2011 in paleontology
Encyclopedia

Lobopods

Name Novelty Status Authors Age Unit Location Notes Images
Diania cactiformis
Diania
Diania is an extinct genus of animal found in the Lower Cambrian Maotianshan shale of China, represented by a single species D. cactiformis. Known during its investigation by the nickname "walking cactus", this remarkable organism belongs to a group known as the armoured lobopodians and has a...


Valid
  • Liu
  • Steiner
  • Dunlop
  • et al.

Lower Cambrian

Maotianshan shale

 Mainland China

A lobopodia
Lobopodia
Lobopodia is a group of poorly understood animals, which mostly fall as a stem group of arthropods. Their fossil range dates back to the Early Cambrian. Lobopods are segmented and typically bear legs with hooked claws on their ends....

n.

Siberion lenaicus
Siberion
Siberion is an extinct genus of lobopod from the Sinsk biota of Russia. Its anatomy, including the proboscis-like organ projecting from the face and prominent grasping first pair of appendages, suggests that xenusians like this organism may have been phylogenetically related to anomalocaridids,...


Valid
  • Dzik

Early Cambrian

Sinsk Formation

 Russia

A lobopodia
Lobopodia
Lobopodia is a group of poorly understood animals, which mostly fall as a stem group of arthropods. Their fossil range dates back to the Early Cambrian. Lobopods are segmented and typically bear legs with hooked claws on their ends....

n.

Arachnids

Name Novelty Status Authors Age Unit Location Notes Images

Nephila jurassica
Nephila jurassica
Nephila jurassica is an extinct species of spider in the family Nephilidae, which contains the golden silk orb-weavers. The species is known only from the Middle Jurassic Jiulongshan Formation, part of the Daohugou Beds, near the village of Daohugou in Ningcheng County, northeastern China.-History...


sp nov

valid

Selden, Shih & Ren

Middle Jurassic
Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from 176-161 million years ago. In European lithostratigraphy, rocks of this Middle Jurassic age are called the Dogger....


Jiulongshan Formation

 Mainland China

largest fossil spider described

Insects

Name Novelty Status Authors Age Unit Location Notes Images

Agulla protomaculata
Agulla protomaculata
Agulla protomaculata is an extinct species of snakefly in the raphidiid genus Agulla. The species is solely known from the Middle Eocene, Lutetian stage, Parachute Member, part of the Green River Formation, in the Piceance Creek Basin and Uinta Basin, Garfield County, northwestern Colorado,...


sp nov

valid

Engel
Michael S. Engel
Michael S. Engel is an American paleontologist and entomologist. He has undertaken field work in Central Asia, Asia Minor, and the Western Hemisphere, and published more than 300 papers in scientific journals. He was trained at the University of Kansas where in 1993 he received a B.S. in Cellular...


Lutetian
Lutetian
The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between and . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene subepoch...


Green River Formation
Green River Formation
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in winter. Each pair of layers is called a varve and...


 United States

Only snakefly from the Green River Formation

Baltimartyria rasnitsyni

sp nov

valid

Mey

Early Eocene

Baltic amber
Baltic amber
The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite, with about 80% of the world's known amber found there. It dates from 44 million years ago...


 Europe

the second species in Baltimartyria


Darekia sanguinea
Darekia
Darekia is a genus of stem group Neoptera which is due to be published by Acta Palaeontologica Polonica. It existed in what is now Poland, during the upper Carboniferous period. It was described by Jakub Prokop, Wieslaw Krzemiński, Ewa Krzemińska, and Dariusz Wojciechowski in 2011, and the type...


nomen nudum
  • Prokop
  • Krzemiński
  • Krzemińska
  • Wojciechowski

Upper Carboniferous

Upper Silesian Coal Basin

 Poland

A stem winged insect.

Diochus electrus
Diochus electrus
Diochus electrus is an extinct species of rove beetle in genus Diochus, the only definitive fossil species in subfamily Staphylininae. The species is known only from the middle Eocene, Lutetian stage Baltic amber found in the Blaue Erde deposits, Baltic region, Northern Europe.-History and...


sp nov

valid

Chatzimanolis & Engel
Michael S. Engel
Michael S. Engel is an American paleontologist and entomologist. He has undertaken field work in Central Asia, Asia Minor, and the Western Hemisphere, and published more than 300 papers in scientific journals. He was trained at the University of Kansas where in 1993 he received a B.S. in Cellular...


Lutetian
Lutetian
The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between and . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene subepoch...


Baltic amber
Baltic amber
The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite, with about 80% of the world's known amber found there. It dates from 44 million years ago...


 Europe

first valid Staphylininae species from the fossil record


Dryinus rasnitsyni
Dryinus rasnitsyni
Dryinus rasnitsyni is an extinct species of wasp in the dryinid genus Dryinus. The species is solely known from the early Miocene, Burdigalian stage, Dominican amber deposits on the island of Hispaniola.-History and classification:...


sp nov

valid

Olmi & Guglielmino

Burdigalian
Burdigalian
The Burdigalian is, in the geologic timescale, an age or stage in the early Miocene. It spans the time between 20.43 ± 0.05 Ma and 15.97 ± 0.05 Ma...


Dominican amber
Dominican amber
Dominican amber is amber from the Dominican Republic. Resin from the extinct species Hymenaea protera is the source of Dominican amber and probably of most amber found in the tropics....


 Dominican Republic

second fossil Dryinus lamellatus group species


Leptolingia calonervis

sp.
  • Shi Chaofan
  • Yang Qiang
  • Ren Dong

Middle Jurassic

Jiulongshan Formation

 Mainland China

Leptolingia imminuta

sp.
  • Yushuang Liu
  • Chaofan Shi
  • Dong Ren

Middle Jurassic

Daohugou Beds
Daohugou Beds
The Daohugou Beds are a series of fossil-bearing rock deposits located in northeastern China, with the type locality around Daohugou village of Ningcheng County south of Chifeng, Inner Mongolia, but extending into southwest Liaoning. The rocks are grey, finely bedded, lacustrine, sandy mudstones...


 Mainland China

the smallest species known in the family Grammolingiidae

Litholingia ptesa

sp.
  • Shi Chaofan
  • Yang Qiang
  • Ren Dong

Middle Jurassic

Jiulongshan Formation

 Mainland China

Myanmymar
Myanmymar
Myanmymar is an extinct genus of fairyfly preserved in amber from Myanmar. It has only one species, Myanmymar aresconoides. It is dated to the Early Cretaceous, at least a hundred million years ago. As of 2011, it is the oldest known fossil mymarid.-Discovery and geologic time range:Myanmymar...


gen et sp nov

valid

Huber

Albian
Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous epoch/series. Its approximate time range is 112.0 ± 1.0 Ma to 99.6 ± 0.9 Ma...


Burmese amber

 Myanmar

oldest mymarid described


Ordralfabetix
Ordralfabetix
Ordralfabetix is an extinct genus of planthopper in the family Lophopidae and containing the single species Ordralfabetix sirophatanis...


gen et sp.

valid

Szwedo

Ypresian
Ypresian
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between and , is preceded by the Thanetian age and is followed by the Eocene Lutetian age....


Oise amber

 Early Modern France

Second fossil lophopid planthopper from Europe

Plumalexius
Plumalexius
Plumalexius is a genus of wasp in the extinct monotypic family Plumalexiidae, containing a single species, Plumalexius rasnitsyni. The species is known only from the Late Cretaceous White Oaks Pit in Sayreville, New Jersey, USA....


fam, gen et sp.

vaild

Brothers

Turonian
Turonian
The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous epoch, or a stage in the Upper Cretaceous series. It spans the time between 93.5 ± 0.8 Ma and 89.3 ± 1 Ma...


New Jersey amber

 United States

sister genus to Plumariidae

Titanomyrma lubei
Gen. et sp.

valid
  • Archibald
  • Johnson
  • Mathewes
  • Greenwood

Early Eocene

Green River Formation
Green River Formation
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in winter. Each pair of layers is called a varve and...

 USA

Messel shales
Messel pit
The Messel Pit is a disused quarry near the village of Messel, about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of fossils, it has significant geological and scientific importance...

, Germany

Eckfeld Maar, Germany

 United States

 Germany

new genus for the largest known ants

Zdenekia silesiensis
Zdenekia
Zdenekia is a genus of prehistoric winged insect. It contains the species Z. grandis from the Czech Republic, Z. occidentalis from Belgium, and Z. silesiensis from Poland....


nomen nudum
  • Prokop
  • Krzemiński
  • Krzemińska
  • Wojciechowski

Upper Carboniferous

Upper Silesian Coal Basin

 Poland

A stem winged insect.
Mickoleitia longimanus
Coxoplectoptera
Coxoplectoptera or "chimera wings" is a primitive, extinct order of winged insects containing one family, Mickoleitiidae, described by Staniczek, Bechly & Godunko in 2011...


ord., fam., gen. et sp.
  • Staniczek
  • Bechly
  • Godunko

Lower Cretaceous

Crato Formation
Crato Formation
The Crato Formation is a geologic formation of Early Cretaceous age in northeastern Brazil's Araripe Basin. It is an important Lagerstätte for palaeontologists. The strata were laid down mostly during the early Albian age, about 108 million years ago, in a shallow inland sea...


 Brazil

A strange stem group representative of mayflies that has been attributed to a new family Mickoleitiidae and a new fossil insect order Coxoplectoptera
Coxoplectoptera
Coxoplectoptera or "chimera wings" is a primitive, extinct order of winged insects containing one family, Mickoleitiidae, described by Staniczek, Bechly & Godunko in 2011...

.


Newly named jawless vertebrates

Name Novelty Status Authors Age Unit Location Notes Images

Althaspis tarloi
Althaspis
Althaspis is an extinct genus of jawless fish....


sp.

Valid
  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Djurinaspis secunda
Djurinaspis
Djurinaspis is an extinct genus of jawless fish which existed during the early Devonian period. It was originally described by Novitskaya in 1983. A new species, D. secunda, from Ukraine was described by Victor Voichyshyn in 2011....


sp.

Valid
  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Palanasaspis
Palanasaspis
Palanasaspis chekhivensis is an extinct species of jawless fish which existed during the early Devonian period....


Gen. et sp.

Valid
  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Podolaspis danieli
Podolaspis
Podolaspis is an extinct genus of jawless fish which existed during the early Devonian period. It was originally described by Zych in 1931, and contains the species P. lerichei, and P. danieli.-External links:* at the Paleobiology Database...


sp.

Valid
  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Semipodolaspis
Semipodolaspis
Semipodolaspis is an extinct genus of jawless vertebrate....


Gen. et sp.

Valid
  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Shuyu
Shuyu
Shuyu is an extinct genus of early jawless vertebrate from the Early or Middle Silurian period . It is the basalmost known eugaleaspidiform galeaspid and it lived in what is now northwestern Zhejiang Province, Southeast China. It is known from more than 20 headshields and at least 20 of them...


Gen. et comb.

Valid
  • Gai
  • Donoghue
  • et al.

Late Telychian/Early Wenlock

Maoshan Formation

The basalmost known eugaleaspidiform galeaspid.

Zenaspis kasymyri
Zenaspis
Zenaspis is an extinct genus of jawless fish which existed during the early Devonian period. It contains the species Z. powriei, and a species from Ukraine, Z. kasymyri, described by Victor Voichyshyn in 2011....


sp.

Valid
  • Voichyshyn

Early Devonian

Dniester Formation

A osteostracan

Newly named acanthodian

Name Novelty Status Authors Age Unit Location Notes Images

Nerepisacanthus
Nerepisacanthus
Nerepisacanthus is an extinct genus of acanthodian, probably acritolepid, from Middle Silurian deposits of New Brunswick, Canada. Nerepisacanthus is known from many incomplete but articulated specimens. It was collected from the Cunningham Creek Formation, near Nerepis, southern New Brunswick. It...


Gen. et sp.

Valid
  • Burrow

?Telychian/Ludlow

Cunningham Creek Formation

An acritolepid ischnacanthiform
Ischnacanthiformes
Ischanacanthiformes is an order of Acanthodii of spiny sharks. It contains Ischnacanthus. This order of sharks have large teeth on their Palatoquadrate....

.

Newly named cartilaginous fishes

Name Novelty Status Authors Age Unit Location Notes Images

Kawichthys
Kawichthys
Kawichthys is an extinct genus of symmoriiform cartilaginous fish from Upper Pennsylvanian deposits of Kansas, USA. Kawichthys is known from two well preserved three-dimensional neurocrania: the holotype KUVP 152144 is associated with some disturbed and broken postcranial elements, but the...


Gen. et sp.

Valid
  • Pradel
  • Tafforeau
  • et al.

Late Virgilian

Douglas Group

A symmoriiform.

Reginaselache
Reginaselache
Reginaselache is an extinct genus of xenacanthiform shark known from the Early Carboniferous of central Queensland, Australia. It was found in the middle the Ducabrook Formation, northwest of Springsure. It was first named by Susan Turner and Carole J. Burrow in 2011 and the type species is...


Gen. et sp.

Valid
  • Turner
  • Burrow

Middle Viséan

Ducabrook Formation

A xenacanthiform
Xenacanthida
Xenacanthida is an order of prehistoric sharks that appeared during the Lower Carboniferous period. The family includes the families Xenacanthidae, Diplodoselachidae, and Orthacanthidae. The most notable members of the group are the genera Xenacanthus and Orthacanthus. Some Xenacanthus may have...

.

Newly named bony fishes

Name Novelty Status Authors Age Unit Location Notes Images

Beishanichthys
Beishanichthys
Beishanichthys is an extinct genus of scanilepiform bony fish which existed in Gansu Province, China during the early Triassic period. It is based on fossils from the Lower Triassic lake deposits exposed in Beishan area. It was first named by Guang-Hui Xu, Ke-Qin Gao in 2011 and the type species is...


