Black-footed Albatross
Encyclopedia
The Black-footed Albatross, Phoebastria nigripes, is a large seabird
Seabird
Seabirds are birds that have adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same environmental problems and feeding niches have resulted in similar adaptations...

 from the North Pacific of the albatross family Diomedeidae. It is one of three species of albatross that range in the northern hemisphere
Northern Hemisphere
The Northern Hemisphere is the half of a planet that is north of its equator—the word hemisphere literally means “half sphere”. It is also that half of the celestial sphere north of the celestial equator...

, nesting on isolated tropical islands. Unlike many albatrosses, it is dark plumaged.

Taxonomy

Black-footed Albatrosses are a type of Albatross that belong to Diomedeidae family and come from the Procellariiformes
Procellariiformes
Procellariiformes is an order of seabirds that comprises four families: the albatrosses, petrels and shearwaters, storm petrels, and diving petrels...

 order, along with Shearwater
Shearwater
Shearwaters are medium-sized long-winged seabirds. There are more than 30 species of shearwaters, a few larger ones in the genus Calonectris and many smaller species in the genus Puffinus...

s, Fulmar
Fulmar
Fulmars are seabirds of the family Procellariidae. The family consists of two extant species and two that are extinct.-Taxonomy:As members of Procellaridae and then the order Procellariiformes, they share certain traits. First, they have nasal passages that attach to the upper bill called...

s, Storm-petrel
Storm-petrel
Storm petrels are seabirds in the family Hydrobatidae, part of the order Procellariiformes. These smallest of seabirds feed on planktonic crustaceans and small fish picked from the surface, typically while hovering. The flight is fluttering and sometimes bat-like.Storm petrels have a cosmopolitan...

s, and Diving-petrels. They share certain identifying features. First, they have nasal passages that attach to the upper bill called naricorns. Although the nostrils on the Albatross are on the sides of the bill. The bills of Procellariiformes are also unique in that they are split into between 7 and 9 horny plates. Finally, they produce a stomach oil made up of wax ester
Wax ester
An ester of a fatty acid and a long-chain alcohol. These oils have similar properties as triglycerides, but are indigestible. They are found in some fish such as orange roughy, oilfish, escolar, black oreo, smooth oreo and other deep water fish. They are also present in marine copepods. Wax...

s and triglycerides that is stored in the proventriculus
Proventriculus
The proventriculus is part of the digestive system of birds, invertebrates and insects.-Birds:The proventriculus is a standard part of avian anatomy...

. This is used against predators as well as an energy rich food source for chicks and for the adults during their long flights. They also have a salt gland that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nose.

The specific epithet of nigripes is derived from two Latin
Latin
Latin is an Italic language originally spoken in Latium and Ancient Rome. It, along with most European languages, is a descendant of the ancient Proto-Indo-European language. Although it is considered a dead language, a number of scholars and members of the Christian clergy speak it fluently, and...

 words, niger means black, and pes means foot.

Description

The Black-footed Albatross is a small member of the family
Family (biology)
In biological classification, family is* a taxonomic rank. Other well-known ranks are life, domain, kingdom, phylum, class, order, genus, and species, with family fitting between order and genus. As for the other well-known ranks, there is the option of an immediately lower rank, indicated by the...

 (while still large compared to most other seabirds) that has almost all black plumage
Plumage
Plumage refers both to the layer of feathers that cover a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage vary between species and subspecies and can also vary between different age classes, sexes, and season. Within species there can also be a...

. Some adults show white undertail coverts, and all adults have white markings around the base of the beak
Beak
The beak, bill or rostrum is an external anatomical structure of birds which is used for eating and for grooming, manipulating objects, killing prey, fighting, probing for food, courtship and feeding young...

 and below the eye
Eye
Eyes are organs that detect light and convert it into electro-chemical impulses in neurons. The simplest photoreceptors in conscious vision connect light to movement...

. As the birds age they acquire more white at the base of the beak. Its beak and feet are also all dark. They have only the one plumage. They measure 71 to 91 cm (28 to 35.8 in) in length, have a wingspan of 190 to 220 cm (6.2 to 7.2 ft), and weigh 2.6–4.3 kg (5.7–9.5 lb).. Males, at an average weight of 3.4 kg (7.5 lb) are larger than females, at an average of 3 kg (6.6 lb).

