Aspen parkland
Encyclopedia
Aspen parkland refers to a very large area of transitional biome between prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 and boreal forest in two sections; the Peace River Country
Peace River Country
The Peace River Country is an aspen parkland region around the Peace River in Canada. It spans from northwestern Alberta to the Rocky Mountains in northeastern British Columbia, where the region is also referred to as the Peace River Block.- Geography :The Peace River Country includes the...

 of northwestern Alberta
Alberta
Alberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...

 crossing the border into British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

, and a much larger area stretching from central Alberta
Alberta
Alberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...

, all across central Saskatchewan
Saskatchewan
Saskatchewan is a prairie province in Canada, which has an area of . Saskatchewan is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, and on the south by the U.S. states of Montana and North Dakota....

 to south central Manitoba
Manitoba
Manitoba is a Canadian prairie province with an area of . The province has over 110,000 lakes and has a largely continental climate because of its flat topography. Agriculture, mostly concentrated in the fertile southern and western parts of the province, is vital to the province's economy; other...

 near the United States
United States
The United States of America is a federal constitutional republic comprising fifty states and a federal district...

 border. Aspen parkland consists of groves of aspen
Aspen
Populus section Populus, of the Populus genus, includes the aspen trees and the white poplar Populus alba. The five typical aspens are all native to cold regions with cool summers, in the north of the Northern Hemisphere, extending south at high altitudes in the mountains. The White Poplar, by...

 poplars and spruce interspersed with areas of prairie grasslands, also intersected by large stream and river valleys lined with aspen-spruce forests and dense shrubbery. This is the largest boreal-grassland transition zone in the world and is a zone of constant competition and tension as prairie and woodlands struggle to overtake each other within the parkland.

This article focuses on this biome in North America. Similar biomes also exist in Russia
Russia
Russia or , officially known as both Russia and the Russian Federation , is a country in northern Eurasia. It is a federal semi-presidential republic, comprising 83 federal subjects...

 north of the steppes (forest steppe
Forest steppe
A forest steppe is a temperate-climate ecotone and habitat type composed of grassland interspersed with areas of woodland or forest.-Locations:...

) and in northern Europe
Europe
Europe is, by convention, one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally 'divided' from Asia to its east by the watershed divides of the Ural and Caucasus Mountains, the Ural River, the Caspian and Black Seas, and the waterways connecting...

.

Setting

The aspen parkland biome runs in a thin band no wider than 500 km through the Prairie Provinces, although it gets broader to the west, especially in Alberta
Alberta
Alberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...

. This is a hilly landscape with many small lakes and ponds. The cities of Edmonton
Edmonton
Edmonton is the capital of the Canadian province of Alberta and is the province's second-largest city. Edmonton is located on the North Saskatchewan River and is the centre of the Edmonton Capital Region, which is surrounded by the central region of the province.The city and its census...

 and Saskatoon
Saskatoon
Saskatoon is a city in central Saskatchewan, Canada, on the South Saskatchewan River. Residents of the city of Saskatoon are called Saskatonians. The city is surrounded by the Rural Municipality of Corman Park No. 344....

 are the largest cities completely in this biome while Winnipeg is bordered by tallgrass prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 to the west and south and the aspen parkland to the northeast, and Calgary
Calgary
Calgary is a city in the Province of Alberta, Canada. It is located in the south of the province, in an area of foothills and prairie, approximately east of the front ranges of the Canadian Rockies...

 is bordered by prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 to the east and the Foothills Parkland to the west.

There are three main sections of aspen parkland: Peace River, Central, and Foothills. The Central Parkland is the largest section and is part of main band of aspen parkland extending across Alberta
Alberta
Alberta is a province of Canada. It had an estimated population of 3.7 million in 2010 making it the most populous of Canada's three prairie provinces...

, Saskatchewan
Saskatchewan
Saskatchewan is a prairie province in Canada, which has an area of . Saskatchewan is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, and on the south by the U.S. states of Montana and North Dakota....

 and Manitoba
Manitoba
Manitoba is a Canadian prairie province with an area of . The province has over 110,000 lakes and has a largely continental climate because of its flat topography. Agriculture, mostly concentrated in the fertile southern and western parts of the province, is vital to the province's economy; other...

, bordered by prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 to the south and the boreal forest to the north. The Peace River Country
Peace River Country
The Peace River Country is an aspen parkland region around the Peace River in Canada. It spans from northwestern Alberta to the Rocky Mountains in northeastern British Columbia, where the region is also referred to as the Peace River Block.- Geography :The Peace River Country includes the...

 is located along the Peace River
Peace River (Canada)
The Peace River is a river in Canada that originates in the Rocky Mountains of northern British Columbia and flows to the northeast through northern Alberta. The Peace River flows into the Slave River, a tributary of the Mackenzie River. The Mackenzie is the 12th longest river in the world,...

 region of the province, extending across the border into northeastern British Columbia
British Columbia
British Columbia is the westernmost of Canada's provinces and is known for its natural beauty, as reflected in its Latin motto, Splendor sine occasu . Its name was chosen by Queen Victoria in 1858...