Gen. et sp.

Valid
  • Xu
  • Gao

Early Triassic

A scanilepiform.

Bourbonnella
Bourbonnella
Bourbonnella is an extinct genus of fish in the family Aeduellidae....


Gen. et sp.

Valid
  • Mickle

Carboniferous

Manning Canyon Shale Formation

An aeduellid.

Caucasichthys
Caucasichthys
Caucasichthys is an extinct genus of perciform bony fish which existed in Russia during the middle Eocene epoch . It is known from the Gorny Luch locality of northern Caucasus. It was first named by Alexandre F. Bannikov, Giorgio Carnevale and N. V. Parin in 2011 and the type species is...


Gen. et sp.

Valid
  • Bannikov
  • Carnevale
  • Parin

Bartonian

A perciform
Perciformes
The Perciformes, also called the Percomorphi or Acanthopteri, is one of the largest orders of vertebrates, containing about 40% of all bony fish. Perciformes means perch-like. They belong to the class of ray-finned fish and comprise over 7,000 species found in almost all aquatic environments...

.

Guntherichthys
Guntherichthys
Guntherichthys is an extinct genus of palaeoniscoid....


Gen. et sp.

Valid
  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Leufuichthys
Leufuichthys
Leufuichthys is an extinct genus of clupeomorph bony fish which existed in Patagonia, Argentina during the late Cretaceous epoch...


Gen. et sp.

Valid
  • Gallo
  • Calvo
  • Kellner

Turonian/Coniacian

Portezuelo Formation
Portezuelo Formation
The Portezuelo Formation is a geologic formation outcropping in the Mendoza, Río Negro and Neuquén provinces of Argentina. It is the fourth-oldest formation in the Neuquén Group and the older of the two formations in the Río Neuquén Subgroup...


A clupeomorph.

Kyphosichthys
Kyphosichthys
Kyphosichthys is an extinct genus of basal actinopterygian bony fish known from the lower Middle Triassic marine deposits in Luoping, eastern Yunnan Province, southwestern China. The this species is the first known fossil record of highly deep-bodied Neopterygii ginglymodians....


Gen. et sp.

In press
  • Xu
  • Wu

Middle Anisian

Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....


A ginglymodian actinopterygian
Actinopterygii
The Actinopterygii or ray-finned fishes constitute a class or sub-class of the bony fishes.The ray-finned fishes are so called because they possess lepidotrichia or "fin rays", their fins being webs of skin supported by bony or horny spines , as opposed to the fleshy, lobed fins that characterize...

.

Luoxiongichthys
Luoxiongichthys
Luoxiongichthys is an extinct genus of basal actinopterygian bony fish known from the lower Middle Triassic of Luoxiong Town, Luoping County of Yunnan Province, southwestern China...


Gen. et sp.

In press
  • Wen
  • Zhang
  • et al.

Middle Anisian

Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....


A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 actinopterygian
Actinopterygii
The Actinopterygii or ray-finned fishes constitute a class or sub-class of the bony fishes.The ray-finned fishes are so called because they possess lepidotrichia or "fin rays", their fins being webs of skin supported by bony or horny spines , as opposed to the fleshy, lobed fins that characterize...

.

Pietschellus
Pietschellus
Pietschellus is an extinct genus of enigmatic bony fish which existed in northern Italy during the early Eocene epoch . It is known from a single well-preserved nearly complete specimen recovered from the Monte Postale site of the Monte Bolca locality. It was first named by Alexandre F...


Gen. et sp.

Valid
  • Bannikov
  • Carnevale

Ypresian

Monte Bolca
Monte Bolca
Monte Bolca is a lagerstätte near Verona, Italy that was one of the first fossil sites with high quality preservation known to Europeans, and is still an important source of fossils from the Eocene....


A percomorph
Perciformes
The Perciformes, also called the Percomorphi or Acanthopteri, is one of the largest orders of vertebrates, containing about 40% of all bony fish. Perciformes means perch-like. They belong to the class of ray-finned fish and comprise over 7,000 species found in almost all aquatic environments...

.

Sinosaurichthys
Sinosaurichthys
Sinosaurichthys is an extinct genus of saurichthyid ray-finned fish which existed in southwestern China, during the middle Triassic . Fossils have been found in the Upper Member of the Guanling Formation of two localities: Yangjuan of Panxian County, Guizhou Province, and Dawazi of Luoping, Yunnan...


Gen. et 3 sp.

Valid
  • Wu
  • Sun
  • et al.

Anisian

Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....


A saurichthyid.

Spinofacia
Spinofacia
Spinofacia is an extinct genus of palaeoniscoid....


Gen. et sp.

Valid
  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Newly named lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Tambaroter
Tambaroter
Tambaroter is an extinct genus of ostodolepid microsaur from the Early Permian of Germany. The type species T. carrolli was named in 2011. Tambaroter is known from a single skull found in the Tambach Formation, which is the lowermost unit of the Upper Rotliegend...


Gen. et sp.

Valid
  • Henrici
  • Martens
  • et al.

Early Permian

Tambach Formation

An ostodolepid.

Newly named temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Arachana
Arachana
Arachana is an extinct genus of rhinesuchid-like temnospondyl known from the Late Permian or Early Triassic of northeastern Uruguay. Arachana was first named by Graciela Piñeiro, Alejandro Ramos and Claudia Marsicano in 2011 and the type species is A. nigra...


Gen. et sp.

In press
  • Piñeiro
  • Ramos
  • Marsicano

? Late Permian

Buena Vista Formation

A rhinesuchid-like stereospondyl.

Calmasuchus
Calmasuchus
Calmasuchus is a genus of capitosaurian temnospondyl which lived during the middle Triassic . Fossils of Calmasuchus have been recovered from the La Mora site of the Catalonian basin in Barcelona of Spain...


Gen. et sp.

Valid
  • Fortuny
  • Galobart
  • De Santisteban

early-middle Anisian

Catalonian basin

A capitosauria
Capitosauria
Capitosauria is a clade of stereospondyl temnospondyls. Capitosauria and its sister taxon Trematosauria are the only major temnospondyl clades that existed during the Mesozoic....

n.

Rubeostratilia
Rubeostratilia
Rubeostratilia is an extinct genus of amphibamid temnospondyl from the Early Permian of Texas. It is known from a single skull. This genus was named by Hélène Bourget and Jason S. Anderson in 2011, and the type species is Rubeostratilia texensis...


Gen. et sp.

Valid
  • Bourget
  • Anderson

Early Permian

An amphibamid
Amphibamidae
Amphibamidae is an extinct family of dissorophoid euskelian temnospondyls. The earliest amphibamids such as Amphibamus are known from Early Permian strata in the United States, while the last known amphibamid, Micropholis, is known from the Early Triassic Karoo Basin of South Africa...

.

Newly named lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Seminobatrachus
Seminobatrachus
Seminobatrachus is an extinct genus of neotenic salamander from late Paleocene to early Eocene deposits of Cherkassy Region, central Ukraine. It is known from 14 skeletons from a drill hole near Boltyshka village. It was first named by Pavel P. Skutschas and Yuri M. Gubin in 2011 and the type...


Gen. et sp.

In press
  • Skutschas
  • Gubin

late Paleocene - early Eocene

A urodelan salamander
Salamander
Salamander is a common name of approximately 500 species of amphibians. They are typically characterized by a superficially lizard-like appearance, with their slender bodies, short noses, and long tails. All known fossils and extinct species fall under the order Caudata, while sometimes the extant...

.

Wesserpeton
Wesserpeton
Wesserpeton is an extinct genus of albanerpetontid amphibian known from the Isle of Wight, southern England.-Description:Wesserpeton is known from the holotype NHMUK PV R36521, nearly complete fused frontals and from the referred materials NHMUK PV R36522–36568 and R36595–36611...


Gen. et sp.

In press
  • Sweetman
  • Gardner

Barremian

Wessex Formation
Wessex Formation
The Wessex Formation is an English fossil site and geological formation that dates to the Barremian stage of the Early Cretaceous. It is part of the Wealden Group and underlies the younger Vectis Formation.-Invertebrates:...


An albanerpetontid.

Newly named basal anapsids

Name Novelty Status Authors Age Unit Location Notes Images

Gansurhinus
Gansurhinus
Gansurhinus is an extinct genus of moradisaurine captorinid known from the Middle Permian of China. It was first named by Robert R. Reisz, Jun Liu, Jin-Ling Li and Johannes Müller in 2011 and the type species is Gansurhinus qingtoushanensis....


Gen. et sp.

Valid
  • Reisz
    Robert R. Reisz
    Robert Rafael Reisz is a Canadian paleontologist and specialist in the study of early amniote and tetrapod evolution.Robert Reisz was born August 27, 1947, in Oradea, Romania. He received his B.Sc. , M.Sc. and Ph.D. from McGill University as Robert L. Carroll’s first doctoral graduate...

  • Liu
  • et al.

Middle Permian

A moradisaurine captorinid
Captorhinidae
Captorhinidae is one of the earliest and most basal reptile families.-Description:...

.

Newly named turtles

Name Novelty Status Authors Age Unit Location Notes Images

Gilmoremys
Gilmoremys
Gilmoremys is an extinct genus of softshell turtle which lived during the late Cretaceous of North Dakota, Montana and Wyoming, USA. It is known from five skulls, a mandible and an incomplete postcranial skeleton...


Gen. et comb.

Valid
  • Joyce
  • Lyson

Maastrichtian

Lance Formation
Lance Formation
The Lance Formation is a division of Late Cretaceous rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas...

 

Hell Creek Formation
Hell Creek Formation
The Hell Creek Formation is an intensely-studied division of Upper Cretaceous to lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana...


A softshell turtle
Trionychidae
Trionychidae is a taxonomic family which comprises a number of turtle genera commonly known as softshells. They are also sometimes called pancake turtles. Softshells consist of some of the world's largest fresh water turtles, though many can adapt to living in highly brackish areas. Members of this...

, new genus for "Aspideretes
Aspideretes
Aspideretes is a genus of soft-shelled turtles . It has been found to be so closely related to the Burmese Peacock Softshell that it is probably better united with this species under Nilssonia, the older name...

" lancensis (Gilmore, 1928).

Hoyasemys
Hoyasemys
Hoyasemys is an extinct genus of basal eucryptodiran freshwater turtle from Lower Cretaceous deposits of Cuenca Province, Spain. It is known from the holotype MCCM-LH 84, a nearly complete and articulated skeleton including the skull...


Gen. et sp.

In press
  • Pérez-García
  • Fuente
  • Ortega

Late Barremian

Calizas de La Huérguina Formation

A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 eucryptodiran.

Patagoniaemys
Patagoniaemys
Patagoniaemys is an extinct genus of stem turtle which existed in central Patagonia, Chubut Province, Argentina during the late Cretaceous epoch . It is known from skull fragments and several postcranial elements including a nearly complete vertebral column recovered from the La Colonia Formation....


Gen. et sp.

Valid
  • Sterli
  • De la Fuente

Campanian - Maastrichtian

La Colonia Formation
La Colonia Formation
The La Colonia Formation is a geological formation in Argentina whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation....


A stem
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 turtle
Turtle
Turtles are reptiles of the order Testudines , characterised by a special bony or cartilaginous shell developed from their ribs that acts as a shield...

.

Selenemys
Selenemys
Selenemys is an extinct genus of pleurosternid turtle from the Late Jurassic of Central West of Portugal. It is known from several specimens recovered from the Lusitanian Basin, dating to the upper Kimmeridgian age...


Gen. et sp.

Valid
  • Pérez-García
  • Ortega

Upper Kimmeridgian

Lusitanian Basin

A pleurosternid.

Newly named basal diapsids

Name Novelty Status Authors Age Unit Location Notes Images

Orovenator
Orovenator
Orovenator is an extinct genus of diapsid from Lower Permian deposits of Oklahoma, USA. It is known from two partial skulls from the Richards Spur locality in Oklahoma...


Gen. et sp.

In press
  • Reisz
    Robert R. Reisz
    Robert Rafael Reisz is a Canadian paleontologist and specialist in the study of early amniote and tetrapod evolution.Robert Reisz was born August 27, 1947, in Oradea, Romania. He received his B.Sc. , M.Sc. and Ph.D. from McGill University as Robert L. Carroll’s first doctoral graduate...

  • Modesto
  • Scott

Sakmarian

The oldest and most basal neodiapsid
Neodiapsida
Neodiapsida is a clade or major branch of the reptilian family tree, that includes all Diapsids apart from some early primitive types known as Araeoscelidia....

.

Newly named ichthyosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Sveltonectes
Sveltonectes
Sveltonectes is an extinct genus of ophthalmosaurid ichthyosaur known from Ul’yanovsk region, western Russia.-Description:Sveltonectes is known from the holotype IRSNB R269, an almost complete three-dimensionally preserved skeleton which preserved partial skull...


Gen. et sp.

Valid
  • Fischer
  • Masure
  • et al.

Barremian

An ophthalmosaurid.

Newly named saurosphargids

Name Novelty Status Authors Age Unit Location Notes Images

Sinosaurosphargis
Sinosaurosphargis
Sinosaurosphargis is an extinct genus of non-sauropterygian marine reptile known from the lower Middle Triassic of Yangmazhai, Luoping County of Yunnan Province, southwestern China...


Gen. et sp.

Valid
  • Li
  • Rieppel
  • et al.

Middle Anisian

Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....


An incertae sedis
Incertae sedis
, is a term used to define a taxonomic group where its broader relationships are unknown or undefined. Uncertainty at specific taxonomic levels is attributed by , , and similar terms.-Examples:*The fossil plant Paradinandra suecica could not be assigned to any...

.

