Black Kite
Encyclopedia
The Black Kite is a medium-sized bird of prey
Bird of prey
Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. They are defined as birds that primarily hunt vertebrates, including other birds. Their talons and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....

 in the family Accipitridae, which also includes many other diurnal raptor
Bird of prey
Birds of prey are birds that hunt for food primarily on the wing, using their keen senses, especially vision. They are defined as birds that primarily hunt vertebrates, including other birds. Their talons and beaks tend to be relatively large, powerful and adapted for tearing and/or piercing flesh....

s. Unlike others of the group, they are opportunistic hunters and are more likely to scavenge. They spend a lot of time soaring and gliding in thermals in search of food. Their angled wing and distinctive forked tail make them easy to identify. This kite
Kite (bird)
Kites are raptors with long wings and weak legs which spend a great deal of time soaring. Most feed mainly on carrion but some take various amounts of live prey.They are birds of prey which, along with hawks and eagles, are from the family Accipitridae....

 is widely distributed through the temperate and tropical parts of Eurasia
Eurasia
Eurasia is a continent or supercontinent comprising the traditional continents of Europe and Asia ; covering about 52,990,000 km2 or about 10.6% of the Earth's surface located primarily in the eastern and northern hemispheres...

 and parts of Australasia
Australasia
Australasia is a region of Oceania comprising Australia, New Zealand, the island of New Guinea, and neighbouring islands in the Pacific Ocean. The term was coined by Charles de Brosses in Histoire des navigations aux terres australes...

 and Oceania
Oceania
Oceania is a region centered on the islands of the tropical Pacific Ocean. Conceptions of what constitutes Oceania range from the coral atolls and volcanic islands of the South Pacific to the entire insular region between Asia and the Americas, including Australasia and the Malay Archipelago...

, with the temperate region populations tending to be migratory. Several subspecies are recognized and formerly with their own English names. The European populations are small, but the South Asian population is very large.

Description

The Black Kite can be distinguished from the Red Kite
Red Kite
The Red Kite is a medium-large bird of prey in the family Accipitridae, which also includes many other diurnal raptors such as eagles, buzzards, and harriers. The species is currently endemic to the Western Palearctic region in Europe and northwest Africa, though formerly also occurred just...

 by its slightly smaller size, less forked tail, visible in flight and generally dark plumage
Plumage
Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

 without any rufous. The sexes are alike. The upper plumage is brown but the head and neck tend to be paler. The patch behind the eye appears darker. The outer flight feathers are black and the feathers have dark cross bars and are mottled at the base. The lower parts of the body are pale brown, becoming lighter towards the chin. The body feathers have dark shafts giving it a streaked appearance. The cere and gape are yellow but the bill is black (unlike in the Yellow-billed Kite
Yellow-billed Kite
The Yellow-billed Kite is the Afrotropic counterpart of the Black Kite , of which it is most often considered a subspecies. However, recent DNA studies suggest that the Yellow-billed Kite differs significantly from Black Kites in the Eurasian clade, and should be considered as a separate,...

). The legs are yellow and the claws are black. They have a distinctive shrill whistle followed by a rapid whinnying call.

Systematics and taxonomy

The Red Kite
Red Kite
The Red Kite is a medium-large bird of prey in the family Accipitridae, which also includes many other diurnal raptors such as eagles, buzzards, and harriers. The species is currently endemic to the Western Palearctic region in Europe and northwest Africa, though formerly also occurred just...

 has been known to hybridize with the Black Kite (in captivity where both species were kept together, and in the wild on the Cape Verde Islands).

Recent DNA studies suggest that the yellow-billed, African races, parasitus and aegyptius, differ significantly from Black Kites in the Eurasian clade
Clade
A clade is a group consisting of a species and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological...

, and should be considered as a separate, allopatric species Yellow-billed Kite
Yellow-billed Kite
The Yellow-billed Kite is the Afrotropic counterpart of the Black Kite , of which it is most often considered a subspecies. However, recent DNA studies suggest that the Yellow-billed Kite differs significantly from Black Kites in the Eurasian clade, and should be considered as a separate,...