, and is completely surrounded by boreal forest, cutting it off from the Central Parkland. The Foothills parkland covers the uplands of southwest Manitoba.

Climate

The climate is considered to be continental
Humid continental climate
A humid continental climate is a climatic region typified by large seasonal temperature differences, with warm to hot summers and cold winters....

 with long cold winters and short summers which tend to be cool to warm. Seasonal temperature extremes are common, and are greatest in Saskatchewan and Manitoba. Alberta's aspen parkland is fairly moderate in seasonal extremes by comparison, however, winters can still be long and harsh. Precipitation is generally lower than in the boreal forest, but is still higher than in the semi-arid
Semi-arid
A semi-arid climate or steppe climate describes climatic regions that receive precipitation below potential evapotranspiration, but not extremely...

 prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

s. Hence, groves of forests and lakes are able to thrive where they cannot do so in the prairies.

The Peace River Parkland is generally cooler than the Central Parkland, but still supports extensive farmland. The Foothills Parkland is located to the west of the grasslands in the southern section of the province and just east of the Canadian Rockies
Canadian Rockies
The Canadian Rockies comprise the Canadian segment of the North American Rocky Mountains range. They are the eastern part of the Canadian Cordillera, extending from the Interior Plains of Alberta to the Rocky Mountain Trench of British Columbia. The southern end borders Idaho and Montana of the USA...

. The Foothills Parkland can get very windy. Chinook wind
Chinook wind
Chinook winds , often called chinooks, commonly refers to foehn winds in the interior West of North America, where the Canadian Prairies and Great Plains meet various mountain ranges, although the original usage is in reference to wet, warm coastal winds in the Pacific Northwest.Chinook is claimed...

s are also common there during the winter.

Flora

Four significantly different habitats are common in the aspen parklands: The fescue
Fescue
Festuce is a genus of about 300 species of perennial tufted grasses, belonging to the grass family Poaceae . The genus has a cosmopolitan distribution, although the majority of the species are found in cool temperate areas...

 prairie, the woodlands, the ravines and the wetlands and lakes. A rarer habitat type, tallgrass aspen parkland
Tallgrass Aspen Parkland
The Tallgrass Aspen Parkland is a Conservation area located in southeastern Manitoba/northwestern Minnesota. The area is characterized by a mosaic of habitat types, including tallgrass prairie, aspen woodland, sedge meadow wetlands, riparian woodland, and oak savannah...

, occurs only in the extreme southeastern corner of the aspen parklands biome (southeastern Manitoba/northwestern Minnesota). The fescue prairie is a meadowland rich in vegetation variety which forms the cover for the development of the richer soils that underlie the parklands. The close association with woodlands and wetlands makes this a choice location for many plants and a preferred range or home site for a wide diversity of wildlife. The richer soil and increased precipitation favours the natural growth of fescue grass. However, varying conditions such as moisture level and grazing pressures allow for the invasion of secondary plant species.

There are numerous grasses and sedges in the fescue prairie. Gravelly and rocky terrain is a good location for parry oat grass. Dry areas favour June, porcupine and spear grass. Wet areas are often covered with slender wheat grass and timber oat grass. Prairie rose and snowberry are common shrubs found in these grasslands.

The forested, or woodlands area is dominated by trembling aspen Populus tremuloides
Populus tremuloides
Populus tremuloides is a deciduous tree native to cooler areas of North America, commonly called quaking aspen, trembling aspen, American aspen, and Quakies,. The trees have tall trunks, up to 25 metres, with smooth pale bark, scarred with black. The glossy green leaves, dull beneath, become golden...

, balsam poplar (Populus balsamifera
Populus balsamifera
Populus balsamifera, commonly called balsam poplar, bamtree, eastern balsam poplar, hackmatack, tacamahac poplar, tacamahaca,...

)
, other poplars and spruce
Spruce
A spruce is a tree of the genus Picea , a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal regions of the earth. Spruces are large trees, from tall when mature, and can be distinguished by their whorled branches and conical...

s, although other species of trees do occur. Pines, mostly jack pine
Jack Pine
Jack pine is a North American pine with its native range in Canada east of the Rocky Mountains from Northwest Territories to Nova Scotia, and the northeast of the United States from Minnesota to Maine, with the southernmost part of the range just into northwest Indiana...

 and lodgepole pine
Lodgepole Pine
Lodgepole Pine, Pinus contorta, also known as Shore Pine, is a common tree in western North America. Like all pines, it is evergreen.-Subspecies:...

 will often grow in areas that have sandy soil conditions. Other native species may include box elder
Box Elder
Box Elder may refer to:*The Box elder, Acer negundo, a species of maple*The Box elder bug, an insect which feeds on maple trees*Box Elders, a punk band from Omaha, Nebraska*Places named "Box Elder" include:*Box Elder, Montana, United States of America...

, tamarack and willow
Willow
Willows, sallows, and osiers form the genus Salix, around 400 species of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere...