Distribution and habitat

The Black-footed Albatross, along with the Laysan Albatross
Laysan Albatross
The Laysan Albatross, Phoebastria immutabilis, is a large seabird that ranges across the North Pacific. This small two-tone gull-like albatross is the second most common seabird in the Hawaiian Islands, with an estimated population of 2.5 million birds, and is currently expanding its range to new...

 and the rare Short-tailed Albatross
Short-tailed Albatross
The Short-tailed Albatross or Steller's Albatross, Phoebastria albatrus, is a large rare seabird from the North Pacific. Although related to the other North Pacific albatrosses, it also exhibits behavioural and morphological links to the albatrosses of the Southern Ocean...

, are the three species
Species
In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are...

 of albatross that range in the northern hemisphere, as opposed to the rest of the family which range from the Equator
Equator
An equator is the intersection of a sphere's surface with the plane perpendicular to the sphere's axis of rotation and containing the sphere's center of mass....

 south. There are at least 12 known breeding locations. They nest colonially on isolated islands of the Northwestern Hawaiian Islands
Northwestern Hawaiian Islands
The Northwestern Hawaiian Islands or the Leeward Islands are the small islands and atolls in the Hawaiian island chain located northwest of the islands of Kauai and Niihau. They are administered by the U.S. state of Hawaii except Midway Atoll, which has temporary residential facilities and is...

, from Kure Atoll
Kure Atoll
Kure Atoll or Ocean Island is an atoll in the Pacific Ocean beyond Midway Atoll in the Northwestern Hawaiian Islands at . The only land of significant size is called Green Island and is habitat for hundreds of thousands of seabirds...

 to Kaula Island, (such as Laysan
Laysan
Laysan , located northwest of Honolulu at N25° 42' 14" W171° 44' 04", is one of the Northwestern Hawaiian Islands. It comprises one land mass of , about 1 by 1.5 miles in size . It is an atoll of sorts, although the land completely surrounds a shallow central lake some above sea level that has...

, Midway
Midway Atoll
Midway Atoll is a atoll in the North Pacific Ocean, near the northwestern end of the Hawaiian archipelago, about one-third of the way between Honolulu, Hawaii, and Tokyo, Japan. Unique among the Hawaiian islands, Midway observes UTC-11 , eleven hours behind Coordinated Universal Time and one hour...

, and the French Frigate Shoals
French Frigate Shoals
The French Frigate Shoals is the largest atoll in the Northwestern Hawaiian Islands. Its name commemorates French explorer Jean-François de La Pérouse, who nearly lost two frigates when attempting to navigate the shoals...

), the Japan
Japan
Japan is an island nation in East Asia. Located in the Pacific Ocean, it lies to the east of the Sea of Japan, China, North Korea, South Korea and Russia, stretching from the Sea of Okhotsk in the north to the East China Sea and Taiwan in the south...

ese islands of Tori Shima
Torishima (Izu Islands)
, literally meaning "Bird Island", is an uninhabited volcanic island at the south end of the Izu Islands in the Pacific Ocean, administered by Japan.-Geography:...

, Bonin, and Senkaku
Senkaku Islands
The , also known as the Diaoyu Islands or Diaoyutai Islands or the Pinnacle Islands, are a group of disputed uninhabited islands in the East China Sea...

, and off the Mexican
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 coast, primarily on Isla Guadalupe
Guadalupe Island
Guadalupe Island, or Isla Guadalupe is a volcanic island located 241 kilometers off the west coast of Mexico's Baja California peninsula and some 400 kilometers southwest of the city of Ensenada in Baja California state, in the Pacific Ocean...

. They are extirpated from the Iwo Jima
Iwo Jima
Iwo Jima, officially , is an island of the Japanese Volcano Islands chain, which lie south of the Ogasawara Islands and together with them form the Ogasawara Archipelago. The island is located south of mainland Tokyo and administered as part of Ogasawara, one of eight villages of Tokyo...

, Agrihan
Agrihan
Agrihan is a stratovolcano which forms an island in the Northern Mariana Islands in the Pacific Ocean. The entire island is a massive volcano which rises over from the ocean floor, and is the fifth largest in the Marianas volcanic arc. At , its summit is the highest point in Micronesia...