, M. aegyptius. They occur throughout Africa
Africa
Africa is the world's second largest and second most populous continent, after Asia. At about 30.2 million km² including adjacent islands, it covers 6% of the Earth's total surface area and 20.4% of the total land area...

 except for the Congo
Congo River
The Congo River is a river in Africa, and is the deepest river in the world, with measured depths in excess of . It is the second largest river in the world by volume of water discharged, though it has only one-fifth the volume of the world's largest river, the Amazon...

 basin and the Sahara Desert. There have been some suggestions that the Black-eared Kite (M. m. lineatus) should be elevated to full species status as M. lineatus, but this is not well supported.

Subspecies

  • Milvus migrans migrans (Boddaert
    Pieter Boddaert
    Pieter Boddaert was a Dutch physician and naturalist.Boddaert was the son of a Middelburg jurist and poet by the same name . Pieter Jr. obtained his M.D. at the University of Utrecht in 1764 and there became a lecturer on natural history. Fourteen letters survive of his correspondence with Carl...

    , 1783)
    : European Black Kite
Breeds central, southern and eastern Europe to Tien Shan and south to northwest Pakistan
Pakistan
Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

. Winters in Sub-Saharan Africa
Sub-Saharan Africa
Sub-Saharan Africa as a geographical term refers to the area of the African continent which lies south of the Sahara. A political definition of Sub-Saharan Africa, instead, covers all African countries which are fully or partially located south of the Sahara...

. The head is whitish.
  • Milvus migrans lineatus (J. E. Gray
    John Edward Gray
    John Edward Gray, FRS was a British zoologist. He was the elder brother of George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray ....

    , 1831)
    : Black-eared Kite
Siberia
Siberia
Siberia is an extensive region constituting almost all of Northern Asia. Comprising the central and eastern portion of the Russian Federation, it was part of the Soviet Union from its beginning, as its predecessor states, the Tsardom of Russia and the Russian Empire, conquered it during the 16th...

 to Amurland S around Himalaya to N India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

, N Indochina
Indochina
The Indochinese peninsula, is a region in Southeast Asia. It lies roughly southwest of China, and east of India. The name has its origins in the French, Indochine, as a combination of the names of "China" and "India", and was adopted when French colonizers in Vietnam began expanding their territory...

 and S China
China
Chinese civilization may refer to:* China for more general discussion of the country.* Chinese culture* Greater China, the transnational community of ethnic Chinese.* History of China* Sinosphere, the area historically affected by Chinese culture...

; Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

. Northern inland birds migrate to E Persian Gulf
Persian Gulf
The Persian Gulf, in Southwest Asia, is an extension of the Indian Ocean located between Iran and the Arabian Peninsula.The Persian Gulf was the focus of the 1980–1988 Iran-Iraq War, in which each side attacked the other's oil tankers...

 coast and S Asia in winter. This has a larger pale carpal patch.
  • Milvus migrans govinda (Sykes
    William Henry Sykes
    Colonel William Henry Sykes, FRS was an Indian Army officer, politician and ornithologist.Sykes was born near Bradford in Yorkshire, and joined the Bombay Army, a part of the armed forces of the Honourable East India Company, in 1804, returning to Britain in 1837...

    , 1832)
    : Small Indian Kite (formerly Pariah Kite)
Eastern Pakistan
Pakistan
Pakistan , officially the Islamic Republic of Pakistan is a sovereign state in South Asia. It has a coastline along the Arabian Sea and the Gulf of Oman in the south and is bordered by Afghanistan and Iran in the west, India in the east and China in the far northeast. In the north, Tajikistan...

 east through tropical India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

 and Sri Lanka
Sri Lanka
Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka is a country off the southern coast of the Indian subcontinent. Known until 1972 as Ceylon , Sri Lanka is an island surrounded by the Indian Ocean, the Gulf of Mannar and the Palk Strait, and lies in the vicinity of India and the...

 to Indochina and Malay Peninsula
Malay Peninsula
The Malay Peninsula or Thai-Malay Peninsula is a peninsula in Southeast Asia. The land mass runs approximately north-south and, at its terminus, is the southern-most point of the Asian mainland...