Newly named sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Dianopachysaurus
Dianopachysaurus
Dianopachysaurus is an extinct genus of pachypleurosaur known from the lower Middle Triassic of Yunnan Province, southwestern China. It was found in the Middle Triassic Lagerstatte of the Guanling Formation. It was first named by Jun Liu, Olivier Rieppel, Da-Yong Jiang, Jonathan C...


Gen. et sp.

Valid
  • Liu
  • Rieppel
  • et al.

Middle Anisian

Guanling Formation
Guanling Formation
The Guanling Formation is a Mesozoic geologic formation. Plesiosaur remains are among the fossils that have been recovered from its strata....


A keichousaurid pachypleurosaur
Pachypleurosaur
Pachypleurosaurs were primitive sauropterygian reptiles that vaguely resembled aquatic lizards, and were limited to the Triassic period. They were elongate animals, ranging in size from 20 cm to about a meter in length, with small heads, long necks, paddle-like limbs, and long deep tails. The...

.

Hauffiosaurus tomistomimus

Species

Valid
  • Benson
  • Ketchum
  • et al.

Early Toarcian

Whitby Limestone Formation

A second species of Hauffiosaurus.

Marmornectes
Marmornectes
Marmornectes is an extinct genus of pliosaurid known from the Bedfordshire of United Kingdom.-Description:Marmornectes is known from the holotype BEDFM 1999.201, articulated partial skeleton which preserved the skull...


Gen. et sp.

Valid
  • Ketchum
  • Benson

Callovian

Oxford Clay Formation

A pliosaurid.

Lusonectes
Lusonectes
Lusonectes is an extinct genus of plesiosaurid from Lower Jurassic deposits of Alhadas, Portugal. It is known from the holotype MG33, a partial skull and articulated mandible. It was found by Henri Émile Sauvage, a french paleontologist, in the 19th century from the São Gião Formation, near...


Gen. et sp.

In press
  • Smith
  • Araújo
  • Mateus

Toarcian

São Gião Formation

A plesiosaurid plesiosaur.

Westphaliasaurus
Westphaliasaurus
Westphaliasaurus is an extinct genus of plesiosaurid from Lower Jurassic deposits of Westfalia, northwestern Germany. It is known from a nearly complete and articulated skeleton missing the skull and about 38% of the upper neck vertebrae...


Gen. et sp.

Valid
  • Schwermann
  • Sander

Pliensbachian

Höxter district
Höxter (district)
Höxter is a Kreis in the east of North Rhine-Westphalia, Germany. Neighboring districts are Holzminden, Northeim, Kassel, Waldeck-Frankenberg, Hochsauerland, Paderborn, and Lippe.-History:...


A plesiosaurid plesiosaur.

Zarafasaura
Zarafasaura
Zarafasaura is an extinct genus of elasmosaurid known from the Oulad Abdoun Basin of Morocco.-Description:Zarafasaura is known from the holotype OCP-DEK/GE 315, an articulated incomplete dorsoventrally crushed skull and mandible and from the paratype OCP-DEK/GE 456, a complete mandible...


Gen. et sp.

Valid
  • Vincent
  • Bardet
  • et al.

Maastrichtian

An elasmosaurid
Elasmosauridae
Elasmosauridae was the family of plesiosaurs. They had the longest necks of the plesiosaurs and survived from the Late Triassic to the end of the Cretaceous. They had a diet of fish and shelless cephalopods.-Size:...

 plesiosaur.

Newly named lizards

Name Novelty Status Authors Age Unit Location Notes Images

Brasiliguana
Brasiliguana
Brasiliguana is a genus of iguanian lizard which lived during the late Cretaceous period in what is now Brazil. It is known from the holotype MN 7230-V, an isolated left maxilla with partially preserved teeth, which was found in the Upper Cretaceous Adamantina Formation, part of the Bauru...


Gen. et sp.

Valid
  • Nava
  • Martinelli

Turonian/Santonian

Adamantina Formation
Adamantina Formation
The Adamantina Formation is a geological formation in Brazil whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation...


A iguania
Iguania
Iguania is the suborder of Squamata that contains the iguanas, chameleons, agamids, and "New World lizards" such as anoles and Phrynosomatidae...

n lizard.

Cryptolacerta
Cryptolacerta
Cryptolacerta is an extinct genus of lacertibaenian lizard which lived during the Eocene epoch in what is now Germany. It is known from a nearly complete and articulated skeleton including the skull, which was found in the Messel locality of Germany. Cryptolacerta was named by Johannes Müller,...


Gen. et sp.

Valid
  • Müller
  • Hipsley
  • et al.

Lutetian

Messel pit
Messel pit
The Messel Pit is a disused quarry near the village of Messel, about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of fossils, it has significant geological and scientific importance...


A amphisbaenia
Amphisbaenia
The Amphisbaenia are a usually legless suborder of squamates closely related to lizards and snakes. As many species possess a pink body coloration and scales arranged in rings, they have a superficial resemblance to earthworms. They are very poorly understood, due to their burrowing lifestyle...

n lizard.

Newly named basal archosauromorphs

Name Novelty Status Authors Age Unit Location Notes Images

Archeopelta
Archeopelta
Archeopelta is an extinct genus of carnivorous archosauriform from the late Middle or early Late Triassic period . It is a doswelliid which lived in what is now southern Brazil. It is known from the holotype CPEZ-239a, which consists of partial skeleton and braincase...


Gen. et sp.

Valid
  • Desojo
  • Ezcurra
  • Schultz

Late Ladinian/Early Carnian

Santa Maria Formation
Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a late Ladinian – early Carnian age , and is notable for its fossils of early dinosaurs, including the herrerasaur Staurikosaurus, the basal saurischian Teyuwasu, and the basal sauropodomorph...


A doswelliid
Doswelliidae
Doswelliidae is a family of carnivore archosauriforms. Doswelliids existed in North and South America during the Middle–Late Triassic period and were among the most derived non-archosaurian archosauriforms. The family was named by R. E. Weems in 1980 and it has been placed in its own suborder,...


Augustaburiania
Augustaburiania
Augustaburiania is an extinct genus of tanystropheid archosauromorph which lived during the lower Triassic of what is now Eastern Europe. It is recovered from the Don River Basin in western Russia. It was named by Sennikov in 2011 and the type species is Augustaburiania vatagini. Augustaburiania is...


Gen. et sp.

Valid
  • Sennikov

Early Triassic

Don River Basin
Don River (Russia)
The Don River is one of the major rivers of Russia. It rises in the town of Novomoskovsk 60 kilometres southeast from Tula, southeast of Moscow, and flows for a distance of about 1,950 kilometres to the Sea of Azov....


The earliest tanystropheid.

Protanystropheus
Protanystropheus
Protanystropheus is an extinct genus of tanystropheid archosauromorph which lived during the Triassic of what is now central and western Europe. It was named by Sennikov in 2011 and the type species is Protanystropheus antiquus. P. antiquus was previously referred to as Tanystropheus antiquus, and...


Gen. et comb.

Valid
  • Sennikov

Middle - Late Triassic

A tanystropheid previously referred to Tanystropheus
Tanystropheus
Tanystropheus , was a 6 metre long reptile that dated from the Middle Triassic period. It is recognisable by its extremely elongated neck, which measured 3 metres long - longer than its body and tail combined. Despite this length, it had only ten neck vertebrae, each quite long...

.

Smok
Smok (archosaur)
Smok is an extinct genus of large carnivorous archosaur. It lived during the latest Triassic period in what is now Lisowice, southern Poland. The type species is Smok wawelski and was named in 2011...


Gen. et sp.

In press
  • Niedźwiedzki
  • Sulej
  • Dzik

Latest Norian/Early Rhaetian

Lipie Śląskie clay-pit

The largest known predatory archosaur
Archosaur
Archosaurs are a group of diapsid amniotes whose living representatives consist of modern birds and crocodilians. This group also includes all extinct non-avian dinosaurs, many extinct crocodilian relatives, and pterosaurs. Archosauria, the archosaur clade, is a crown group that includes the most...

 from the Late Triassic and Early Jurassic of central Europe.

Newly named crurotarsans

Name Novelty Status Authors Age Unit Location Notes Images

Acherontisuchus
Acherontisuchus
Acherontisuchus is an extinct genus of dyrosaurid neosuchian from Middle to Late Paleocene deposits of Colombia. The only known species is A. guajiraensis, whose name means "Acheron crocodile of the Guajira Peninsula".-Description:...


Gen. et sp.

Valid
  • Hastings
  • Bloch
  • Jaramillo

Middle/Late Paleocene

Cerrejón Formation
Cerrejón Formation
The Cerrejón Formation is a geologic formation in Colombia dating back to the Middle-Late Paleocene. It is found in the El Cerrejón subbasin of the Racheria basin of La Guajira. The formation consists of bituminous coal fields that are an important economic resource. Coal from the Cerrejón...


A dyrosaurid
Dyrosauridae
Dyrosauridae is a family of extinct neosuchian crocodyliforms that lived from the Late Cretaceous to the Eocene. Fossils of this group have been found in almost every continent, specifically Africa, Asia, Europe, North America and South America....

.


Arenysuchus
Arenysuchus
Arenysuchus is an extinct genus of crocodyloid from Late Cretaceous deposits of north Spain. It is known from the holotype MPZ ELI-1, a partial skull from Elías site and from the referred material MPZ2010/948, MPZ2010/949, MPZ2010/950 and MPZ2010/951, four teeth from Blasi 2 site...


Gen. et sp.

Valid
  • Puértolas
  • Canudo
  • Cruzado-Caballero

Maastrichtian

Tremp Formation
Tremp Formation
The Tremp Formation is a geological formation in Huesca, Spain whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation....


A stem crocodyloid
Crocodyloidea
Crocodyloidea is a superfamily of crocodilians that evolved in the Late Cretaceous period. Cladistically, it is defined as Crocodylus niloticus and all crocodylians more closely related to C. niloticus than to either Alligator mississippiensis or Gavialis gangeticus .- Phylogeny :Cladogram after...

.

Caipirasuchus
Caipirasuchus
Caipirasuchus is an extinct genus of sphagesaurid notosuchian known from the Late Cretaceous of São Paulo State, southeastern Brazil.-Discovery:...


Gen. et sp.

Valid
  • Iori
  • Carvalho

Turonian/Santonian

Adamantina Formation
Adamantina Formation
The Adamantina Formation is a geological formation in Brazil whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation...


A sphagesaurid.

Campinasuchus
Campinasuchus
Campinasuchus is an extinct genus of baurusuchid mesoeucrocodylian which existed in the Minas Gerais State, Brazil during the late Cretaceous period . It is known from four partial skulls from the Honorópolis District of the Adamantina Formation, Bauru Basin. The specimens were collected at Três...


Gen. et sp.

Valid
  • Carvalho
  • Teixeira
  • et al.

Turonian/Santonian

Adamantina Formation
Adamantina Formation
The Adamantina Formation is a geological formation in Brazil whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation...


A baurusuchid.

Decuriasuchus
Decuriasuchus
Decuriasuchus is an extinct genus of prestosuchid rauisuchian from the Middle Triassic period . It is a carnivorous archosaur that lived in what is now southern Brazil. It was first named by Marco Aurélio G. França, Jorge Ferigolo and Max C. Langer in 2011 and the type species is Decuriasuchus...


Gen. et sp.

Valid
  • França
  • Ferigolo
  • Langer

Ladinian

Santa Maria Formation
Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a late Ladinian – early Carnian age , and is notable for its fossils of early dinosaurs, including the herrerasaur Staurikosaurus, the basal saurischian Teyuwasu, and the basal sauropodomorph...


Skeletons indicate possible group behavior.

Neptunidraco
Neptunidraco
Neptunidraco is an extinct genus of carnivorous marine crocodyliform which lived during the middle Jurassic period in what is now northeastern Italy. It is known from a partial skeleton recovered from the nodular limestone of the Rosso Ammonitico Veronese Formation...


Gen. et sp.

Valid
  • Cau
  • Fanti

late Bajocian/early Bathonian

Rosso Ammonitico Veronese Formation

The oldest known metriorhynchid
Metriorhynchidae
Metriorhynchidae is an extinct family of metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. Metriorhynchids are fully aquatic crocodyliforms. Their forelimbs were small and paddle-like, and unlike living crocodilians,...

.

Pepesuchus
Pepesuchus
Pepesuchus is an extinct genus of carnivorous metasuchian from the late Cretaceous period. It is a peirosaurid which lived during the Campanian and Maastrichtian stages of the Late Cretaceous in what is now state of São Paulo, Brazil.-Etymology:...


Gen. et sp.

Valid
  • Campos
  • Oliveira
  • et al.

Campanian/Maastrichtian

Presidente Prudente Formation

A peirosaurid.

Pissarrachampsa
Pissarrachampsa
Pissarrachampsa is an extinct genus of baurusuchid mesoeucrocodylian from the Late Cretaceous of Brazil...


Gen. et sp.

Valid
  • Montefeltro
  • Larsson
  • Langer

Campanian/Maastrichtian

Vale do Rio do Peixe Formation

A baurusuchid. The type species is Pissarrachampsa sera.

Theriosuchus grandinaris
Theriosuchus
Theriosuchus is an extinct genus of atoposaurid mesoeucrocodylian from the Late Jurassic of Europe, the Early Cretaceous of Asia and the Late Cretaceous of Romania. The five species are T. pusillus from England , T. ibericus from Spain , T. guimarotae from Portugal , T. sympiestodon from Romania...


Species

Valid
  • Lauprasert
  • Laojumpon
  • et al.

Berriasian/Barremian

Khua Formation

A fifth species of Theriosuchus.