, while the foothills area in the southeast of the region, such as Turtle Mountain
Turtle Mountain (plateau)
Turtle Mountain, or the Turtle Mountains, is an area in central North America, in the north-central portion of the U.S. state of North Dakota and southwestern portion of the Canadian province of Manitoba...

 or Spruce Woods Provincial Park
Spruce Woods Provincial Park
Spruce Woods Provincial Park is located in south-central Manitoba, Canada. This park has large sand dunes and the Assiniboine River passes through it.-History:The park was established in 1970...

, have woodland of white spruce
White Spruce
Picea glauca is a species of spruce native to boreal forests in the north of North America, from central Alaska east to Newfoundland, and south to northern Montana, Minnesota, Wisconsin, Michigan, upstate New York, Vermont, New Hampshire, and Maine; there is also an isolated population in the...

 and balsam fir
Balsam Fir
The balsam fir is a North American fir, native to most of eastern and central Canada and the northeastern United States .-Growth:It is a small to medium-size evergreen tree typically tall, rarely to tall, with a narrow conic crown...

 but quaking aspen will domninate where the woodland has been cleared by fire. The proportion of forests to grasslands has increased somewhat over the prairie in areas not affected by agriculture in the last 100 years. This increase is partly due to the reduction of prairie fires which used to destroy the new saplings on the fringes of the aspen groves. Also, it was a common practice for farmers to plant stands of trees as windbreaks.

Aspen woodlands support an extensive understory
Understory
Understory is the term for the area of a forest which grows at the lowest height level below the forest canopy. Plants in the understory consist of a mixture of seedlings and saplings of canopy trees together with understory shrubs and herbs...

 consisting of mid-sized and small shrubs, some herbs and ground cover. Spruce-dominated woodlands usually do not support a dense understory due to more acidic and nutrient-poor soils and a denser canopy, which reduces sunlight reaching the forest floor below. However, in areas where a mixture of aspen and spruce occur, a fairly dense understory can still thrive. In fact, the mixed wood understory, as it is called, supports the greatest diversity of forest wildlife in the aspen parkland.

Large shrubs such as red-osier dogwood, beaked willow, saskatoon, chokecherry
Chokecherry
Prunus virginiana, commonly called chokecherry, bitter-berry, or Virginia bird cherry, is a species of bird cherry native to North America, where it is found almost throughout the continent except for the Deep South and the far north.-Growth:It is a suckering shrub or small tree growing to 5 m tall...

 and pincherry, along with the smaller shrubs including prickly rose, snowberry, beaked hazelnut and low bush cranberry, form a dense entangled understory. Dense shrubbery is a typical feature in aspen-dominated forests. Common herbs found in the woodlands include: Lindley’s aster (Aster ciliolatus), northern bedstraw (Galium boreale), pea vine, Western Canada violet (Viola canadensis), dewberry
Dewberry
The dewberries are a group of species in the genus Rubus, section Rubus, closely related to the blackberries. They are small trailing brambles with berries reminiscent of the raspberry, but are usually purple to black instead of red.Dewberries are common throughout most of the Northern Hemisphere,...

 and bunchberry
Bunchberry
Bunchberry is a common name for two species of dwarf dogwoods:*Cornus canadensis - Canadian or Eastern Bunchberry*Cornus suecica - Eurasian or Northern Bunchberry...

. Mosses appear at the base of trees and on the ground.

Wetlands are very common in this biome, including lakes, shallow open water, marshes, and grassy wetlands. Glacial erosion has contributed to such features by creating depressions in which standing water can collect. In the larger depressions, permanent lakes or ponds of water remain. Many of the lakes have a saline character, thus most shore vegetation has a high tolerance of salty soils. These lakes are known as alkali lakes. Wet meadows are flooded in the spring and dry by fall. They contain rushes
Juncaceae
Juncaceae, the rush family, are a monocotyledonous family of flowering plants. There are eight genera and about 400 species. Members of the Juncaceae are slow-growing, rhizomatous, herbaceous plants, and they may superficially resemble grasses. They often grow on infertile soils in a wide range...

, sedge
Cyperaceae
Cyperaceae are a family of monocotyledonous graminoid flowering plants known as sedges, which superficially resemble grasses or rushes. The family is large, with some 5,500 species described in about 109 genera. These species are widely distributed, with the centers of diversity for the group...

s and grasses and provide excellent opportunities to study the similarities and differences of these forms of vegetation.

Rivers and streams erode valleys throughout the parkland ecoregion. Steep hills and ravines result in a unique topography. Southwest slopes with increased exposure to the sun are dry and often more grass covered while the shaded north and east exposures retain more moisture and tend to have greater forest cover. Some forms of vegetation unique to the ravines include: poplar
Poplar
Populus is a genus of 25–35 species of deciduous flowering plants in the family Salicaceae, native to most of the Northern Hemisphere. English names variously applied to different species include poplar , aspen, and cottonwood....