. Resident. A dark brown kite found throughout the subcontinent. Can be seen circling and soaring in urban areas. Easily distinguished by the shallow forked tail. The name Pariah originates from the Indian caste system and usage of this name is deprecated.
  • Milvus migrans affinis (Gould
    John Gould
    John Gould was an English ornithologist and bird artist. The Gould League in Australia was named after him. His identification of the birds now nicknamed "Darwin's finches" played a role in the inception of Darwin's theory of evolution by natural selection...

    , 1838)
    : Fork-tailed Kite
Sulawesi
Sulawesi
Sulawesi is one of the four larger Sunda Islands of Indonesia and is situated between Borneo and the Maluku Islands. In Indonesia, only Sumatra, Borneo, and Papua are larger in territory, and only Java and Sumatra have larger Indonesian populations.- Etymology :The Portuguese were the first to...

 and possibly Lesser Sunda Islands
Lesser Sunda Islands
The Lesser Sunda Islands or Nusa Tenggara are a group of islands in the southern Maritime Southeast Asia, north of Australia. Together with the Greater Sunda Islands to the west they make up the Sunda Islands...

; Papua New Guinea
Papua New Guinea
Papua New Guinea , officially the Independent State of Papua New Guinea, is a country in Oceania, occupying the eastern half of the island of New Guinea and numerous offshore islands...

 except mountains; NE and E Australia.
  • Milvus migrans formosanus (Kuroda
    Nagamichi Kuroda
    was a Japanese ornithologist. His works included Birds of the Island of Java and Parrots of the World in Life Colours . He described the Crested Shelduck in 1917....

    , 1920)
    : Taiwan Kite
Taiwan
Taiwan
Taiwan , also known, especially in the past, as Formosa , is the largest island of the same-named island group of East Asia in the western Pacific Ocean and located off the southeastern coast of mainland China. The island forms over 99% of the current territory of the Republic of China following...

 and Hainan
Hainan
Hainan is the smallest province of the People's Republic of China . Although the province comprises some two hundred islands scattered among three archipelagos off the southern coast, of its land mass is Hainan Island , from which the province takes its name...

; resident.

Distribution

The species is found in Europe, Asia, Africa and Australia. The temperate populations of this kite tend to be migratory while the tropical ones are resident. Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

an and central Asia
Asia
Asia is the world's largest and most populous continent, located primarily in the eastern and northern hemispheres. It covers 8.7% of the Earth's total surface area and with approximately 3.879 billion people, it hosts 60% of the world's current human population...

n birds (subspecies
Subspecies
Subspecies in biological classification, is either a taxonomic rank subordinate to species, ora taxonomic unit in that rank . A subspecies cannot be recognized in isolation: a species will either be recognized as having no subspecies at all or two or more, never just one...

 M. m. milvus and Black-eared Kite M. m. lineatus, respectively) are migratory
Bird migration
Bird migration is the regular seasonal journey undertaken by many species of birds. Bird movements include those made in response to changes in food availability, habitat or weather. Sometimes, journeys are not termed "true migration" because they are irregular or in only one direction...

, moving to the tropics in winter, but races in warmer regions such as the India
India
India , officially the Republic of India , is a country in South Asia. It is the seventh-largest country by geographical area, the second-most populous country with over 1.2 billion people, and the most populous democracy in the world...

n M. m. govinda (Pariah Kite), or the Australasian M. m. affinis (Fork-tailed Kite), are resident. In some areas such as in the United Kingdom
United Kingdom
The United Kingdom of Great Britain and Northern IrelandIn the United Kingdom and Dependencies, other languages have been officially recognised as legitimate autochthonous languages under the European Charter for Regional or Minority Languages...

, the Black Kite occurs only as a wanderer on migration. These birds are usually of the nominate race, but in November 2006 a juvenile of the eastern lineatus, not previously recorded in western Europe, was found in Lincolnshire
Lincolnshire
Lincolnshire is a county in the east of England. It borders Norfolk to the south east, Cambridgeshire to the south, Rutland to the south west, Leicestershire and Nottinghamshire to the west, South Yorkshire to the north west, and the East Riding of Yorkshire to the north. It also borders...

.