Newly named birds

Name Novelty Status Authors Age Unit Location Notes Images

Bohaiornis
Bohaiornis
Bohaiornis is an extinct genus of enantiornithine bird. Fossils have been found from the Lower Cretaceous Yixian Formation of western Liaoning, China. The type species B. guoi was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved...


Gen. et sp.

Valid
  • Hu
  • Li
  • et al.

Barremian/Aptian

Yixian Formation
Yixian Formation
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...


An enantiornithine.

Hoazinavis
Hoazinavis
Hoazinavis is an extinct genus of early hoatzin from Late Oligocene and Early Miocene deposits of Brazil. It was collected in 2008 from the Tremembé Formation of São Paulo, Brazil. It was first named by Gerald Mayr, Herculano Alvarenga and Cécile Mourer-Chauviré in 2011 and the type species is...


Gen. et sp.

In press
  • Mayr
  • Alvarenga
  • Mourer-Chauviré

Late Oligocene/Early Miocene

Tremembé Formation

An early hoatzin
Hoatzin
The Hoatzin , also known as the Hoactzin, Stinkbird, or Canje Pheasant, is a species of tropical bird found in swamps, riverine forest and mangrove of the Amazon and the Orinoco delta in South America...

.

Jamna

Gen. et sp.

In press
  • Bocheński
  • Tomek
  • et al.

Oligocene

Carpathian flysch

A passerine
Passerine
A passerine is a bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds or, less accurately, as songbirds, the passerines form one of the most diverse terrestrial vertebrate orders: with over 5,000 identified species, it has roughly...

.

Kaiika
Kaiika
Kaiika is an extinct genus of basal penguin from Early Eocene deposits of South Canterbury, New Zealand. It is known only from a single humerus. It was found in 1998 by Dr. Phillip Maxwell, a paleontologist and stratigrapher, from the Kauru Formation of the Canterbury Basin, near the Waihao...


Gen. et sp.

Valid
  • Fordyce
  • Thomas

Early Eocene

Kauru Formation

A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 penguin
Penguin
Penguins are a group of aquatic, flightless birds living almost exclusively in the southern hemisphere, especially in Antarctica. Highly adapted for life in the water, penguins have countershaded dark and white plumage, and their wings have become flippers...

.

Lavocatavis
Lavocatavis
Lavocatavis is an extinct genus of phorusrhacoid or "terror bird" from the Eocene of Algeria. A fossilized femur was described from the Glib Zegdou Formation in 2011 and is the only known specimen of Lavocatavis. The species was designated L...


Gen. et sp.

In press
  • Mourer-Chauviré
  • Tabuce
  • et al.

Eocene

Glib Zegdou Formation

A phorusrhacoid.

Mystiornis
Mystiornis
Mystiornis is an extinct genus of bird which existed in what is now western Siberia, Russia during the early Cretaceous period . It is known from an isolated metatarsus found in the Shestakovo-1 locality of southern Western Siberia. It was named by Evgeny N. Kurochkin, Nikita V. Zelenkov, Alexandr...


Gen. et sp.

Valid
  • Kurochkin
  • Zelenkov
  • et al.

Barremian/Aptian

Shestakovo-1 locality
,  Russia

Nambashag
Nambashag
Nambashag is an extinct genus of early cormorant from Late Oligocene and Early Miocene deposits of South Australia. The genus was first named by Trevor H. Worthy in 2011 and the type species is Nambashag billerooensis. A second species, N. microglaucus, was also named. The type species is known...


Gen. et 2 sp.

Valid
  • Worthy

Late Oligocene/Early Miocene

Etadunna Formation 

Namba Formation

A cormorant
Cormorant
The bird family Phalacrocoracidae is represented by some 40 species of cormorants and shags. Several different classifications of the family have been proposed recently, and the number of genera is disputed.- Names :...

.

Qiliania
Qiliania
Qiliania is an extinct genus of early bird from the lower Cretaceous . It is a enantiornithine which lived in what is now Gansu Province, north-western China. It is known from two incomplete, semi-articulated and three-dimensionally preserved skeletons, which was found in the Xiagou Formation of...


Gen. et sp.

Valid
  • Ji
  • Atterholt
  • et al.

Aptian

Xiagou Formation
Xiagou Formation
The Xiagou Formation is the middle unit of the Xinminpu Group. It is named for its type site in Xiagou, in the Changma Basin of Gansu Province, northwestern China and is considered Early Cretaceous in age. It is known outside the specialized world of Chinese geology as the site of a lagerstätte...


An enantiornithine.

Samrukia
Samrukia
"Samrukia" is an as-of-yet unpublished name for a genus of large Cretaceous archosaurs known only from a single lower jaw discovered in Kazakhstan. The holotype and only known specimen was collected from the Santonian-Campanian age Bostobynskaya Formation in Kyzylorda District...


Gen. et sp.

In press
  • Naish
  • Dyke
  • et al.

Late Cretaceous

Kyzylorda District

A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 ornithuromorph.

Sanshuiornis
Sanshuiornis
Sanshuiornis is an extinct genus of bird from Middle Eocene deposits of Guangdong Province, south China. It is known from the holotype IVPP V18116 an articulated complete right leg and foot. It was found from the black oil shales of the Huayong Formation of the Sanshui Basin, near Foshan...


Gen. et sp.

In press
  • Wang
  • Mayr
  • et al.

Middle Eocene

Huayong Formation

Newly named non-avian dinosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Acristavus
Acristavus
Acristavus is a genus of hadrosaurine hadrosaurid dinosaur. Fossils have been found from the Campanian Two Medicine Formation in Montana and Wahweap Formation in Utah. The type species A. gagslarsoni was named in 2011. Unlike nearly all hadrosaurids except Edmontosaurus, Acristavus lacked...


Gen. et sp.

Valid
  • Gates
  • Horner
  • et al.

Campanian

Two Medicine Formation
Two Medicine Formation
The Two Medicine Formation is a geologic formation, or rock body, that was deposited between 83.5 ± 0.7 Ma to 70.6 ± 0.6 Ma , during Campanian time, and is located in northwestern Montana...

 

Wahweap Formation
Wahweap Formation
The Wahweap Formation of the Grand Staircase-Escelante National Monument is a geological formation in southern Utah and northern Arizona, around the Lake Powell region, whose strata date back to the Late Cretaceous...


A brachylophosaurin hadrosaurine.


Ahshislepelta
Ahshislepelta
Ahshislepelta is a genus of herbivorousankylosaurine ankylosaurid dinosaur from Late Cretaceous deposits of San Juan Basin, New Mexico....


Gen. et sp.

Valid
  • Burns
  • Sullivan

Late Campanian

Kirtland Formation
Kirtland Formation
The Kirtland Formation is a sedimentary geological formation. It is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that existed on the inland seashore of North America, in the late Cretaceous period. It overlies the Fruitland Formation...


An ankylosaurine ankylosaurid
Ankylosauridae
An ankylosaurid is a member of the Ankylosauridae family of armored dinosaurs that evolved 125 million years ago and became extinct 65 million years ago during the Cretaceous-Tertiary extinction event...

.

Albinykus
Albinykus
Albinykus is a genus of alvarezsaurid dinosaur. Fossils have been found from Late Cretaceous-age deposits in the Gobi Desert of Mongolia. The type species A. baatar was named by Sterling J. Nesbitt, Julia A. Clarke, Alan H. Turner and Mark A...


Gen. et sp.

Valid
  • Nesbitt
  • Clarke
  • et al.

Late Cretaceous

An alvarezsaurid
Alvarezsauridae
Alvarezsauridae is an enigmatic family of small, long-legged running dinosaurs. Although originally thought to represent the earliest known flightless birds, a consensus of recent work suggests that they are primitive members of the Maniraptora. Other work found them to be the sister group to the...

 theropod.

Amtocephale
Amtocephale
Amtocephale is a genus of pachycephalosaurid dinosaur from early Late Cretaceous deposits of southern Gobi Desert, Mongolia....


Gen. et sp.

Valid
  • Watabe
  • Tsogtbaatar
  • Sullivan

Turonian-Santonian

Baynshire Formation

A pachycephalosaurid
Pachycephalosauria
Pachycephalosauria is a clade of ornithischian dinosaurs. Well-known genera include Pachycephalosaurus, Stegoceras, Stygimoloch, and Dracorex. Most lived during the Late Cretaceous Period, in what is now North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls...

.

Angolatitan
Angolatitan
Angolatitan is a genus of long-necked, plant eating sauropod dinosaur from Angola. It is known from a partial right arm including a shoulder blade, upper arm , the two bones of the lower arm , and three metacarpals...


Gen. et sp.

Valid
  • Mateus
  • Jacobs
  • et al.

Late Turonian

Itombe Formation

A somphospondylian sauropod.

Arcusaurus
Arcusaurus
Arcusaurus is an extinct genus of sauropodomorph dinosaur from the Early Jurassic of South Africa.Arcusaurus was first named by Adam Yates, Matthew Bonnan and Johann Neveling in 2011 and the type species is Arcusaurus pereirabdalorum. The generic name is derived from Latin arcus, "rainbow", a...


Gen. et sp.

Valid
  • Yates
  • Bonnan
  • Neveling

Lower Jurassic

Elliot Formation
Elliot Formation
The Elliot Formation is a geological formation dating to roughly 210 to 190 million years ago and covering the Norian to Sinemurian stages. The Elliot Formation is found in South Africa and Lesotho and is a member of the Stormberg Group. It consists mainly of limestone, sandstone, and mudstone...


Basal Sauropodomorph

Atacamatitan
Atacamatitan
Atacamatitan is a genus of sauropod dinosaur. It is a titanosaur which lived during the late Cretaceous in what is now northern Chile. It is known from the holotype SGO-PV-961, which includes two dorsal vertebrae, caudal vertebrae, ribs, a possible sternum, part of an upper arm, the right femur,...


Gen. et sp.

Valid
  • Kellner
  • Rubilar-Rogers
  • et al.

Late Cretaceous

Tolar Formation

A titanosaur
Titanosaur
Titanosaurs were a diverse group of sauropod dinosaurs, which included Saltasaurus and Isisaurus. It includes some of the heaviest creatures ever to walk the earth, such as Argentinosaurus and Paralititan — which some believe have weighed up to 100 tonnes...

.

Brontomerus
Brontomerus
Brontomerus is a genus of camarasauromorph sauropod which lived during the early Cretaceous . It was named in 2011 and the type species is Brontomerus mcintoshi...


Gen. et sp.

Valid
  • Taylor
  • Wedel
  • Cifelli

Aptian/Albian

Cedar Mountain Formation
Cedar Mountain Formation
The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation . It is composed of non-marine sediments, that is, sediments deposited in rivers, lakes and on flood plains...


A camarasauromorph sauropod.

Daemonosaurus
Daemonosaurus
Daemonosaurus is a genus of theropod dinosaur from the Late Triassic of New Mexico. Fossils have been found from deposits in the Chinle Formation, which is latest Triassic in age. While theropods had diversified into several specialized groups by this time, Daemonosaurus is a basal theropod that...


Gen. et sp.
  • Sues
  • Nesbitt
  • Berman
  • Henrici

Late Triassic

Chinle Formation
Chinle Formation
The Chinle is a geologic formation that is spread across the U.S. states of northern Arizona, Nevada, Utah, western New Mexico, and western Colorado. The Chinle is controversially considered to be synonymous to the Dockum Group in eastern Colorado, eastern New Mexico, southwestern Kansas, the...


A primitive theropod.

Delapparentia
Delapparentia
Delapparentia is a genus of iguanodont dinosaur from the Early Cretaceous period of Galve, Teruel Province, Spain.The type specimen, MPT/I.G, was found in the Spring of 1958 by amateur palaeontologist José María Herrero Marzo...


Gen. et sp.

Valid
  • Ruiz-Omeñaca

Early Barremian

Camarillas Formation
Camarillas Formation
The Camarillas Formation is a geological formation in Teruel and La Rioja, Spain whose strata date back to the Early Cretaceous. Dinosaur remains have been recovered from the formation.-Vertebrate paleofauna:* cf Pleurocoelus sp....


An iguanodontoid.

Demandasaurus
Demandasaurus
Demandasaurus is a genus of rebbachisaurid sauropod from early Cretaceous deposits of Spain. Demandasaurus is known from an incomplete but associated skeleton that includes cranial and postcranial remains. It was collected from the Castrillo la Reina Formation in Burgos Province of Spain...


Gen. et sp.

Valid
  • Fernández-Baldor
  • Canudo
  • et al.

Late Barremian-early Aptian

Castrillo la Reina Formation

A rebbachisaurid
Rebbachisauridae
Rebbachisauridae is a family of sauropod dinosaurs known from fragmentary fossil remains from the Cretaceous of South America, Africa, and Europe.-Taxonomy:...

 sauropod.

Drusilasaura
Drusilasaura
Drusilasaura is an extinct genus of possible lognkosaurian titanosaur sauropod which lived during the late Cretaceous of Santa Cruz Province of southern Patagonia, Argentina....


Gen. et sp.

Valid
  • Navarrete
  • Casal
  • Martínez

Cenomanian/Turonian

Bajo Barreal Formation
Bajo Barreal Formation
The Bajo Barreal Formation is a geological formation in Chubut, Argentina whose strata date back to the Late Cretaceous.-Dinosaurs:Dinosaur remains are among the fossils that have been recovered from the formation...


A possible lognkosauria
Lognkosauria
Lognkosauria is a group of giant long-necked sauropod dinosaurs within the clade Titanosauria, and includes some of the largest and heaviest dinosaurs to ever walk the earth.-Defining characteristics:...

n titanosaur.

Eodromaeus
Eodromaeus
Eodromaeus is an extinct genus of basal theropod dinosaur known from the Late Triassic period of Argentina. It has been cited by Sereno as resembling a supposed common ancestor to all dinosaurs, the "Eve" of the dinosaurs....