, spruce
Spruce
A spruce is a tree of the genus Picea , a genus of about 35 species of coniferous evergreen trees in the Family Pinaceae, found in the northern temperate and boreal regions of the earth. Spruces are large trees, from tall when mature, and can be distinguished by their whorled branches and conical...

, birch
Birch
Birch is a tree or shrub of the genus Betula , in the family Betulaceae, closely related to the beech/oak family, Fagaceae. The Betula genus contains 30–60 known taxa...

, willow
Willow
Willows, sallows, and osiers form the genus Salix, around 400 species of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere...

, and river alder.

Wildflower
Wildflower
A wildflower is a flower that grows wild, meaning it was not intentionally seeded or planted. Yet "wildflower" meadows of a few mixed species are sold in seed packets. The term "wildflower" has been made vague by commercial seedsmen who are interested in selling more flowers or seeds more...

s are an important component of the grassland association of the parkland. Look for common yarrow
Yarrow
Achillea millefolium or yarrow is a flowering plant in the family Asteraceae, native to the Northern Hemisphere. In New Mexico and southern Colorado, it is called plumajillo, or "little feather", for the shape of the leaves. In antiquity, yarrow was known as herbal militaris, for its use in...

, cut-leaf anemone
Anemone
Anemone , is a genus of about 120 species of flowering plants in the buttercup family Ranunculaceae in the north and south temperate zones...

, rock cress, creeping white prairie aster, milk vetch, late yellow loco weed, goldenrod
Goldenrod
Solidago, commonly called goldenrods, is a genus of about 100 species of flowering plants in the family Asteraceae. Most are herbaceous perennial species found in the meadows and pastures, along roads, ditches and waste areas in North America. There are also a few species native to Mexico, South...

, prairie rose
Wild Prairie Rose
Rosa arkansana is a species of rose native to a large area of central North America, between the Appalachian and Rocky Mountains from Alberta, Manitoba, and Saskatchewan south to New Mexico, Texas and Indiana.There are two varieties:*Rosa arkansana var...

, prairie crocus, and tiger lily
Tiger lily
-Botany:* Lilium catesbaei, a lily species native to southeastern North America* Lilium columbianum, a lily native to western North America* Lilium henryi, an orange lily native to central China...

.

The aspen understory

There are three main factors which influence the understory vegetation in the aspen stands of the mixed wood forest.

1. Good sun exposure encourages a dense vegetation growth below the canopy. This is of particular importance in the early spring before the trees are in leaf.

2. Warm soil and air temperature at the base level result in rapid melting process in spring which favours the growth of shrubs.

3. A large percentage of precipitation passes through the canopy. This provides a protective snow cover in winter and in warm seasons precipitation percolates through the leaf cover to nourish plants which require surface soil moisture.

The result of the above factors is an extensive understory of vegetation in the aspen forest. Common shrubs and herbs are: saskatoon, red-osier dogwood, raspberry
Raspberry
The raspberry or hindberry is the edible fruit of a multitude of plant species in the genus Rubus, most of which are in the subgenus Idaeobatus; the name also applies to these plants themselves...

, wild rose
Wild Rose
Wild Rose is the name given to certain flowering shrubs:*Genus Rosa:** Rosa acicularis, or Wild Rose, a rose species which occurs in Asia, Europe, and North America...

, currants
Ribes
Ribes is a genus of about 150 species of flowering plants native throughout the temperate regions of the Northern Hemisphere. It is usually treated as the only genus in the family Grossulariaceae. Seven subgenera are recognized....

 and bracted honeysuckle, wild sarsaparilla, hairy lungwort, asters, and peavine
Peavine
Peavine may refer to:*Merrimac, California, formerly Peavine*Peavine, Oklahoma*Peavine Mountain*West Peavine, Oklahoma*any of several plants in the genus Lathyrus...

. Twinflower
Twinflower
Linnaea is a plant genus in the family Caprifoliaceae . The genus includes a single, generally boreal to subarctic woodland subshrub species, Linnaea borealis, commonly known as Twinflower .This plant was a favorite of Carl Linnaeus, founder of the modern system of biological nomenclature, for...

, strawberries, bunchberries, horsetail
Horsetail
Equisetum is the only living genus in the Equisetaceae, a family of vascular plants that reproduce by spores rather than seeds.Equisetum is a "living fossil", as it is the only living genus of the entire class Equisetopsida, which for over one hundred million years was much more diverse and...

s and wintergreen
Wintergreen
Wintergreen is a group of plants. Wintergreen once commonly referred to plants that continue photosynthesis throughout the winter...

 form an attractive grown cover.

The mineral soil is covered by a decaying cover of organic matter. Numerous consumer
Consumer
Consumer is a broad label for any individuals or households that use goods generated within the economy. The concept of a consumer occurs in different contexts, so that the usage and significance of the term may vary.-Economics and marketing:...

s and decomposer
Decomposer
Decomposers are organisms that break down dead or decaying organisms, and in doing so carry out the natural process of decomposition. Like herbivores and predators, decomposers are heterotrophic, meaning that they use organic substrates to get their energy, carbon and nutrients for growth and...

s create humus materials. Burrowing animals mix the new fertile materials with the soil to form a rich rooting compound.