The species is not found in the Indonesia
Indonesia
Indonesia , officially the Republic of Indonesia , is a country in Southeast Asia and Oceania. Indonesia is an archipelago comprising approximately 13,000 islands. It has 33 provinces with over 238 million people, and is the world's fourth most populous country. Indonesia is a republic, with an...

n archipelago
Archipelago
An archipelago , sometimes called an island group, is a chain or cluster of islands. The word archipelago is derived from the Greek ἄρχι- – arkhi- and πέλαγος – pélagos through the Italian arcipelago...

 between the South East Asian mainland and the Wallace Line
Wallace Line
The Wallace Line separates the ecozones of Asia and Wallacea, a transitional zone between Asia and Australia. West of the line are found organisms related to Asiatic species; to the east, a mixture of species of Asian and Australian origin is present...

. Vagrants, most likely of the Black-eared Kite, on occasion range far into the Pacific, out to the Hawaiian islands
Hawaiian Islands
The Hawaiian Islands are an archipelago of eight major islands, several atolls, numerous smaller islets, and undersea seamounts in the North Pacific Ocean, extending some 1,500 miles from the island of Hawaii in the south to northernmost Kure Atoll...

.
In India, the population of M. m. govinda is particularly large especially in areas of high human population. Here the birds avoid heavily forested regions. A survey in 1967 in the 150 square kilometres of the city of New Delhi produced an estimate of about 2200 pairs or roughly 15 per square kilometer.

Food and foraging

Black Kites are most often seen gliding and soaring on thermals as they search for food. The flight is buoyant and the bird glides with ease, changing directions easily. They will swoop down with their legs lowered to snatch small live prey, fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

, household refuse and carrion
Carrion
Carrion refers to the carcass of a dead animal. Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters include vultures, hawks, eagles, hyenas, Virginia Opossum, Tasmanian Devils, coyotes, Komodo dragons, and burying beetles...

, for which behaviour they are known in British military slang
Military slang
Military slang is an array of colloquial terminology used commonly by military personnel, including slang which is unique to or originates with the armed forces. It often takes the form of abbreviations/acronyms or derivations of the NATO Phonetic Alphabet, or otherwise incorporates aspects of...

 as the shite-hawk
Shite-hawk
Shite-hawk or shit-hawk or shitty hawk is a slang name applied to various birds of prey that exhibit scavenging behaviour, originally and primarily the Black Kite, although the term has also been applied to other birds such as the Herring Gull...

. They are opportunist hunters and have been known to take birds, bats and rodents. They are attracted to smoke and fires, where they seek escaping prey. This behaviour has led to Australian native beliefs that kites spread fires by picking up burning twigs and dropping them on dry grass. The Indian populations are well adapted to living in cities and are found in densely populated areas. Large numbers may be seen soaring in thermal
Thermal
A thermal column is a column of rising air in the lower altitudes of the Earth's atmosphere. Thermals are created by the uneven heating of the Earth's surface from solar radiation, and are an example of convection. The sun warms the ground, which in turn warms the air directly above it...

s over cities. In some places they will readily swoop and snatch food held by humans. Black Kites in Spain prey on nestling waterfowl especially during summer to feed their young. Predation of nests of other pairs of Black Kites has also been noted. Kites have also been seen to tear and carry away the nests of Baya Weaver
Baya Weaver
The Baya Weaver is a weaverbird found across South and Southeast Asia. Flocks of these birds are found in grasslands, cultivated areas, scrub and secondary growth and they are best known for their hanging retort shaped nests woven from leaves...

s in an attempt to obtain eggs or chicks.

Flocking and roosting

In winter, kites form large communal roosts. Flocks may fly about before settling at the roost. When migrating, the Black Kite has a greater propensity to form large flocks than other migratory raptors, particularly prior to making a crossing across water. In India, the subspecies govinda shows large seasonal fluctuations with the highest numbers seen from July to October, after the Monsoons, and it has been suggested that they make local movements in response to high rainfall.