Gen. et sp.

Valid
  • Martinez
  • Sereno
  • et al.

Middle Carnian

Ischigualasto Formation

A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 theropod.

Epichirostenotes
Epichirostenotes
Epichirostenotes is a genus of oviraptorosaurian dinosaur from the late Cretaceous. Epichirostenotes is known from an incomplete skeleton found in 1923 at the Horseshoe Canyon Formation, in strata dated to about 72 million years ago. It was first named by Robert M. Sullivan, Steven E. Jasinski and...


Gen. et sp.

Valid
  • Sullivan
  • Jasinski
  • Van Tomme

Late Campanian

Horseshoe Canyon Formation
Horseshoe Canyon Formation
The Horseshoe Canyon Formation is part of the Edmonton Group and is up to 230m in thickness. It is Late Campanian to Early Maastrichtian in age and is composed of mudstone, sandstone, and carbonaceous shales...


A caenagnathid.

Haya
Haya (dinosaur)
Haya is an extinct genus of basal ornithopod dinosaur known from Mongolia.-Description:Haya is known from several well-preserved specimens which collected in the Khugenetslavkant locality by the joint Mongolian Academy of Sciences from 2002 to 2007, from the Javkhlant Formation. The locality...


Gen. et sp.

Valid
  • Makovicky
  • Kilbourne
  • et al.

Late Cretaceous

Javkhlant Formation

Basal ornithopod with many gastroliths.

Jaklapallisaurus
Jaklapallisaurus
Jaklapallisaurus is a genus of herbivore plateosaurian sauropodomorph dinosaur. It is a plateosaurid which lived during the Late Triassic period in what is now Andhra Pradesh, central India...


Gen. et. sp.

Valid
  • Novas
  • Ezcurra
  • et al.

Late Norian-earliest Rhaetian

Upper Maleri Formation
Upper Maleri Formation
The Upper Maleri Formation is a sedimentary rock formation found in Andhra Pradesh, India. It is one of the formations of the Pranhita–Godavari Basin. It is of late Norian and possibly earliest Rhaetian ages , and is notable for its fossils of early dinosaurs.-Vertebrate fauna:Angistorhinus sp., cf...

 

Lower Dharmaram Formation
Lower Dharmaram Formation
The Lower Dharmaram Formation is a sedimentary rock formation found in Andhra Pradesh, India. It is one of the formations of the Pranhita–Godavari Basin. It is of latest Norian and Rhaetian ages , and is notable for its fossils of early dinosaurs.-Vertebrate fauna:cf. Paratypothorax, cf...


A plateosaurid
Plateosauridae
Plateosauridae is a family of plateosaurian sauropodomorphs. Plateosaurids were early sauropodomorph dinosaurs which existed in Asia, Europe and South America during the Late Triassic period. Although several dinosaurs have been classified as plateosaurids over the years, a 2007 study by Adam M....

.

Koreaceratops
Koreaceratops
Koreaceratops is a genus of basal ceratopsian dinosaur discovered in Albian-age Lower Cretaceous rocks of South Korea. It is based on KIGAM VP 200801, an articulated series of 36 caudal vertebrae associated with partial hind limbs and ischia...


Gen. et sp.

Valid
  • Lee
  • Ryan
  • Kobayashi

Albian

"Tando beds"

A basal
Basal (phylogenetics)
In phylogenetics, a basal clade is the earliest clade to branch in a larger clade; it appears at the base of a cladogram.A basal group forms an outgroup to the rest of the clade, such as in the following example:...

 ceratopsia
Ceratopsia
Ceratopsia or Ceratopia is a group of herbivorous, beaked dinosaurs which thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, Yinlong downsi, lived between 161.2 and 155.7...

n.

Koreanosaurus
Koreanosaurus (ornithopod)
Koreanosaurus is a genus of ornithopod dinosaur. In May 2003 three fossils of Koreanosaurus were found in the Late Cretaceous-age Seonso Conglomerate from the southern coast of the Bi-bong Dinosaur Fossil Findspot, Boseong, Korean Peninsula: the holotype KDRC-BB2, a partial skeleton lacking the...


Gen. et sp.

Valid
  • Huh
  • Lee
  • et al.

?Santonian/Campanian

Seonso Conglomerate

A hypsilophodontid ornithopod.

Leonerasaurus
Leonerasaurus
Leonerasaurus is a basal genus of sauropodomorph dinosaur. Currently, there is only one species known, named L. taquetrensis by Diego Pol, Alberto Garrido and Ignacio A. Cerda in 2011. The fossil, an incomplete subadult individual, was found in the Las Leoneras Formation in Argentina. This...


Gen. et sp.

Valid
  • Pol
  • Garrido
  • Cerda

Pliensbachian/Toarcian

Las Leoneras Formation

An anchisaurian sauropodomorph.

Leyesaurus
Leyesaurus
Leyesaurus is an extinct genus of massospondylid sauropodomorph known from the San Juan Province, northwestern Argentina.-Description:...


Gen. et sp.

Valid
  • Apaldetti
  • Martinez
  • et al.

Lower Jurassic

Quebrada del Barro Formation
Quebrada del Barro Formation
The Quebrada del Barro Formation is a geological formation in South America. It dates back to the Late Triassic.-Vertebrate fauna:...


A massospondylid sauropodomorph.

Linhenykus
Linhenykus
Linhenykus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous of Inner Mongolia, China. It is the most basal known member of the Parvicursorinae. The genus gets its name from Linhe, a city near the site where fossils were first found and Greek onyx, "claw"...


Gen. et sp.

Valid
  • Xu
  • Sullivan
  • et al.

Campanian

Wulansuhai Formation

An alvarezsaurid.

Linhevenator
Linhevenator
Linhevenator is a genus of short-armed troodontid theropod dinosaur from the Late Cretaceous Bayan Mandahu Formation of Bayan Mandahu, Inner Mongolia, China....


Gen. et sp.

Valid
  • Xu
  • Tan
  • et al.

Campanian

Bayan Mandahu Formation
Bayan Mandahu Formation
The Bayan Mandahu Formation is a geological unit of "redbeds" located near the village of Bayan Mandahu in Inner Mongolia, China Asia and dates from the Late Cretaceous Period...


An short-armed troodontid
Troodontidae
Troodontidae is a family of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and scrappy and they have therefore been allied, at various times, with many dinosaurian lineages...

.

Manidens
Manidens
Manidens is a genus of heterodontosaurid dinosaur from the Middle Jurassic of Patagonia. Fossils have been found from the Cañadon Asfalto Formation in Chubut Province, Argentina, dating to the Bajocian.-Etymology:...


Gen. et sp.

Vald
  • Pol
  • Rauhut
  • Becerra

Aalenian/Early Bathonian

Cañadón Asfalto Formation
Cañadon Asfalto Formation
-Paleofauna:-References:* Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka : The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN 0-520-24209-2....


A heterodontosaurid.

Nambalia
Nambalia
Nambalia is a genus of basal sauropodomorph dinosaur. It lived during the Late Triassic period in what is now Andhra Pradesh, central India...


Gen. et. sp.

Valid
  • Novas
  • Ezcurra
  • et al.

Late Norian-earliest Rhaetian

Upper Maleri Formation
Upper Maleri Formation
The Upper Maleri Formation is a sedimentary rock formation found in Andhra Pradesh, India. It is one of the formations of the Pranhita–Godavari Basin. It is of late Norian and possibly earliest Rhaetian ages , and is notable for its fossils of early dinosaurs.-Vertebrate fauna:Angistorhinus sp., cf...


A basal sauropodomorph.

Narambuenatitan
Narambuenatitan
Narambuenatitan is a genus of lithostrotian titanosaur sauropod from late Cretaceous deposits of northern Patagonia of Argentina. Narambuenatitan is known from the holotype MAU-Pv-N-425, an incomplete skeleton. It was collected in 2005 and 2006 from the Anacleto Formation in northern Patagonia....


Gen. et sp.

Valid
  • Filippi
  • García
  • Garrido

lower-middle Campanian

Anacleto Formation
Anacleto Formation
The Anacleto Formation is a geologic formation with outcroppings in the Argentine Patagonian provinces of Mendoza, Río Negro, and Neuquén. It is the youngest formation within the Neuquén Group and belongs to the Río Colorado Subgroup...


A lithostrotian titanosaur.

Ojoraptorsaurus
Ojoraptorsaurus
Ojoraptorsaurus is a genus of oviraptorosaurian dinosaur from the late Cretaceous. Ojoraptorsaurus is known from the holotype SMP VP-1458 an incomplete pair of fused pubes found at the Naashoibito Member of the Ojo Alamo Formation dating from the early Maastrichtian, about 69 million years...


Gen. et sp.

Valid
  • Sullivan
  • Jasinski
  • Van Tomme

Lower Maastrichtian

Ojo Alamo Formation
Ojo Alamo Formation
The Ojo Alamo Formation is a geologic formation spanning the Mesozoic/Cenozoic boundary. Dinosaur remains are among the fossils that have been recovered from the formation, though all dinosaur remains come from the lowest part of the formation, the Naashoibito member The Ojo Alamo Formation is a...


A caenagnathid.

Osmakasaurus
Osmakasaurus
Osmakasaurus is a genus of herbivorous iguanodontian dinosaur. It is a basal iguanodontian which lived during the lower Cretaceous period in what is now Buffalo Gap of South Dakota, USA. It is known from the Chilson Member of the Lakota Formation. This genus was named by Andrew T. McDonald in 2011...


Gen. et comb.

Valid
  • McDonald

Valanginian

Lakota Formation
Lakota Formation
The Lakota Formation is a sequence of rocks from the Barremian epoch from Western North America. Located in South Dakota, the name is derived from the Lakota Native American tribe....


A new genus for "Camptosaurus" depressus (Gilmore, 1909).

Oxalaia
Oxalaia
Oxalaia is a genus of carnivorous theropod. It is a spinosaurine spinosaurid which lived during the late Cretaceous in what is now Brazil....


Gen. et sp.

Valid
  • Kellner
  • Azvedo
  • et al.

Cenomanian

Alcântara Formation
Alcântara Formation
The Alcântara Formation is a geological formation in northeastern Brazil whose strata date back to the Upper Cretaceous.-Fishes:-Crurotarsans:-Ornithodirans:...


Huge spinosaur 12–14 m long.

Pachyrhinosaurus perotorum

Species

In press
  • Fiorillo
  • Tykoski

Maastrichtian

Prince Creek Formation
Prince Creek Formation
The Prince Creek Formation is a geological formation in Alaska with strata from the early Maastrichtian stage of the Upper Cretaceous, dating to between 70 and 69 million years ago...


Third known species of Pachyrhinosaurus.

Pampadromaeus
Pampadromaeus
Pampadromaeus is an extinct genus of basal sauropodomorph dinosaurs known from the Triassic of Rio Grande do Sul, southern Brazil.-Discovery:...


Gen. et sp.

Valid
  • Cabreira
  • Schultz
  • et al.

Carnian

Santa Maria Formation
Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a late Ladinian – early Carnian age , and is notable for its fossils of early dinosaurs, including the herrerasaur Staurikosaurus, the basal saurischian Teyuwasu, and the basal sauropodomorph...


A very basal sauropodomorph.

Pamparaptor
Pamparaptor
Pamparaptor is an extinct genus of carnivorous deinonychosaur from the late Cretaceous period. It is a basal dromaeosaurid with troodontid-like pes which lived during the late Cretaceous period in what is now Neuquén province, Patagonia, Argentina. It is known from the holotype MUCPv-1163, an...


Gen. et sp.

Valid
  • Porfiri
  • Calvo
  • Santos

Turonian/Coniacian

Portezuelo Formation
Portezuelo Formation
The Portezuelo Formation is a geologic formation outcropping in the Mendoza, Río Negro and Neuquén provinces of Argentina. It is the fourth-oldest formation in the Neuquén Group and the older of the two formations in the Río Neuquén Subgroup...


A deinonychosaur.

Petrobrasaurus
Petrobrasaurus
Petrobrasaurus is a genus of herbivorous sauropod dinosaur. It is a titanosaur which lived during the upper Cretaceous period in what is now Rincón de los Sauces, Patagonia, Argentina. It is known from the holotype MAU-Pv-PH-449 — a partial disarticulated skeleton recovered from the Plottier...


Gen. et sp.

Valid
  • Filippi
  • Canudo
  • et al.

Santonian

Plottier Formation
Plottier Formation
The Plottier Formation is a geologic formation that outcrops in the Argentine Patagonian provinces of Río Negro and Neuquén. It is the younger of two formations belonging to the Río Neuquén Subgroup within the Neuquén Group, with the oldest rocks dating from the late Coniacian and its youngest...


A titanosaur.

Qiupalong
Qiupalong
Qiupalong is a genus of ornithomimid dinosaur. Qiupalong lived during the late Cretaceous period in what is now Henan Province, China. It is known from a single partial skeleton recovered from the Qiupa Formation...


Gen. et sp.

Valid
  • Xu
  • Kobayashi
  • et al.

Late Cretaceous

Qiupa Formation

An ornithomimid.

Ratchasimasaurus
Ratchasimasaurus
Ratchasimasaurus is a genus of non-hadrosaurid iguanodontian ornithopod dinosaur from Early Cretaceous Khok Kruat Formation of Nakhon Ratchasima Province in northeastern Thailand. One autapomorphy of Ratchasimasaurus is its elongated and flat ramus of the dentary...


Gen. et sp.

Valid
  • Shibata
  • Jintasakul
  • Azuma

Aptian

Khok Kruat Formation
Khok Kruat Formation
The Khok Kruat Formation is a rock formation found in northeastern Thailand. It is one of the formations of the Khorat Group. It is of Aptian stage , and is notable for its fossils of dinosaurs...