The spruce forest understory

Factors which influence the understory vegetation of spruce stands in the boreal forest association include:

1. Year round reduced sun exposure below the canopy restricts the forest undergrowth to shade tolerant species.

2. A large percentage of the precipitation is trapped in the upper tree boughs of the spruce forest and is released through evaporation. The ground cover of feather moss
Feather moss
Feather moss is a term applied to a number of moss species, many within the family Hypnaceae. Feather moss is a term applied to a moss species growing in a Boreal forest with the appearance of frond or feather like foliage. Alternatively the term Boreal forest moss is sometimes applied...

 quickly absorbs most of the moisture which does penetrate the canopy. These factors combine to cause drier conditions in the underlying mineral soils.

3. The fallen acidic spruce needles are not fully decomposed and combine with the moss base. Water held in the moss carries the acid from the spruce needles into the mineral soil and leaches out soil nutrients – leaving a highly acidic, low nutrient soil base which is unsuitable for most boreal vegetation.

As a result of the above factors the forest floor ranges from nearly devoid of vegetation to a dense carpet of feather moss. A sparse community of shade tolerant shrubs exists in this environment. Some species of plants in the understory are Green Alder
Green Alder
Alnus viridis is an alder with a wide range across the cooler parts of the Northern Hemisphere.-Description:It is a large shrub or small tree 3–12 m tall with smooth grey bark even in old age. The leaves are shiny green, ovoid, 3–8 cm long and 2–6 cm broad...

, low bush cranberry
Cranberry
Cranberries are a group of evergreen dwarf shrubs or trailing vines in the subgenus Oxycoccus of the genus Vaccinium. In some methods of classification, Oxycoccus is regarded as a genus in its own right...

, prickly rose, bunchberry, twinflower
Twinflower
Linnaea is a plant genus in the family Caprifoliaceae . The genus includes a single, generally boreal to subarctic woodland subshrub species, Linnaea borealis, commonly known as Twinflower .This plant was a favorite of Carl Linnaeus, founder of the modern system of biological nomenclature, for...

, wild lily-of-the-valley, northern Comandra and wintergreens.

The mixedwood stands understory

There are several factors which influence the mixedwood stands in the boreal forest.

Where there are stands of aspen and spruce forests in close association with each other, a mixed wood forest occurs. Each group forms its own microassociation as described previously.

When the spruce and aspen forest types are mixed, the result can be quite different from the aspen or spruce stands. Animal and vegetation associations from each type combine to create considerable diversity of habitat which is typical of either spruce or aspen stands. The mixture of the transition soils provides an attractive environment with either pure spruce or pure aspen woodlands. An example of a bird which prefers a mixed wood habitat is the yellow-rumped warbler
Yellow-rumped Warbler
Four closely related North American bird forms—the eastern Myrtle Warbler , its western counterpart, Audubon's Warbler , the northwest Mexican Black-fronted Warbler , and the Guatemalan Goldman's Warbler —are periodically lumped as the Yellow-rumped Warbler .-Classification:Since...

.

The mixedwood forest wetlands consist mainly of bog
Bog
A bog, quagmire or mire is a wetland that accumulates acidic peat, a deposit of dead plant material—often mosses or, in Arctic climates, lichens....

s, fen
Fen
A fen is a type of wetland fed by mineral-rich surface water or groundwater. Fens are characterised by their water chemistry, which is neutral or alkaline, with relatively high dissolved mineral levels but few other plant nutrients...

s and marshes. Black spruce
Black Spruce
Picea mariana is a species of spruce native to northern North America, from Newfoundland west to Alaska, and south to northern New York, Minnesota and central British Columbia...

, tamarack, willow
Willow
Willows, sallows, and osiers form the genus Salix, around 400 species of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere...

 and bog and sphagnum mosses are the major vegetation types found in these lowlands. Dwarf birch
Dwarf Birch
Betula nana is a species of birch in the family Betulaceae, found mainly in the tundra of the Arctic region.-Description:...

 and sedges cover large, wet areas with jack pine
Jack Pine
Jack pine is a North American pine with its native range in Canada east of the Rocky Mountains from Northwest Territories to Nova Scotia, and the northeast of the United States from Minnesota to Maine, with the southernmost part of the range just into northwest Indiana...

 occurring on the sandy ridges.

Fauna

Wildlife in the parklands include moose
Moose
The moose or Eurasian elk is the largest extant species in the deer family. Moose are distinguished by the palmate antlers of the males; other members of the family have antlers with a dendritic configuration...

 (Alces alces), white-tailed deer
White-tailed Deer
The white-tailed deer , also known as the Virginia deer or simply as the whitetail, is a medium-sized deer native to the United States , Canada, Mexico, Central America, and South America as far south as Peru...

 (Odocoileus virginianus), black bear
American black bear
The American black bear is a medium-sized bear native to North America. It is the continent's smallest and most common bear species. Black bears are omnivores, with their diets varying greatly depending on season and location. They typically live in largely forested areas, but do leave forests in...