, Taongi Atoll, Marcus Island, Wake Island
Wake Island
Wake Island is a coral atoll having a coastline of in the North Pacific Ocean, located about two-thirds of the way from Honolulu west to Guam east. It is an unorganized, unincorporated territory of the United States, administered by the Office of Insular Affairs, U.S. Department of the Interior...

, and Johnston Island. Their range at sea
Sea
A sea generally refers to a large body of salt water, but the term is used in other contexts as well. Most commonly, it means a large expanse of saline water connected with an ocean, and is commonly used as a synonym for ocean...

 varies during the seasons (straying farther from the breeding islands when the chicks are older or they don't have chicks) but they make use of great areas of the North Pacific, feeding from Alaska
Alaska
Alaska is the largest state in the United States by area. It is situated in the northwest extremity of the North American continent, with Canada to the east, the Arctic Ocean to the north, and the Pacific Ocean to the west and south, with Russia further west across the Bering Strait...

 to California
California
California is a state located on the West Coast of the United States. It is by far the most populous U.S. state, and the third-largest by land area...

 and Japan; however they do prefer the northeastern Pacific Ocean. They overlap greatly in breeding and feeding range with the other two species of northern albatross, although the other two will range further north into the Bering Sea
Bering Sea
The Bering Sea is a marginal sea of the Pacific Ocean. It comprises a deep water basin, which then rises through a narrow slope into the shallower water above the continental shelves....

 than the Black-footed will. They have, on occasion, been sighted in the southern hemisphere.

Behavior

Their vocalization will range from shrieks and squeals whilst fighting over food to bill-clapping, whistles, groans, and quacks while courting.

Breeding

The Black-footed Albatross, like the rest of its family, forms long term pair-bonds that last for life. After fledging
Fledge
Fledge is the stage in a young bird's life when the feathers and wing muscles are sufficiently developed for flight. It also describes the act of a chick's parents raising it to a fully grown state...

 the birds return to the colony after three years, and spend two years building nests
Bird nest
A bird nest is the spot in which a bird lays and incubates its eggs and raises its young. Although the term popularly refers to a specific structure made by the bird itself—such as the grassy cup nest of the American Robin or Eurasian Blackbird, or the elaborately woven hanging nest of the...

, dancing and being with prospective mates, a behaviour that probably evolved to ensure maximum trust between the birds (raising an albatross chick is a massive energetic investment, and a long courting period establishes for both birds that the other is committed). They will start reproducing after about seven years.

Nests are simple depressions scraped in the sand
Sand
Sand is a naturally occurring granular material composed of finely divided rock and mineral particles.The composition of sand is highly variable, depending on the local rock sources and conditions, but the most common constituent of sand in inland continental settings and non-tropical coastal...

, into which one dull white with reddish brown spots egg
Egg (biology)
An egg is an organic vessel in which an embryo first begins to develop. In most birds, reptiles, insects, molluscs, fish, and monotremes, an egg is the zygote, resulting from fertilization of the ovum, which is expelled from the body and permitted to develop outside the body until the developing...

 is laid. The egg is incubated for just over two months (65 days). Both birds incubate the egg, the male incubating more as the female leaves soon after hatching to recoup reserves used for egg-laying. The average time spent on incubating shifts is 18 days. However, mates can wait up to 38 days to be relieved, and if something happens to the mate the other has been recorded incubating for 49 days without food or water.

The chick is brooded for 20 days by its parents, after which both parents leave the nest and return to feed the chick. The chick is fed regurgitated food by sticking its bill inside that of its parent. Fledging
Fledge
Fledge is the stage in a young bird's life when the feathers and wing muscles are sufficiently developed for flight. It also describes the act of a chick's parents raising it to a fully grown state...

 occurs after 140 days.

Feeding

The Black-footed Albatross feeds in pelagic waters, taking fish
Fish
Fish are a paraphyletic group of organisms that consist of all gill-bearing aquatic vertebrate animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish, as well as various extinct related groups...

, mostly the eggs of flying fish, squid
Squid
Squid are cephalopods of the order Teuthida, which comprises around 300 species. Like all other cephalopods, squid have a distinct head, bilateral symmetry, a mantle, and arms. Squid, like cuttlefish, have eight arms arranged in pairs and two, usually longer, tentacles...

 and to a lesser extent crustacean
Crustacean
Crustaceans form a very large group of arthropods, usually treated as a subphylum, which includes such familiar animals as crabs, lobsters, crayfish, shrimp, krill and barnacles. The 50,000 described species range in size from Stygotantulus stocki at , to the Japanese spider crab with a leg span...

s. It will also consume floating debris
Debris
Debris is rubble, wreckage, ruins, litter and discarded garbage/refuse/trash, scattered remains of something destroyed, or, in geology, large rock fragments left by a melting glacier etc. The singular form of debris is debris...