A non-hadrosaurid iguanodontian.

Sarahsaurus
Sarahsaurus
Sarahsaurus is a genus of basal sauropodomorph dinosaur which lived during the lower Jurassic period in what is now northeastern Arizona, USA....


Gen. et sp.

Valid
  • Rowe
  • Sues
    Hans-Dieter Sues
    Hans-Dieter Sues is a German-born paleontologist who is Senior Scientist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. He received his education at Johannes Gutenberg-Universität Mainz , University of Alberta, and...

  • Reisz
    Robert R. Reisz
    Robert Rafael Reisz is a Canadian paleontologist and specialist in the study of early amniote and tetrapod evolution.Robert Reisz was born August 27, 1947, in Oradea, Romania. He received his B.Sc. , M.Sc. and Ph.D. from McGill University as Robert L. Carroll’s first doctoral graduate...


Sinemurian/Pliensbachian

Kayenta Formation
Kayenta Formation
The Kayenta Formation is a geologic layer in the Glen Canyon Group that is spread across the Colorado Plateau province of the United States, including northern Arizona, northwest Colorado, Nevada, and Utah. This rock formation is particularly prominent in southeastern Utah, where it is seen in the...


A basal sauropodomorph

Siamodon
Siamodon
Siamodon is a genus of iguanodontian ornithopod dinosaur from Early Cretaceous deposits of northeastern Thailand. Siamodon is known from the holotype PRC-4, a well-preserved left maxilla and from the referred materials PRC-5, an isolated maxillary tooth and PRC-6, a braincase...


Gen. et sp.

Valid
  • Buffetaut
  • Suteethorn

Aptian

Khok Kruat Formation
Khok Kruat Formation
The Khok Kruat Formation is a rock formation found in northeastern Thailand. It is one of the formations of the Khorat Group. It is of Aptian stage , and is notable for its fossils of dinosaurs...


An advanced iguanodontian.

Stegoceras novomexicanum

Species

Valid
  • Jasinski
  • Sullivan

Late Campanian

Fruitland Formation
Fruitland Formation
The Fruitland Formation is a sedimentary geological formation containing layers of sandstone, shale, and coal. It was laid down in marshy delta conditions, with poor drainage and frequent flooding, under a warm, humid and seasonal climate...

 

Kirtland Formation
Kirtland Formation
The Kirtland Formation is a sedimentary geological formation. It is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that existed on the inland seashore of North America, in the late Cretaceous period. It overlies the Fruitland Formation...


Second known species of Stegoceras.

Talos

Gen. et sp.

Valid
  • Zanno
  • Varricchio
  • et al.

Late Campanian

Kaiparowits Formation
Kaiparowits Formation
The Kaiparowits Formation is a sedimentary rock formation found in the Kaiparowits Plateau in Grand Staircase-Escalante National Monument, in the southern part of Utah in the western United States. It is over 2800 feet thick, and is Campanian in age...


A troodontid theropod.

Tapuiasaurus
Tapuiasaurus
Tapuiasaurus is a genus of herbivorous sauropod dinosaur. It is a nemegtosaurid titanosaur which lived during the Lower Cretaceous period in what is now Brazil. Its fossils, including a partial skeleton with a nearly complete skull, have been recovered from the Quiricó Formation at Minas Gerais,...


Gen. et sp.

Valid
  • Zaher
  • Pol
  • et al.

Aptian

Quiricó Formation
Quiricó Formation
The Quiricó Formation is a geological formation in Brazil whose strata date back to the Lower Cretaceous. Many occurrences of fossils are reported in the lacustrine deposits of the Quiricó Formation.* Tapuiasaurus macedoi* Dastilbe moraesi...


A titanosaur.

Teratophoneus
Teratophoneus
Teratophoneus is a genus of carnivorous tyrannosaurid theropod dinosaur which lived during the late Cretaceous period in what is now Utah, USA. It is known from an incomplete skull and postcranial skeleton recovered from the Kaiparowits Formation. Teratophoneus was named by Thomas D. Carr, Thomas E...


Gen. et sp.

Valid
  • Carr
  • Williamson
  • et al.

Late Campanian

Kaiparowits Formation
Kaiparowits Formation
The Kaiparowits Formation is a sedimentary rock formation found in the Kaiparowits Plateau in Grand Staircase-Escalante National Monument, in the southern part of Utah in the western United States. It is over 2800 feet thick, and is Campanian in age...


A tyrannosaurine tyrannosaurid.

Thescelosaurus assiniboiensis

Species

Valid
  • Brown
  • Boyd
  • Russell

Late Maastrichtian

Frenchman Formation
Frenchman Formation
The Frenchman Formation is a division of Upper Cretaceous rocks found in Saskatchewan, Canada. More accurately described as Late Maastrichtian, these rocks contain the youngest of dinosaur genera, much like the Hell Creek Formation in the United States....


Third known species of Thescelosaurus.

Titanoceratops
Titanoceratops
Titanoceratops is a genus of herbivorous ceratopsian dinosaur. It was a giant chasmosaurine ceratopsian which lived during the Late Cretaceous period in what is now New Mexico, and the earliest known triceratopsin. It is known from the holotype OMNH 10165, a partial skeleton including...


Gen. et sp.

Valid
  • Longrich

Late Campanian

Fruitland
Fruitland Formation
The Fruitland Formation is a sedimentary geological formation containing layers of sandstone, shale, and coal. It was laid down in marshy delta conditions, with poor drainage and frequent flooding, under a warm, humid and seasonal climate...

 or Kirtland Formation
Kirtland Formation
The Kirtland Formation is a sedimentary geological formation. It is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that existed on the inland seashore of North America, in the late Cretaceous period. It overlies the Fruitland Formation...


A giant chasmosaurine
Chasmosaurinae
Chasmosaurinae is a subfamily of ceratopsid dinosaurs. Triceratops is a well-known example. They were one of the most successful groups of herbivores of their time. Chasmosaurines appeared in the early Campanian, and became extinct, along with all other non-avian dinosaurs, during the K-T extinction...

 ceratopsian from New Mexico.

Traukutitan
Traukutitan
Traukutitan is a genus of possible lognkosaurian titanosaur sauropod which lived during the late Cretaceous .Fossils of Traukutitan were recovered from the lowermost section of the Bajo de la Carpa Formation in North Patagonia of Argentina and described but not named in 1993 by Leonardo Salgado...


Gen. et sp.

Valid
  • Juárez Valieri
  • Calvo

Santonian

Bajo de la Carpa Formation
Bajo de la Carpa Formation
The Bajo de la Carpa Formation is a geologic formation that outcrops in Patagonia, in the provinces of Río Negro and Neuquén, Argentina. It is the first of two formations belonging to the Río Colorado Subgroup within the Neuquén Group...


A possible lognkosaurian titanosaur.

Uteodon
Uteodon
Uteodon is a genus of herbivorous iguanodontian dinosaur. It is a basal iguanodontian which lived during the upper Jurassic period in what is now Uintah County, Utah. It is known from the middle of the Brushy Basin Member, Morrison Formation. This genus was named by Andrew T. McDonald in 2011 and...


Gen. et comb.

Valid
  • McDonald

Late Jurassic

Morrison Formation
Morrison Formation
The Morrison Formation is a distinctive sequence of Late Jurassic sedimentary rock that is found in the western United States, which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltstone and limestone and is light grey, greenish...


A new genus for "Camptosaurus" aphanoecetes (Carpenter & Wilson, 2008).

Xiaotingia
Xiaotingia
Xiaotingia is a genus of bird-like theropod dinosaur from early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi.-Discovery:...


Gen. et sp.

Valid
  • Xu
  • You
  • et al.

early Late Jurassic

Tiaojishan Formation
Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period . It is known for its fossil plants, and is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks...


An Archaeopteryx
Archaeopteryx
Archaeopteryx , sometimes referred to by its German name Urvogel , is a genus of theropod dinosaur that is closely related to birds. The name derives from the Ancient Greek meaning "ancient", and , meaning "feather" or "wing"...

-like paravian.

Xuwulong
Xuwulong
Xuwulong is a genus of iguanodontoid dinosaur from the Early Cretaceous period. It is a basal hadrosauriform which lived during the early Cretaceous period in what is now Yujingzi Basin in the Jiuquan area, Gansu Province of northwestern China...


Gen. et sp.

Valid
  • You
  • Li
  • Liu

Early Cretaceous

Xinminpu Group

A hadrosauriform.

Zhuchengtyrannus
Zhuchengtyrannus
Zhuchengtyrannus is an extinct genus of carnivorous theropod dinosaur from the Late Cretaceous period. It is a tyrannosaurine tyrannosaurid which lived during the late Cretaceous period in what is now Zhucheng, Shandong Province of China...


Gen. et sp.

Valid
  • Hone
  • Wang
  • et al.

Late Cretaceous

Wangshi Group
Wangshi Group
The Wangshi Group is a geological formation in Shandong, China whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.-Dinosaurs:-References:...


A tyrannosaurine tyrannosaurid.

Newly named pterosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Archaeoistiodactylus
Archaeoistiodactylus
Archaeoistiodactylus is a extinct genus of istiodactylid pterosaur from the Middle Jurassic of China. It is known from an incomplete skeleton with a partial skull and lower jaw recovered from rocks of the Tiaojishan Formation in western Liaoning, China. It was first named by Lü Junchang and Fucha...


Gen. et sp.

Valid
  • Fucha

Middle Jurassic

Tiaojishan Formation
Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period . It is known for its fossil plants, and is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks...


An istiodactylid
Istiodactylidae
Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus Istiodactylus was discovered not to be a member of the genus Ornithodesmus....

.

Aurorazhdarcho
Aurorazhdarcho
Aurorazhdarcho is an extinct genus of azhdarchoid pterosaur known from the Late Jurassic of Bavaria, southern Germany.-Discovery and naming:...


Gen. et sp.

In press
  • Frey
  • Meyer
  • Tischlinger

Tithonian

Solnhofen limestone
Solnhofen limestone
The Solnhofen Plattenkalk is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organisms such as sea jellies...


An azhdarchoid.

Aussiedraco
Aussiedraco
Aussiedraco is a genus of basal ornithocheiroid pterodactyloid pterosaur from the early Cretaceous of Australia.Aussiedraco is known from holotype QM F10613, a partial mandibular symphysis housed at the Queensland Museum, recovered from rocks of the Toolebuc Formation, about 70 km east of...


Gen. et sp.

Valid
  • Kellner
  • Rodrigues
  • Costa

Albian

Toolebuc Formation
Toolebuc Formation
The Toolebuc Formation is a geological formation in Queensland, Australia whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.-Vertebrate paleofauna:...


An ornithocheiroid.

Barbosania
Barbosania
Barbosania is an extinct genus of crestless ornithocheirid pterosaur from the Cretaceous Romualdo Member of the Santana Formation of Northeastern Brazil, dating to the Albian-Cenomanian stage....


Gen. et sp.

In press
  • Elgin
  • Frey

Cenomanian

Santana Formation
Santana Formation
The Santana Formation is a geologic Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau...


An ornithocheirid.

Cuspicephalus
Cuspicephalus
Cuspicephalus is an extinct genus of monofenestratan pterosaur known from Dorset in England.-Discovery:Cuspicephalus is known from the holotype MJML K1918, a partial skull which is missing the mandible and dentition, preserved on a single slab. It was collected in December 2009 by Steve Etches...


Gen. et sp.

In press
  • Martill
  • Etches

Kimmeridgian

Kimmeridge Clay Formation

A basal monofenestratan.

Darwinopterus robustodens
Darwinopterus
Darwinopterus is a genus of pterosaur, discovered in China and named after biologist Charles Darwin. Between 30 and 40 fossil specimens have been identified, all collected from the Tiaojishan Formation, which dates to the middle Jurassic period. The type species, D. modularis, was described in...


Species

Valid
  • Xu
  • et al.

Middle Jurassic

Tiaojishan Formation
Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period . It is known for its fossil plants, and is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks...


A darwinopterid.

Gwawinapterus
Gwawinapterus
Gwawinapterus is a genus of istiodactylid pterosaur from the Late Cretaceous of North America.In May 2005 amateur paleontologist Sharon Hubbard found a rock with bones and teeth visible on the surface on a beach near Collishaw Point at Hornby Island, British Columbia, Canada...


Gen. et sp.

Valid
  • Arbour
  • Currie

Campanian

Northumberland Formation

An istiodactylid.

Jianchangopterus
Jianchangopterus
Jianchangopterus is a genus of scaphognathine rhamphorhynchid pterosaur from the Middle Jurassic of western Liaoning, China. Jianchangopterus is known from a nearly complete skeleton with skull preserved. It was collected from the Tiaojishan Formation. It was first named by Lü Junchang and Bo Xue...


Gen. et sp.

Valid
  • Bo

Middle Jurassic

Tiaojishan Formation
Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period . It is known for its fossil plants, and is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks...


A rhamphorhynchid
Rhamphorhynchidae
Rhamphorhynchidae is a group of early "rhamphorhynchoid" pterosaurs named after Rhamphorhynchus, that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley.-Classification:...

.

Navajodactylus
Navajodactylus
Navajodactylus is an extinct genus of pterosaur from Late Cretaceous deposits of San Juan Basin, New Mexico and Alberta, Canada....


Gen. et sp.

Valid
  • Sullivan
  • Fowler

Late Campanian

Kirtland Formation
Kirtland Formation
The Kirtland Formation is a sedimentary geological formation. It is the product of alluvial muds and overbank sand deposits from the many channels draining the coastal plain that existed on the inland seashore of North America, in the late Cretaceous period. It overlies the Fruitland Formation...