 (Ursus americanus), coyote
Coyote
The coyote , also known as the American jackal or the prairie wolf, is a species of canine found throughout North and Central America, ranging from Panama in the south, north through Mexico, the United States and Canada...

 (Canis latrans), northern pocket gopher
Northern Pocket Gopher
The Northern Pocket Gopher, Thomomys talpoides, was first discovered by Lewis and Clark on April 9, 1805 at the mouth of the Knife River, North Dakota. These animals are often rich brown or yellowish brown, but also grayish or closely approaching local soil color and have white markings under chin...

s (Thomomys talpoides), thirteen-lined ground squirrel
Thirteen-lined ground squirrel
The thirteen-lined ground squirrel , also known as the striped gopher, leopard ground squirrel, squinney, and as the leopard-spermophile in Audubon’s day, is a ground squirrel....

s, Richardson’s ground squirrels, North American Beaver (Castor canadensis), snowshoe hare
Snowshoe Hare
The Snowshoe Hare , also called the Varying Hare, or Snowshoe Rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet and the marks its tail leaves. The animal's feet prevent it from sinking into the snow when it hops and walks...

 (Lepus americanus), weasel
Weasel
Weasels are mammals forming the genus Mustela of the Mustelidae family. They are small, active predators, long and slender with short legs....

s and gray wolf
Gray Wolf
The gray wolf , also known as the wolf, is the largest extant wild member of the Canidae family...

 (canis lupus). Bear, moose, foxes, coyotes, beaver, snowshoe hare and red squirrels are found most often in the mixedwood stands compared to the aspen forests and spruce forests.

Burrowing rodents such as Richardson’s ground squirrels, thirteen-lined ground squirrels, and pocket gophers play a major role in the balance between the aspen groves and the grassland. These excavators make mounds of fresh soil which are ideal locations for the germination of poplar seeds. Once established, these trees spread by suckering, thus creating a new aspen groves.

White-tailed deer
White-tailed Deer
The white-tailed deer , also known as the Virginia deer or simply as the whitetail, is a medium-sized deer native to the United States , Canada, Mexico, Central America, and South America as far south as Peru...

 finds shelter in the aspen and graze on the grasslands; coyote
Coyote
The coyote , also known as the American jackal or the prairie wolf, is a species of canine found throughout North and Central America, ranging from Panama in the south, north through Mexico, the United States and Canada...

s and fox
Fox
Fox is a common name for many species of omnivorous mammals belonging to the Canidae family. Foxes are small to medium-sized canids , characterized by possessing a long narrow snout, and a bushy tail .Members of about 37 species are referred to as foxes, of which only 12 species actually belong to...

es hunt the resident rodents. Historically, bison
Bison
Members of the genus Bison are large, even-toed ungulates within the subfamily Bovinae. Two extant and four extinct species are recognized...

 grazed on the grassland and helped to prevent the spread of aspen groves. However, bison are now mostly absent due to overhunting during settlement in the 19th Century and extensive loss of habitat due to agriculture. Bison, however, can still be seen in protected areas such as Elk Island National Park
Elk Island National Park
Elk Island National Park , is one of 43 national parks and park reserves administered by the Parks Canada Agency. This “island of conservation” is located 35 km east of Edmonton, Alberta along the Yellowhead Highway, which nearly bisects the park...

 east of Edmonton
Edmonton
Edmonton is the capital of the Canadian province of Alberta and is the province's second-largest city. Edmonton is located on the North Saskatchewan River and is the centre of the Edmonton Capital Region, which is surrounded by the central region of the province.The city and its census...

 and in farms, where they are raised for meat.

Wildlife in the woodlands is varied and abundant. The varying hare, weasel, fox, coyote, and white-tailed deer make their homes in this region, while water dependent mammals who make the ravines and wetland areas of the ecoregion their home are beaver, muskrat
Muskrat
The muskrat , the only species in genus Ondatra, is a medium-sized semi-aquatic rodent native to North America, and introduced in parts of Europe, Asia, and South America. The muskrat is found in wetlands and is a very successful animal over a wide range of climates and habitats...

s, otters and mink.

Birds of the aspen parkland include kingfisher
Kingfisher
Kingfishers are a group of small to medium sized brightly coloured birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species being found in the Old World and Australia...

s, ruffed grouse
Ruffed Grouse
The Ruffed Grouse is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is non-migratory.The Ruffed Grouse is frequently referred to as a "partridge"...

, magpie
Magpie
Magpies are passerine birds of the crow family, Corvidae.In Europe, "magpie" is often used by English speakers as a synonym for the European Magpie, as there are no other magpies in Europe outside Iberia...

s and northern orioles. and in particular several species of warblers find this a preferred habitat.

