • Moniliophthora roreri
      Moniliophthora roreri
      Moniliophthora roreri is a plant pathogen of Herrania and Theobroma spp., including the cacao tree: on which it is called Frosty pod rot. T. gileri thought to be the indigenous forest host in the upper Amazon basin. Sometimes it is known by the misnomer Monilia...

      ("Frosty Pod Rot")
    • Moniliophthora perniciosa
      Moniliophthora perniciosa
      Moniliophthora perniciosa is a fungus that causes "Witches' Broom Disease" of the cocoa tree...

      ("Witches' Broom")
    • Ceratocystis cacaofunesta ("Mal de machete") or ("Ceratocystis wilt")
    • Verticillium dahliae
      Verticillium dahliae
      Verticillium dahliae is a fungal plant pathogen. It causes verticillium wilt in many plant species, causing leaves to curl and discolor. It may cause death in some plants.- External links :* *...

    • Oncobasidium theobromae
      Oncobasidium theobromae
      Oncobasidium theobromae is a plant pathogen.- External links :* *...

      ("Vascular streak dieback")
  • Oomycetes
    • Phytophthora spp. ("Black Pod") especially Phytophthora megakarya
      Phytophthora megakarya
      Phytophthora megakarya is a plant pathogen that causes black pod disease in cocoa trees.-External links:* *...

       in West Africa
      West Africa
      West Africa or Western Africa is the westernmost region of the African continent. Geopolitically, the UN definition of Western Africa includes the following 16 countries and an area of approximately 5 million square km:-Flags of West Africa:...

  • Virus
    Virus
    A virus is a small infectious agent that can replicate only inside the living cells of organisms. Viruses infect all types of organisms, from animals and plants to bacteria and archaea...

    es
    • CSSV
  • Mistletoe
    Mistletoe
    Mistletoe is the common name for obligate hemi-parasitic plants in several families in the order Santalales. The plants in question grow attached to and within the branches of a tree or shrub.-Mistletoe in the genus Viscum:...

  • Rats
    RATS
    RATS may refer to:* RATS , Regression Analysis of Time Series, a statistical package* Rough Auditing Tool for Security, a computer program...

     and other vertebrate
    Vertebrate
    Vertebrates are animals that are members of the subphylum Vertebrata . Vertebrates are the largest group of chordates, with currently about 58,000 species described. Vertebrates include the jawless fishes, bony fishes, sharks and rays, amphibians, reptiles, mammals, and birds...

     pests (squirrel
    Squirrel
    Squirrels belong to a large family of small or medium-sized rodents called the Sciuridae. The family includes tree squirrels, ground squirrels, chipmunks, marmots , flying squirrels, and prairie dogs. Squirrels are indigenous to the Americas, Eurasia, and Africa and have been introduced to Australia...

    s, woodpecker
    Woodpecker
    Woodpeckers are near passerine birds of the order Piciformes. They are one subfamily in the family Picidae, which also includes the piculets and wrynecks. They are found worldwide and include about 180 species....

    s, etc.)




Cacao genome

In September 2010, a team of scientists announced a draft sequence of the cacao genome (Matina1-6 genotype) Retrieved September 24, 2010. A second unrelated project (The International Cocoa Genome Sequencing Consortium) sequenced a different type of cacao (a criollo type discovered in Belize (B97-61/B2) and a manuscript describing the analysis of this data was published December 2010 in the peer reviewed journal Nature Genetics. The cacao genome is about 430,000,000 bp in length consisting of 10 chromosome and has about 28,798 genes compared to the roughly 30,000 genes of the human genome.[11] About 20% of the cacao genome consists of transposable elements, a low proportion compared to other plant species. The genome sequence will help accelerate research on cacao molecular biology and of breeding for elite varieties of cacao through marker assisted selection.

External links

The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
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