 

Dinosaur Park Formation
Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Judith River Group, a major geologic unit in southern Alberta. It was laid down over a period of time between about 76.5 and 75 million years ago. The formation is made up of deposits of a high-sinuosity fluvial system, and is capped...


A possible azhdarchid
Azhdarchidae
Azhdarchidae is a family of pterosaurs known primarily from the late Cretaceous Period, though an isolated vertebrae apparently from an azhdarchid is known from the early Cretaceous as well...

.

Unwindia
Unwindia
Unwindia is a genus of basal ctenochasmatoid pterosaur from the Late Cretaceous of northeastern Brazil. It is known from a partial skull recovered from rocks of the Santana Formation of the Araripe Basin, dating to Cenomanian stage. It was named by David M. Martill in 2011 and the type species is...


Gen. et sp.

Valid
  • Martill

Cenomanian

Santana Formation
Santana Formation
The Santana Formation is a geologic Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau...


A basal ctenochasmatoid.

Newly named non-mammalian synapsids

Name Novelty Status Authors Age Unit Location Notes Images

Bridetherium
Bridetherium
Bridetherium is an extinct genus of morganucodontan from Early Jurassic deposits of southern Wales, United Kingdom. Bridetherium is known from some isolated upper and lower molariform. It was collected in the Pant Quarry, Vale of Glamorgan. It was first named by William A. Clemens in 2011 and the...


Gen. et sp.

Valid
  • Clemens

Early Jurassic

A morganucodonta
Morganucodonta
The morganucodonts are an extinct group of important animals that seem to represent the earliest of mammals. These were shrew-sized, likely warm-blooded and possibly possessed mammary glands....

n.


Candelariodon
Candelariodon
Candelariodon is an extinct genus of carnivorous cynodont from Middle Triassic deposits of Rio Grande do Sul state, Brazil. Candelariodon is known from a partial mandible having some complete teeth. It was collected from the Santa Maria Formation...


Gen. et sp.

Valid
  • De Oliveira
  • Schultz
  • et al.

Middle Triassic

Santa Maria Formation
Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a late Ladinian – early Carnian age , and is notable for its fossils of early dinosaurs, including the herrerasaur Staurikosaurus, the basal saurischian Teyuwasu, and the basal sauropodomorph...


A carnivorous cynodont
Cynodont
Cynodontia or cynodonts are a taxon of therapsids which first appeared in the Late Permian and were eventually distributed throughout all seven continents by the Early Triassic . This clade includes modern mammals and their extinct close relatives. They were one of the most diverse groups of...

.

Cryptovenator
Cryptovenator
Cryptovenator is an extinct genus of sphenacodontid pelycosaur which existed in Germany during the latest Carboniferous...


Gen. et sp.

Valid
  • Fröbisch
  • Schoch
  • et al.

Late Gzhelian

Remigiusberg Formation

A sphenacodontid
Sphenacodontidae
Sphenacodontidae is a family of small to large, advanced, carnivorous, Late Pennsylvanian to middle Permian pelycosaurs. Primitive forms were generally small in size , but during the later part of the early Permian these animals grew progressively larger , to become the top predators of their...

 pelycosaur
Pelycosaur
The pelycosaurs are an informal grouping composed of basal or primitive Late Paleozoic synapsid amniotes. Some species were quite large and could grow up to 3 meters or more, although most species were much smaller...

. The type species is Cryptovenator hirschbergeri.

Euromycter
Euromycter
Euromycter is an extinct genus of large caseid synapsid from middle Early to early Late Permian deposits of Southern France. It is known from the holotype MNHN.F.MCL-2, complete skull with lower jaws and partial postcranial skeleton. It was collected by D. Sigogneau-Russell and D...


Gen. et comb.

Valid
  • Reisz
  • Maddin
  • et al.

Upper Sakmarian
Sakmarian
In the geologic timescale, the Sakmarian is an age or stage of the Permian. It is a subdivision of the Cisuralian epoch or series. The Sakmarian lasted between 294.6 ± 0.8 and 284.4 ± 0.7 million years ago...

 - Lower Lopingian

Grès Rouge Group

A large caseid
Caseidae
The Caseidae were a widespread group of very primitive herbivorous synapsids, which appeared during the later early Permian and persisted until the late middle Permian. Although ranging in size from 1 to 5½ meters in body length, caseids were surprisingly conservative in their skeletal anatomy and...

. A new genus for "Casea" rutena.

Paceyodon
Paceyodon
Paceyodon is an extinct genus of morganucodontan from Early Jurassic deposits of southern Wales, United Kingdom. Paceyodon is known from an isolated molariform that is significantly larger than any morganucodontan molariform yet discovered. It was collected in the Pant Quarry, Vale of Glamorgan. It...


Gen. et sp.

Valid
  • Clemens

Early Jurassic

A morganucodontan.

Ruthenosaurus
Ruthenosaurus
Ruthenosaurus is an extinct genus of large caseid synapsid from middle Early to early Late Permian deposits of Southern France. It is known from the holotype MNHN.F.MCL-1 a partial postcranial skeleton. It was collected by D. Sigogneau-Russell and D...


Gen. et sp.

Valid
  • Reisz
  • Maddin
  • et al.

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid
Caseidae
The Caseidae were a widespread group of very primitive herbivorous synapsids, which appeared during the later early Permian and persisted until the late middle Permian. Although ranging in size from 1 to 5½ meters in body length, caseids were surprisingly conservative in their skeletal anatomy and...

. The type species is Ruthenosaurus russellorum

Tiarajudens
Tiarajudens
Tiarajudens is an extinct genus of saber-toothed herbivorous anomodont which lived during the middle Permian period in what is now Rio Grande do Sul, Brazil. It is known from the holotype UFRGS PV393P, a nearly complete skull. The type species T...


Gen. et sp.

Valid
  • Cisneros
  • Abdala
  • et al.

Capitanian

Rio do Rasto Formation

A basal, saber-toothed herbivorous anomodont
Anomodont
The Anomodontia were a major group of therapsids, an extinct group of animals commonly known as "mammal-like reptiles." They were mostly toothless herbivores. During the Middle Permian they were very diverse, including groups like the Venyukovioidea, the Dromasauria, the Dicynodontia, and early...

. The type species is Tiarajudens eccentricus.

Newly named non-eutherian mammals

Name Novelty Status Authors Age Unit Location Notes Images

Cronopio
Cronopio (mammal)
Cronopio is an extinct genus of dryolestoid known from Río Negro of Argentina.-Description:Cronopio is known from the holotype MPCA PV 454, a partial skull which is missing the skull roof, basicranium and squamosals and from the referred specimens MPCA PV 450, a partial left...


Gen. et sp.

Valid
  • Rougier
  • Apesteguía
  • Gaetano

Cenomanian

Candeleros Formation
Candeleros Formation
The Candeleros Formation is a geologic formation that outcrops in Río Negro, Neuquén, and Mendoza provinces of Argentina. It is the oldest formation in the Neuquén Group and belongs to the Rio Limay Subgroup...


A "saber-toothed" dryolestoid.

Iberica
Iberica (mammal)
Iberica is an extinct genus of eobaatarid or a possible plagiaulacid multituberculate which existed in what is now Galve, Spain, during the early Cretaceous . It was first named by Ainara Badiola, José Ignacio Canudo and Gloria Cuenca-Bescós in 2011 and the type species is Iberica hahni....


Gen. et sp.

Valid
  • Badiola
  • Canudo
  • Cuenca-Bescós

Hauterivian/Barremian

Galve
Galve, Teruel
Galve is a municipality located in the province of Teruel, Aragon, Spain. According to the 2006 census , the municipality has a population of 145 inhabitants. There is an important paleontological site.-Paleontological significance:...


A eobaatarid or a possible plagiaulacid
Plagiaulacidae
Plagiaulacidae is a family of fossil mammals within the order Multituberculata. Remains are known from the Upper Jurassic of North America through the Lower Cretaceous of Europe...

 multituberculate.

Invictokoala
Invictokoala
Invictokoala is an extinct genus of Koala from the middle Pleistocene of central-eastern Queensland, Australia. The holotype was found during cave excavations at Mount Etna. It was first named by Gilbert J. Price and Scott A. Hocknull in 2011 and the type species is Invictokoala monticola....


Gen. et sp.

Valid
  • Price
  • Hocknull

Middle Pleistocene

A Koala
Phascolarctidae
Phascolarctidae is a family of marsupials of the order Diprotodontia, consisting of only one extant species, the Koala, six well known fossil species, with another 5 less well known fossil species, and 2 fossil species whose taxonomy is debatable but is put in this group...

.

Liaoconodon
Liaoconodon
Liaoconodon is an extinct genus of early mammal from the early Cretaceous . It is a eutriconodont which lived in what is now the Jianchang of Liaoning Province, eastern China. It is known from the holotype IVPP V 16051, which consists of nearly complete skeleton and skull...


Gen. et sp.

Valid
  • Meng
  • Wang
  • Li

Aptian

Jiufotang Formation
Jiufotang Formation
The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. . It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with...


An "eutriconodont"
Eutriconodonta
Eutriconodonta is a order of early mammals. Eutriconodonts existed in Asia, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack et al. in 1973 as a replacement name for the paraphyletic Triconodonta....

.

Naraboryctes
Naraboryctes
Naraboryctes is an extinct genus of marsupial mole from early Miocene deposits of Boodjamulla National Park of Riversleigh area, northwestern Queensland, Australia. It was first named by Michael Archer, Robin Beck, Miranda Gott, Suzanne Hand, Henk Godthelp and Karen Black in 2011 and the type...


Gen. et sp.

Valid
  • Archer
  • Beck
  • et al.

Early Miocene

A marsupial mole
Marsupial mole
Marsupial moles is a family of marsupials of the order Notoryctemorphia, consisting of only two extant species:* Notoryctes typhlops * Notoryctes caurinus ...

.

Newly named eutherians

Name Novelty Status Authors Age Unit Location Notes Images

Aegyptocetus
Aegyptocetus
Aegyptocetus is an extinct genus of protocetid archaeocete whale known from Egypt.-Description:Aegyptocetus is known from the articulated holotype MSNTUP I-15459, an almost complete cranium, lower jaws and a partial postcranial skeleton...


Gen. et sp.

Valid
  • Bianucci
  • Gingerich

Middle Eocene

Gebel Hof Formation

A protocetid cetacean.


Antarctodon
Antarctodon
Antarctodon is an extinct genus of meridiungulatan mammal from the early Eocene . It is a basal astrapotherian which lived in what is now Seymour Island, Antarctic Peninsula. The holotype and only specimen MLP 08-XI-30-1, an isolated right p4 or m1, was found in the La Meseta Formation in Cucullaea...


Gen. et sp.

Valid
  • Bond
  • Kramarz
  • et al.

Late Ypresian

La Meseta Formation

An astrapotheria
Astrapotheria
Astrapotheria is an extinct order of South American hoofed animals. The history of this order is enigmatic, but it may taxonomically belong to Meridiungulata . In turn, Meridungulata is believed to belong to the extant superorder Laurasiatheria...

n.

Betonnia
Betonnia
Betonnia is an extinct genus of cimolestid mammal which existed in New Mexico, during the early Paleocene . Its fossils have been recovered from the Nacimiento Formation, San Juan Basin, New Mexico. It was first named by Thomas E. Williamson, Anne Weil and Barbara Standhardt in 2011 and the type...


Gen. et sp.

Valid
  • Williamson
  • Weil
  • Standhardt

Middle/Late Puercan

Nacimiento Formation
Nacimiento Formation
The Nacimiento Formation is a sedimentary rock formation found in the San Juan Basin of western New Mexico and named for the Nacimiento Mountains...


A cimolestid cimolesta
Cimolesta
Cimolesta is an extinct order of mammals. A few experts place the pangolins within Cimolesta, though most other experts prefer to place the pangolins within their own order, Pholidota....

.

Balochititanops
Balochititanops
Balochititanops is an extinct genus of brontotheriid from late Early Eocene deposits of Pakistan. Balochititanops is known from the holotype GSP-UM 6532, right maxilla with teeth, from Kingri area, Balochistan. Many referred materials are known and include cranial and postcranial remains...


Gen. et sp.

Valid
  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid
Brontotheriidae
Brontotheriidae, also called Titanotheriidae, is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially they looked rather like rhinos, although they were not true rhinos and are probably most closely related to...

.

Chacomylus
Chacomylus
Chacomylus is an extinct genus of hyopsodontid condylarth which existed in the Nacimiento Formation, USA during the early Paleocene period . It was first named by Thomas E. Williamson and Anne Weil in 2010 and the type species is Chacomylus sladei....


Gen. et sp.

In press
  • Williamson
  • Weil

Paleocene

Nacimiento Formation
Nacimiento Formation
The Nacimiento Formation is a sedimentary rock formation found in the San Juan Basin of western New Mexico and named for the Nacimiento Mountains...


A hyopsodontid
Hyopsodontidae
Hyopsodontidae is an extinct family of unspecialized, primitive mammals from the Condylarthra order, living from the Paleocene to the Eocene in North America and EurasiaThey were generally small insectivorous animals. The most common genus is Hyopsodus....

 condylarth
Condylarth
Condylarthra is an order of extinct placental mammals known primarily from the Paleocene and Eocene epochs. Condylarths are among the most characteristic Paleocene mammals and they illustrate the evolutionary level of the Paleocene mammal fauna....

.

Chacopterygus
Chacopterygus
Chacopterygus is an extinct genus of cimolestid mammal which existed in New Mexico, during the early Paleocene . Its fossils have been recovered from the Nacimiento Formation, San Juan Basin, New Mexico. It was first named by Thomas E. Williamson, Anne Weil and Barbara Standhardt in 2011 and the...


Gen. et sp.