Extensive cultivation has disturbed the habitats of some birds which nest and feed on the fescue grassland. However, the horned lark and meadowlark
Meadowlark
Meadowlarks are birds belonging to the genus Sturnella in the New World family Icteridae.This genus includes seven species of largely insectivorous grassland birds...

 have managed to adapt to the new conditions. Song sparrow
Song Sparrow
The Song Sparrow is a medium-sized American sparrow.Adults have brown upperparts with dark streaks on the back and are white underneath with dark streaking and a dark brown spot in the middle of the breast. They have a brown cap and a long brown rounded tail. Their face is grey with a streak...

, vesper sparrow
Vesper Sparrow
The Vesper Sparrow is a medium-sized American sparrow. It is the only member of the genus Pooecetes.Adults have light brown upperparts and light underparts, both with darker streaking. They have a white eye ring and a long dark brown tail which shows white outer feathers in flight.Their breeding...

, and American goldfinch
American Goldfinch
The American Goldfinch , also known as the Eastern Goldfinch and Wild Canary, is a small North American bird in the finch family...

 can often be seen in open areas.

The woodlands meanwhile are abundant with a variety of bird species. Black-capped chickadee
Black-capped Chickadee
The Black-capped Chickadee is a small, North American songbird, a passerine bird in the tit family Paridae. It is the state bird of both Maine and Massachusetts in the United States, and the provincial bird of New Brunswick in Canada...

, hairy woodpecker
Hairy Woodpecker
The Hairy Woodpecker is a medium-sized woodpecker, averaging approximately 250 mm in length with a 380 mm wingspan...

s, ruffed grouse
Ruffed Grouse
The Ruffed Grouse is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is non-migratory.The Ruffed Grouse is frequently referred to as a "partridge"...

, magpie
Magpie
Magpies are passerine birds of the crow family, Corvidae.In Europe, "magpie" is often used by English speakers as a synonym for the European Magpie, as there are no other magpies in Europe outside Iberia...

s, and great horned owl
Great Horned Owl
The Great Horned Owl, , also known as the Tiger Owl, is a large owl native to the Americas. It is an adaptable bird with a vast range and is the most widely distributed true owl in the Americas.-Description:...

s can be observed in all seasons. Summer residents include: red-eyed vireo
Red-eyed Vireo
The Red-eyed Vireo, Vireo olivaceus, is a small American songbird, 13–14 cm in length. It is somewhat warbler-like but not closely related to the New World warblers...

, least flycatcher
Least Flycatcher
The Least Flycatcher , , is a small insect-eating bird...

 and northern oriole.

Birds which prefer the wetland habitat include kingfisher
Kingfisher
Kingfishers are a group of small to medium sized brightly coloured birds in the order Coraciiformes. They have a cosmopolitan distribution, with most species being found in the Old World and Australia...

s and bank swallows. Finally there is an abundance of bird life around the wetland marshes. Many species of ducks make their summer homes in these waters and Canada geese nest in the more remote marshes. Blackbirds, marsh wren
Marsh Wren
The Marsh Wren is a small North American songbird of the wren family. It is sometimes called Long-billed Marsh Wren to distinguish it from the Sedge Wren, also known as Short-billed Marsh Wren....

s and black tern
Black Tern
The Black Tern, Chlidonias niger, is a small tern generally found in or near inland water in Europe and North America. As its name suggests, it has predominantly dark plumage.- Description :...

s nest in the reeds. Franklin gulls nest in the marsh vegetation, but range over agricultural fields for grasshoppers, crickets, and mice. Shore birds include: avocet
Avocet
The four species of Avocets are a genus, Recurvirostra, of waders in the same avian family as the stilts.Avocets have long legs and long, thin, upcurved bills which they sweep from side to side when feeding in the brackish or saline wetlands they prefer...

, piping plover
Piping Plover
The Piping Plover is a small sand-colored, sparrow-sized shorebird that nests and feeds along coastal sand and gravel beaches in North America. The adult has yellow-orange legs, a black band across the forehead from eye to eye, and a black ring around the neck...

, spotted sandpiper
Spotted Sandpiper
The Spotted Sandpiper is a small shorebird, 18–20 cm long. Together with its sister species, the Common Sandpiper they make up the genus Actitis...

, willet
Willet
The Willet, Tringa semipalmata , is a large shorebird in the sandpiper family. It is a good-sized and stout scolopacid, the largest of the shanks...

, common snipe
Common Snipe
The Common Snipe is a small, stocky wader native to the Old World. The breeding habitat is marshes, bogs, tundra and wet meadows throughout northern Europe and northern Asia...

 and killdeer
Killdeer
The Killdeer is a medium-sized plover.Adults have a brown back and wings, a white belly, and a white breast with two black bands. The rump is tawny orange. The face and cap are brown with a white forehead. They have an orange-red eyering...

.

The invertebrate population in the woodland is enormous. Some of the most common invertebrates are roundworms, snails, segmented worms, centipedes, mites, spiders and mosquitoes. Poplar gore beetles and forest tent caterpillar
Forest tent caterpillar
The Forest Tent Caterpillar Moth is the larva of a North American moth, found throughout the United States and Canada, and most common in the eastern regions....

s are destructive to the tree cover. Insects of the wetlands in this region include caddis flies, mayflies
Mayfly
Mayflies are insects which belong to the Order Ephemeroptera . They have been placed into an ancient group of insects termed the Palaeoptera, which also contains dragonflies and damselflies...

 and black flies
Black fly
A black fly is any member of the family Simuliidae of the Culicomorpha infraorder. They are related to the Ceratopogonidae, Chironomidae, and Thaumaleidae. There are over 1,800 known species of black flies . Most species belong to the immense genus Simulium...