, including plastics. It has been described as a 'floating pig' by one author for its habit of taking kitchen scraps from ships.

Conservation

Breeding Population and Trends
Breeding Location Breeding Pair Trend
Midway Atoll
Midway Atoll
Midway Atoll is a atoll in the North Pacific Ocean, near the northwestern end of the Hawaiian archipelago, about one-third of the way between Honolulu, Hawaii, and Tokyo, Japan. Unique among the Hawaiian islands, Midway observes UTC-11 , eleven hours behind Coordinated Universal Time and one hour...

 
24,000 pair -9.6% from 1992 to 2001
Laysan Island  21,000 -9.6% from 1992 to 2001
French Frigate Shoals
French Frigate Shoals
The French Frigate Shoals is the largest atoll in the Northwestern Hawaiian Islands. Its name commemorates French explorer Jean-François de La Pérouse, who nearly lost two frigates when attempting to navigate the shoals...

 
Torishima  1,218 Unknown
Bonin Islands  23 Unknown
Islas Guadalupe
Guadalupe Island
Guadalupe Island, or Isla Guadalupe is a volcanic island located 241 kilometers off the west coast of Mexico's Baja California peninsula and some 400 kilometers southwest of the city of Ensenada in Baja California state, in the Pacific Ocean...

 
337 Unknown
Other offshore Mexican
Mexico
The United Mexican States , commonly known as Mexico , is a federal constitutional republic in North America. It is bordered on the north by the United States; on the south and west by the Pacific Ocean; on the southeast by Guatemala, Belize, and the Caribbean Sea; and on the east by the Gulf of...

 islands
63 unknown
Total 64,500 -60% over 56 years

The Black-footed Albatross is considered endangered
Endangered species
An endangered species is a population of organisms which is at risk of becoming extinct because it is either few in numbers, or threatened by changing environmental or predation parameters...

 by the IUCN, because it is taken incidentally by longline fishing. An estimated 4,000 are taken every year, based on the number taken in 1990; other estimates put the number at 8,000, although more recent numbers are at around 6,150 per year with the majority of deaths from Taiwanese and Japanese fishing fleets. It is also vulnerable to oil and ingestion of floating plastics, which reduces the space in the stomach
Stomach
The stomach is a muscular, hollow, dilated part of the alimentary canal which functions as an important organ of the digestive tract in some animals, including vertebrates, echinoderms, insects , and molluscs. It is involved in the second phase of digestion, following mastication .The stomach is...

 available for food to be brought to the chick. Finally volcanic eruptions on Torishima continues to be a threat.

The Black-footed Albatross has an occurrence range of 37600000 km² (14,517,441.2 sq mi) and a breeding range of 28 km² (10.8 sq mi), with a population of 129,000 adult birds. Of these birds 24,000 pair breed on Midway Atoll
Midway Atoll
Midway Atoll is a atoll in the North Pacific Ocean, near the northwestern end of the Hawaiian archipelago, about one-third of the way between Honolulu, Hawaii, and Tokyo, Japan. Unique among the Hawaiian islands, Midway observes UTC-11 , eleven hours behind Coordinated Universal Time and one hour...

 and 21,000 pair breed on Laysan Island. Torishima has 1,218 pair, the Bonin Islands have 23 pair and there are about 400 pair on offshore Mexican islands with 337 on Islas Guadalupe
Guadalupe Island
Guadalupe Island, or Isla Guadalupe is a volcanic island located 241 kilometers off the west coast of Mexico's Baja California peninsula and some 400 kilometers southwest of the city of Ensenada in Baja California state, in the Pacific Ocean...

. All of these numbers come from estimates from 2005 to 2007.

All of its nesting sites in the U.S. are protected, along with a 50 nmi (92.6 km) buffer zone around these islands. Within this buffer zone longline fishing is outlawed. Almost 80% of the breeding population is counted or sampled each year and most fisheries utilize seabird bycatch prevention measures.

External links

The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
x
OK