Valid
  • Williamson
  • Weil
  • Standhardt

Middle Puercan

Nacimiento Formation
Nacimiento Formation
The Nacimiento Formation is a sedimentary rock formation found in the San Juan Basin of western New Mexico and named for the Nacimiento Mountains...


A cimolestid cimolesta.

Coelodonta thibetana
Coelodonta thibetana
The Tibetan woolly rhinoceros is an extinct species of woolly rhinoceros native to western Himalayas that lived during the middle Pliocene epoch. C. thibetana is known from the holotype IVPP V15908, a partially complete skull including incomplete lower jaw preserved with full dentition...


Species

Valid
  • Deng
  • Wang
  • et al.

Middle Pliocene

Woolly
Woolly Rhinoceros
The woolly rhinoceros is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived the last glacial period. The genus name Coelodonta means "cavity tooth"...

 rhinoceros.

Comahuetherium
Comahuetherium
Comahuetherium is an extinct genus of astrapotherian mammal from the early Miocene . It is a basal astrapotheriid which lived in what is now Patagonia, Argentina...


Gen. et sp.

Valid
  • Kramarz
  • Bond

Colhuehuapian

Cerro Bandera Formation

An astrapotheriid.

Cynthiacetus peruvianus
Cynthiacetus
Cynthiacetus is an extinct genus of cetacean archaeocete, belonging to the family Basilosauridae. Lived during the Upper Eocene 38 million years ago. Was found in the southeastern United States, Egypt and Peru. Its skull was similar in size and morphology to Basilosaurus, but unlike it had no...


Species

Valid
  • Martínez-Cáceres
  • Muizon

Late Eocene-Early Oligocene

Otuma Formation

A basilosaurid cetacean.

Eotitanops pakistanensis

Species

Valid
  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid.

Griphotherion
Griphotherion
Griphotherion is an extinct genus of notoungulate mammal from the Eocene of Argentina. A fossil skeleton was found in the Lumbrera Formation and described in 2011 as the holotype of the type species G. peiranoi. Griphotherion is a small "rodent-like" notoungulate similar to members of the families...


Gen. et sp.

Valid
  • López
  • Powell

Eocene

A rodent-like notoungulate.

Hiskatherium
Hiskatherium
Hiskatherium is an extinct genus of small ground sloth from the Miocene of Bolivia. The type species H. saintandrei was named in 2011 on the basis of a lower jaw. Although it has not been placed in a specific family, Hiskatherium is similar to the extinct sloths Hapalops and Xyophorus....


Gen. et sp.

Valid
  • Pujos
  • Iuliis
  • Quispe

Santacrucian
Santacrucian
The Santacrucian age is a period of geologic time within the Miocene epoch of the Neogene used more specifically with South American Land Mammal Ages. It follows the Colhuehuapian and precedes the Friasian age....


A small ground sloth
Ground sloth
Ground sloths are a diverse group of extinct sloths, in the mammalian superorder Xenarthra. Their most recent survivors lived in the Antilles, where it has been proposed they may have survived until 1550 CE; however, the youngest AMS radiocarbon date reported is 4190 BP, calibrated to c. 4700 BP...

.

Hystrix paukensis

Species

Valid
  • Yuichiro Nishioka
  • Zin-Maung-Maung-Thein
  • et al.
late Miocene/early Pliocene new species in the genus Hystrix
Hystrix
Hystrix is a genus of porcupine that contains what are the best known and most distinctive of the Old World porcupines.-Species:*Genus Hystrix**Subgenus Thecurus***Hystrix crassispinis - Thick-spined Porcupine...

 (Old World porcupine
Old World porcupine
The Old World porcupines, or Hystricidae, are large terrestrial rodents, distinguished by the spiny covering from which they take their name. They range over the south of Europe, most of Africa, India, and the Maritime Southeast Asia as far east as Borneo...

).

Juramaia
Juramaia
Juramaia is an extinct genus of very basal eutherian mammal from late Middle Jurassic deposits of western Liaoning, China. Juramaia is known from the holotype BMNH PM1343, an articulated and nearly complete skeleton including incomplete skull preserved with full dentition...


Gen. et sp.

Valid
  • Luo
  • Yuan
  • et al.

Callovian
Callovian
In the geologic timescale, the Callovian is an age or stage in the Middle Jurassic, lasting between 164.7 ± 4.0 Ma and 161.2 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the Oxfordian....

 to Bathonian
Bathonian
In the geologic timescale the Bathonian is an age or stage of the Middle Jurassic. It lasted from approximately 167.7 Ma to around 164.7 Ma...


Tiaojishan Formation
Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period . It is known for its fossil plants, and is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks...


A basal eutherian. The type species is Juramaia sinensis.

Khoratpithecus ayeyarwadyensis
Khoratpithecus
Khoratpithecus is an extinct genus of primates that live during the late Miocene in Southeast Asia.There are three species belong to this genus:...


Species

Valid
  • Jaeger
  • Soe
  • et al.

Late Miocene
Late Miocene
The Late Miocene is a sub-epoch of the Miocene Epoch made up of two stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch....


Irrawaddy Formation

A hominoid.

Kuntinaru
Kuntinaru
Kuntinaru is an extinct genus of tolypeutine dasypodid armadillo which existed in Bolivia, during the late Oligocene . It is known from the holotype MNHN-SAL 1024, skull missing the apex of the rostrum and the paratype MNHN-SAL 3, second skull missing the apex of the rostrum, recovered from the...


Gen. et sp.

In press
  • Billet
  • Hautier
  • et al.

Deseadan

A dasypodid armadillo
Armadillo
Armadillos are New World placental mammals, known for having a leathery armor shell. Dasypodidae is the only surviving family in the order Cingulata, part of the superorder Xenarthra along with the anteaters and sloths. The word armadillo is Spanish for "little armored one"...

.

Lycophocyon
Lycophocyon
Lycophocyon is an extinct genus of caniformian carnivoran from Middle Eocene deposits of San Diego County, California. Lycophocyon is known from the holotype UCMP  85202, a partial left and right dentary...


Gen. et sp.

Valid
  • Tomiya

?Uintan-Duchesnean

Santiago Formation

A caniformia
Caniformia
Caniformia, or Canoidea , is a suborder within the order Carnivora. They typically possess a long snout and non-retractile claws . The Pinnipedia evolved from caniform ancestors and are accordingly assigned to this group...

n carnivora
Carnivora
The diverse order Carnivora |Latin]] carō "flesh", + vorāre "to devour") includes over 260 species of placental mammals. Its members are formally referred to as carnivorans, while the word "carnivore" can refer to any meat-eating animal...

n.

Megaleptictis
Megaleptictis
Megaleptictis is an extinct genus of large insectivore mammal from Late Paleogene deposits of Custer County, South Dakota. It is known from the holotype KUVP 2568 a nearly complete skull including the mandibles...


Gen. et sp.

In press
  • Meehan
  • Martin

Chadronian/?Orellan

White River Group

A large leptictid
Leptictida
Leptictida is an extinct order of placental mammals. According to cladistic studies, they may be related to Euarchontoglires , although they are more often regarded as the first branch to split from basal eutherians.- Description :The leptictids are a characteristic example of the...

.

Mescalerolemur
Mescalerolemur
Mescalerolemur is an extinct genus of adapiform strepsirrhini from late Middle Eocene deposits of Brewster County, Texas. It is known from the holotype TMM 41672-232, associated left and right maxillae with partial and complete teeth and from the referred specimens TMM 41672-230,...


Gen. et sp.

In press
  • Kirk
  • Williams

Uintan

Devil’s Graveyard Formation

A adapiform
Adapiformes
Adapiformes are an extinct group of primitive primates.The adapiformes radiated throughout much of the northern continental mass, reaching as far south as northern Africa and tropical Asia. The adapiformes existed from the Eocene to the Miocene epoch...

 strepsirrhini
Strepsirrhini
The clade Strepsirrhini is one of the two suborders of primates. Madagascar's only non-human primates are strepsirrhines, and others can be found in southeast Asia and Africa...

.

Mionothropus
Mionothropus
Mionothropus is an extinct genus of nothrotheriine nothrotheriid sloth which existed in Peru in western Amazonia, during the late Miocene . It is known from the holotype LACM 4609/117533, which includes cranial remains, recovered from the riverbank deposits of the Río Acre region, were...


Gen. et sp.

Valid
  • De Iuliis
  • Gaudin
  • Vicars

Huayquerian

A megatheriid
Megatheriidae
Megatheriidae is a family of extinct ground sloths that lived from approximately 23 mya—11,000 years ago, existing for approximately .Megatheriids appeared later in the Oligocene, some 30 million years ago, also in South America. The group includes the heavily-built Megatherium and Eremotherium...

 sloth
Sloth
Sloths are the six species of medium-sized mammals belonging to the families Megalonychidae and Bradypodidae , part of the order Pilosa and therefore related to armadillos and anteaters, which sport a similar set of specialized claws.They are arboreal residents of the jungles of Central and South...

.

Mondegodon
Mondegodon
Mondegodon is an extinct genus of basal "mesonychian" which existed in Portugal, during the earliest Eocene . It is known from the holotype – UNLSNC-19, and other parts of dentary, recovered from the locality of Silveirinha, Portugal...


Gen. et sp.

Valid
  • Tabuce
  • Clavel
  • Antunes

Earliest Eocene

A mesonychian.

Nuralagus

Gen. et sp.

Valid
  • Quintana
  • Köhler
  • Moyà-Solà

Early Pliocene

A leporid rabbit
Rabbit
Rabbits are small mammals in the family Leporidae of the order Lagomorpha, found in several parts of the world...

.

Ocucajea
Ocucajea
Ocucajea is an extinct genus of basilosaurid cetacean from Middle Eocene deposits of southern Peru. Ocucajea is known from the holotype MUSM 1442, a partial skeleton. It was collected in the Archaeocete Valley site, from the Paracas Formation of the Pisco Basin about . It was first named by...


Gen. et sp.

Valid
  • Uhen
  • Pyenson
  • et al.

Bartonian

Paracas Formation

A basilosaurid cetacean.

Phoxomylus
Phoxomylus
Phoxomylus is an extinct genus of palaechthonid plesiadapiform which existed in Alberta, Canada, during the late Paleocene . It was first named by Richard C. Fox in 2011 and the type species is Phoxomylus puncticuspis. The holotype exposed in Paskapoo Formation by Gordon P. Youzwyshyn on 10 July,...


Gen. et sp.

Valid
  • Fox

Early Tiffanian

Paskapoo Formation
Paskapoo Formation
The Paskapoo Formation is a stratigraphical unit of Paleocene age in the Western Canadian Sedimentary Basin.It takes the name from the Blindman River , and was first described in outcrops along the river, north of Red Deer by J. B...


A palaechthonid plesiadapiform.

Supayacetus
Supayacetus
Supayacetus is an extinct genus of basilosaurid cetacean from Middle Eocene deposits of southern Peru. Supayacetus is known from the holotype MUSM 1465, a partial skeleton. It was collected in the Archaeocete Valley site, from the Paracas Formation of the Pisco Basin about . It was first...


Gen. et sp.

Valid
  • Uhen
  • Pyenson
  • et al.

Bartonian

Paracas Formation

A basilosaurid cetacean.

Turiasorex
Turiasorex
Turiasorex is an extinct genus of shrew mammal which existed in east central Spain, during the middle and late Miocene. Its fossils have been recovered from the Calatayud-Daroca and Teruel basins, Spain. It was first named by Jan A. van Dam, Lars W. van den Hoek Ostende and Jelle W. F. Reumer in...


Gen. et sp.

Valid
  • van Dam
  • van den Hoek Ostende
  • Reumer

Middle - Late Miocene

A short-snouted shrew
Shrew
A shrew or shrew mouse is a small molelike mammal classified in the order Soricomorpha. True shrews are also not to be confused with West Indies shrews, treeshrews, otter shrews, or elephant shrews, which belong to different families or orders.Although its external appearance is generally that of...

.

Umayodus
Umayodus
Umayodus is an extinct genus of "condylarth" mammal from the late Paleocene or the earliest Eocene. It is a didolodontid which lived in what is now Peru. It is known from the holotype LU3-801, an isolated right third molar, which was found in the Muñami Formation of Laguna Umayo, Peru. It was first...


Gen. et sp.

In press
  • Gelfo
  • Sigé

Late Paleocene/Earliest Eocene

Muñami Formation

A didolodontid "condylarth".

Gymnosperms

Name Novelty Status Authors Age Unit Location Notes Images

Qataniaria
Qataniaria
Qataniaria is an extinct plant genus which existed in what is now Israel during the Albian period. It was described by Valentin Krassilov and Eckart Schrank in 2011, and the type species is Qataniaria noae....


Gen. et sp.

Valid
  • Krassilov
  • Schrank

Albian

Hatira Formation

Angiosperms

Name Novelty Status Authors Age Unit Location Notes Images

Palmoxylon lametaei

Species

Valid
  • Dutta
  • Ambwani
  • Estrada-Ruiz

Maastrichtian

Lameta Formation
Lameta Formation
The Lameta Formation is a sedimentary rock formation found in Madhya Pradesh, Gujarat, and Maharashtra, India. It is of Maastrichtian age , and is notable for its dinosaur fossils...


Leefructus
Leefructus
Leefructus is an extinct eudicot plant which existed in Yixian Formation, China during the early Cretaceous period. It was first named by Ge Sun, David L. Dilcher, Hongshan Wang and Zhiduan Chen in 2011 and the type species is Leefructus mirus....


Gen. et sp.

Valid
  • Sun
  • Dilcher
  • et al.

Early Cretaceous

Yixian Formation
Yixian Formation
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans 11 million years during the early Cretaceous period...


A basal eudicot

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.
The source of this article is wikipedia, the free encyclopedia.  The text of this article is licensed under the GFDL.
 
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