.

Human use, threats and conservation

Before European colonization, there were large areas of western aspen and aspen parkland in the west of what would become Canada and the United States. This was maintained by light to moderate fires with a frequency of 3 to 15 years. Fire also swept the Rocky Mountains
Rocky Mountains
The Rocky Mountains are a major mountain range in western North America. The Rocky Mountains stretch more than from the northernmost part of British Columbia, in western Canada, to New Mexico, in the southwestern United States...

 aspen as frequently as every ten years, creating large areas of parkland. Settlement increased fire frequency in the late 19th century until fire suppression became popular.

Most of the aspen parkland, like the prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

 biome, has been extensively altered by agriculture over the last 100 years since settlement first began in the late 19th Century. While the climate is generally cooler than in the prairie
Prairie
Prairies are considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and grasses, herbs, and shrubs, rather than trees, as the dominant vegetation type...

s, the climate is still mild and dry enough to support large-scale farming of crops such as canola
Canola
Canola refers to a cultivar of either Rapeseed or Field Mustard . Its seeds are used to produce edible oil suitable for consumption by humans and livestock. The oil is also suitable for use as biodiesel.Originally, Canola was bred naturally from rapeseed in Canada by Keith Downey and Baldur R...

 (Brassica napsus), alfalfa
Alfalfa
Alfalfa is a flowering plant in the pea family Fabaceae cultivated as an important forage crop in the US, Canada, Argentina, France, Australia, the Middle East, South Africa, and many other countries. It is known as lucerne in the UK, France, Australia, South Africa and New Zealand, and known as...

 (Medicago sativa) and wheat
Wheat
Wheat is a cereal grain, originally from the Levant region of the Near East, but now cultivated worldwide. In 2007 world production of wheat was 607 million tons, making it the third most-produced cereal after maize and rice...

 (Triticum aestivum), and livestock grazing. The soils in the aspen parkland biome are also quite fertile, especially around Edmonton
Edmonton
Edmonton is the capital of the Canadian province of Alberta and is the province's second-largest city. Edmonton is located on the North Saskatchewan River and is the centre of the Edmonton Capital Region, which is surrounded by the central region of the province.The city and its census...

 and Saskatoon
Saskatoon
Saskatoon is a city in central Saskatchewan, Canada, on the South Saskatchewan River. Residents of the city of Saskatoon are called Saskatonians. The city is surrounded by the Rural Municipality of Corman Park No. 344....

. Oil and natural gas exploration and drilling have also disturbed the natural habitat, especially in Alberta and northeastern British Columbia. As a result, less than 10% of the original habitat remains undisturbed.

The largest blocks of intact parkland can be found in Moose Mountain Provincial Park
Moose Mountain Provincial Park
Moose Mountain Provincial Park is a Provincial Park, located in southeastern Saskatchewan 24 km north of the town of Carlyle. It is one of Saskatchewan's few parks with towns on the inside of the park. The only town in Moose Mountain is Kenosee Lake....

 north of Carlyle, Saskatchewan
Carlyle, Saskatchewan
-Entertainment:The Bear Claw Casino & Hotel is located near Carlyle.The Shackshakers is a recording musical group that was formed in Carlyle by Shawn Nagy in 1988.-Climate:-Notable people:* Brenden Morrow - hockey player* Eliza Beatty - Silver Cross Mother...

 and Bronson Forest in Saskatchewan, and Elk Island National Park
Elk Island National Park
Elk Island National Park , is one of 43 national parks and park reserves administered by the Parks Canada Agency. This “island of conservation” is located 35 km east of Edmonton, Alberta along the Yellowhead Highway, which nearly bisects the park...

 and Canadian Forces Base Wainwright
CFB Wainwright
Canadian Forces Base Wainwright, commonly referred to as CFB Wainwright is a Canadian Forces Base located in Denwood, Alberta, adjacent to the town of Wainwright.-Military Camp Wainwright:...

 in Alberta. The rest of the parkland area does contain fragments of original habitat, some in protected areas such as Spruce Woods Provincial Park
Spruce Woods Provincial Park
Spruce Woods Provincial Park is located in south-central Manitoba, Canada. This park has large sand dunes and the Assiniboine River passes through it.-History:The park was established in 1970...

 and Turtle Mountain Provincial Park
Turtle Mountain Provincial Park
Turtle Mountain Provincial Park is a provincial park located in the southwestern portion of the Canadian province of Manitoba. Within it is the Adam Lake and Max Lake campgrounds. The Park is known for its bike trails, fishing, back country cabins and canoe routes...

 in Manitoba, and Porcupine Provincial Forest in Saskatchewan.

External links

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