Breeding

The breeding season of Black Kites in India begins in winter, the young birds fledging before the monsoon
Monsoon
Monsoon is traditionally defined as a seasonal reversing wind accompanied by corresponding changes in precipitation, but is now used to describe seasonal changes in atmospheric circulation and precipitation associated with the asymmetric heating of land and sea...

s. The nest is a rough platform of twigs and rags placed in a tree. Nest sites may be reused in subsequent years. European birds breed in summer. Birds in the Italian Alps tended to build their nest close to water in steep cliffs or tall trees. Nest orientation may be related to wind and rainfall. The nests may sometimes be decorated with bright materials such as white plastic and a study in Spain suggests that they may have a role in signalling to keep away other kites. After pairing, the male frequently copulates with the female. Unguarded females may be approached by other males, and extra pair copulations are frequent. Males returning from a foraging trip will frequently copulate on return, as this increases the chances of his sperm fertilizing the eggs rather than a different male. Both the male and female take part in nest building, incubation and care of chicks. The typical clutch size is 2 or sometimes 3 eggs. The incubation period varies from 30–34 days. Chicks of the Indian population stayed at the nest for nearly two months. Chicks hatched later in European populations appeared to fledge faster. The care of young by the parents also rapidly decreased with the need for adults to migrate. Siblings show aggression to each other and often the weaker chick may be killed, but parent birds were found to preferentially feed the smaller chicks in experimentally altered nests. Newly hatched young have down (prepennae) which are sepiaon the back and black around the eye and buff on the head, neck and underparts. This is replaced by brownish gray second down (preplumulae). After 9 to 12 days the second down appears on the whole body except the top of the head. Body feathers begin to appear after 18 to 22 days. The feathers on the head become noticeable from the 24th to 29th day. The nestlings initially feed on food fallen at the bottom of the nest and begin to tear flesh after 33–39 days. They are able to stand on their legs after 17–19 days and begin flapping their wings after 27–31 days. After 50 days they begin to move to branches next to the nest. Birds are able to breed after their second year. Parent birds guard their nest and will dive aggressively at intruders. Humans who intrude the nest appear to be recognized by birds and singled out for dive attacks.

Mortality factors

Black-eared Kites in Japan were found to accumulate nearly 70% of mercury accumulated from polluted food in the feathers, thus excreting it in the moult process. Black Kites often perch on electric wires and are frequent victims of electrocution. Their habit of swooping to pick up dead rodent
Rodent
Rodentia is an order of mammals also known as rodents, characterised by two continuously growing incisors in the upper and lower jaws which must be kept short by gnawing....

s or other roadkill
Roadkill
Roadkill is an animal or animals that have been struck and killed by motor vehicles. In the United States of America, removal and disposal of animals struck by motor vehicles is usually the responsibility of the state's state trooper association or department of transportation.-History:During the...

 leads to collisions with vehicles. Instances of mass poisoning as a result of feeding on poisoned voles in agricultural fields have been noted. They are also a major nuisance at some airport
Airport
An airport is a location where aircraft such as fixed-wing aircraft, helicopters, and blimps take off and land. Aircraft may be stored or maintained at an airport...

s, where their size makes them a significant birdstrike hazard.

Like most bird species, they have parasites, several species of endoparasitic trematodes are known and some Digenea
Digenea
Digenea is a subclass within the Platyhelminthes consisting of parasitic flatworms with a syncytial tegument and, usually, two suckers, one ventral and one oral. Adults are particularly common in the digestive tract, but occur throughout the organ systems of all classes of vertebrates...

 species that are transmitted via fishes.

Birds with abnormal development of a secondary upper mandible have been recorded in govinda and lineatus.

Other sources

  • Desai, J. H. & A. K. Malhotra (1982) Annual gonadal cycle of Black Kite Milvus migrans govinda. J. Yamashina Inst. Orn. 14: 143-150.
  • Hardy, J. 1985. Black Kite capturing small passerines. Australasian Raptor Association News 6:14.
  • American Ornithologists' Union
    American Ornithologists' Union
    The American Ornithologists' Union is an ornithological organization in the USA. Unlike the National Audubon Society, its members are primarily professional ornithologists rather than amateur birders...

     (AOU) (2000): Forty-second supplement to the American Ornithologists' Union Check-list of North American Birds. Auk 117(3): 847–858. DOI
    Digital object identifier
    A digital object identifier is a character string used to uniquely identify an object such as an electronic document. Metadata about the object is stored in association with the DOI name and this metadata may include a location, such as a URL, where the object can be found...

    : 10.1642/0004-8038(2000)117[0847:FSSTTA]2.0.CO;2

External links

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