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Barbule
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Barbules are a part of the tree formed by feathers : the trunk, or axis, being the rachis and the barbs the main boughs, barbules are the secondary branches.

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Encyclopedia
Barbules are a part of the tree formed by feathers : the trunk, or axis, being the rachis and the barbs the main boughs, barbules are the secondary branches.
Barbules have minute hooks called barbicels for cross-attachment.
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